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Taxonomy of the family Agamidae
Bibliography of the genus Hypsilurus
Biodiversity of the family Agamidae








Genus
Hypsilurus

Eastern Anglehead Agamas*, Angle-headed Agamas, Angle-headed Lizards, Hump-headed Agamas, Narrow-headed Tree Lizards

Östliche Winkelkopfagamen*

Østlige Vinkelhovedagamer*

1867 Hypsilurus Peters (type species: Lophura godeffroyi Peters 1867)
1874 Arua Doria (type species: Gonyocephalus inornatus Doria 1874; syn. Manthey & Denzer 2006)
1878 Lophosteus Peters & Doria (type species: Gonyocephalus albertisii Peters & Doria 1878; syn. Manthey & Denzer 2006)
1893 Diptychodera Boettger (type species: Diptychodera lobata Boettger 1893; syn. Manthey & Denzer 2006)
Contents: 17 species, of which 10 (58.8%) are endemic.
Remarks: Formerly included in the genus Gonocephalus (e.g., Wermuth 1967). Previously included the genus Lophosaurus. Presence in Palau (type locality of godeffroyi) needs confirmation (see remarks under godeffroyi).
Distribution: E. Malay Archipelago, SW. Pacific Ocean.
Reported from: Indonesia (Aru Islands, Batanta, Biak, Kai Islands, Misool, Numfoor, Salawati, Sulawesi, Supiori, Western New Guinea, Yapen), Papua New Guinea (Admiralty Islands [Rambutyo Island], Bismarck Archipelago [Duke of York, New Britain, New Ireland], d'Entrecasteaux Islands [Fergusson, Normanby], Eastern New Guinea, North Solomon Islands [Bougainville], Trobriand Islands), Solomon Islands.

Hypsilurus auritus

Eared Anglehead Agama*

Ohren-Winkelkopfagame*

Vestpapuansk Vinkelhovedagam*

1874 Gonyocephalus auritus Meyer
Hypsilurus auritus Moody 1980

Distribution: Indonesia (Batanta, Misool, Salawati, Waigeo, Western New Guinea, Yapen), Papua New Guinea (Eastern New Guinea).


Hypsilurus binotatus

Two-marked Anglehead Agama*

Zweifleckiger Winkelkopfagame*

Toplettet Vinkelhovedagam*

1874 Gonyocephalus binotatus Meyer
Hypsilurus binotatus Moody 1980

Distribution: Indonesia (Aru Islands, Kai Islands, Mansinam, Western New Guinea, Yapen), Papua New Guinea (Eastern New Guinea).


Hypsilurus bruijnii

Arfak Mountains Anglehead Agama*

Arfak-Winkelkopfagame*

Arfak-vinkelhovedagam*

1878 Gonyocephalus bruijnii Peters & Doria
Hypsilurus bruijnii Moody 1980
1885 Gonyocephalus bruynii Boulenger [substitute name for Gonyocephalus bruijnii Peters & Doria 1878] (Wermuth 1967)

Distribution: Indonesia (Batanta, Western New Guinea), Papua New Guinea (Eastern New Guinea).


Hypsilurus capreolatus

Chevron-marked Anglehead Agama*

Sparren-Winkelkopfagame*

Sparre-vinkelhovedagam*

2012 Hypsilurus capreolatus Kraus & Myers

Distribution: Papua New Guinea (Eastern New Guinea).


Hypsilurus geelvinkianus

Papuan Anglehead Agama*

Papuanische Winkelkopfagame*

Papuansk Vinkelhovedagam*

1878 Gonyocephalus geelvinkianus Peters & Doria
Hypsilurus geelvinkianus Moody 1980

Distribution: Indonesia (Biak, Gag, Numfoor, Supiori, Western New Guinea), Papua New Guinea (Eastern New Guinea).


Hypsilurus godeffroyi

Greater Crested Anglehead Agama, Great Crested Anglehead Agama, Great Crested Dragon, (Papuan Anglehead Lizard, Godeffroy's Anglehead Lizard)

Große Winkelkopfagame*

Kammet Vinkelhovedagam*

1867 Lophura godeffroyi Peters
Gonyocephalus godeffroyi Boulenger 1885
Hypsilurus godeffroyi Moody 1980

Remarks: Previously included macrolepis (e.g., Wermuth 1967). The type locality, "Pelew Inseln" (=Palau), is presumed to be in error (Crombie & Pregill 1999; Bauer & Watkins-Colwell 2001) and the type is supposed to have been collected in one of the islands off northern New Guinea. The species has never been seen or collected in Palau again. However, skeletal remains of an agamid lizard from Ngeaur in Palau indicate that an agamid lizard, possibly an endemic species, occurred in Palau in pre-European times (Crombie & Pregill 1999). Nevertheless, presence of any species of the family Agamidae in Palau remains unconfirmed. A single record from Queensland, Australia, refer to longii (but see remarks under that species). The distribution given below should be regarded as tentative, as some of these may refer to hikidanus or longii (Manthey & Denzer 2006).
Distribution: Indonesia (Western New Guinea), Papua New Guinea (Bismarck Archipelago [New Britain], Eastern New Guinea, North Solomon Islands [Bougainville]).


Hypsilurus hikidanus

Enarotali Anglehead Agama*

Enarotali-Winkelkopfagame*

Enarotali-vinkelhovedagam*

2006 Hypsilurus hikidanus Manthey & Denzer

Distribution: Indonesia (Western New Guinea), Papua New Guinea (Eastern New Guinea).


Hypsilurus longii

Bismarck Anglehead Agama*

Bismarck-Winkelkopfagame*

Bismarck-vinkelhovedagam*

1877 Tiaris longii Macleay
Hypsilurus longii Manthey & Denzer 2006

Remarks: Previously regarded as a synonym of godeffroyi (e.g., Boulenger 1885; Cogger 1979). Revalidated by Manthey & Denzer (2006). A single, vouchered record from Queensland, Australia, previously questioned (e.g., Cogger 1979, 1996), was considered verified as longii by Manthey & Denzer (2006), but the alleged provenance of this specimen was rejected by Shea (2008).
Distribution: Papua New Guinea (Bismarck Archipelago [Duke of York, New Britain], North Solomon Islands [Bougainville]), Solomon Islands.


Hypsilurus macrolepis

Solomon Islands Large-scaled Anglehead Agama*, Solomons Tree Dragon

Salomonen-Winkelkopfagame*

Salomon-vinkelhovedagam*

1872 Hypsilurus macrolepis Peters

Remarks: Previously regarded as a synonym of godeffroyi (e.g., Boulenger 1885). Revalidated by Moody (1980).
Distribution: Solomon Islands.


Hypsilurus magnus

Gariau Anglehead Agama*

Gariau-Winkelkopfagame*

Gariau-vinkelhovedagam*

2006 Hypsilurus magnus Manthey & Denzer

Distribution: Indonesia (Western New Guinea), Papua New Guinea (Eastern New Guinea).


Hypsilurus modestus

Green Anglehead Agama, Lesser Forest Dragon

Grüne Winkelkopfagame*

Grøn Vinkelhovedagam*

1874 Gonyocephalus modestus Meyer
Hypsilurus modestus Moody 1980
Arua modesta Macey, Schulte, Larson, Ananyeva, Pethiagoda, Rastegar-Pouyani & Papenfuss 2000
1874 Gonyocephalus inornatus Doria (Manthey & Denzer 2006)
1893 Diptychodera lobata Boettger (Manthey & Denzer 2006)
1918 Gonyocephalys modestus carinatus Sternfeld (Manthey & Denzer 2006)

Distribution: Indonesia (Aru Islands, Kai Islands, Western New Guinea, Yapen), Papua New Guinea (Admiralty Islands [Rambutyo Island], Bismarck Archipelago [New Britain, New Ireland], Eastern New Guinea).


Hypsilurus nigrigularis

Black-throated Anglehead Agama*

Schwarzkehl-Winkelkopfagame*

Sortstrubet Vinkelhovedagam*

1874 Gonyocephalus nigrigularis Meyer
Hypsilurus nigrilabris [lapsus for nigrigularis] Moody 1980

Remarks: Known only from the type locality in West Papua. Records from other places most probably refer to magnus (Manthey & Denzer 2006).
Distribution: Indonesia (Western New Guinea).


Hypsilurus ornatus

Ornate Anglehead Agama*

Pracht-Winkelkopfagame*

Pragt-vinkelhovedagam*

2006 Hypsilurus ornatus Manthey & Denzer

Distribution: Papua New Guinea (Eastern New Guinea).


Hypsilurus papuensis

White-cheeked Anglehead Agama

Weißwangen-Streifenagame*

Hvidkindet Vinkelhovedagam*

1877 Tiaris papuensis Macleay
Gonyocephalus papuensis Boulenger 1885
Hypsilurus papuensis Moody 1980
1878 Gonyocephalys albertisii Peters & Doria (Boulenger 1885)
2006 Hypsilurus papuensis longicauda Manthey & Denzer

Distribution: Papua New Guinea (d'Entrecasteaux Islands [Fergusson, Normanby], Eastern New Guinea, Trobriand Islands).


Hypsilurus schoedei

Rambutyo Anglehead Agama*

Rambutyo-Winkelkopfagame*

Rambutyo-vinkelhovedagam*

1932 Gonyocephalus schoedei Vogt
Hypsilurus schoedei Moody 1980

Remarks: Likely to be distributed on other islands in the group, e.g., nearby Manus Island (Manthey & Denzer 2006).
Distribution: Papua New Guinea (Admiralty Islands [Rambutyo Island]).


Hypsilurus schultzewestrumi

Nondugl Anglehead Agama*

Nondugl-Winkelkopfagame*

Nondugl-vinkelhovedagam*

1999 Gonocephalus schultzewestrumi Urban
Hypsilurus schultzewestrumi Manthey & Denzer 2000

Remarks: This species was already described i 1977 in an unpublished thesis (Urban 1977), but being unpublished, it did not meet the formal requirements for a valid species description until such a description was published 22 years later (Urban 1999).
Distribution: Indonesia (Western New Guinea), Papua New Guinea (Eastern New Guinea).


Hypsilurus tenuicephalus

Narrow-headed Anglehead Agama*

Schmalköpfige Winkelkopfagame*

Smalhovedet Vinkelhovedagam*

2006 Hypsilurus tenuicephalus Manthey & Denzer

Distribution: Indonesia (Western New Guinea).