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Taxonomy of the genus Astrochelys Biodiversity of the family Testudinidae

Bibliography of the genus
Astrochelys (Madagascar Tortoises)

(Reptilia: Testudines: Testudinidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Astrochelys in general

Bour, R. 1978. Les tortues actuelles de Madagascar (Republique malgache): liste systematique et description de deux sous-espèces nouvelles (Reptilia - Testudines). Bulletin de la Societe d'Etudes Scientifiques de l'Anjou Nouvelle Serie 10: 141-154.

Schwarz, D.; Newton, D.; Ratzimbazafy, C. 2017. Assessment of the international illicit wildlife trade in the critically endangered Malagasy tortoise species Astrochelys yniphora and Astrochelys radiata. Salamandra 53(1): 163-166.

Walker, R.C.J. 2012. The internet based trade in Madagascar's critically endangered tortoise species: a preliminary study identifying the conservation threats. Testudo 7(4): 3-13.

Astrochelys radiata

Anonymous. 1956. Strikingly-marked tortoise (Testudo radiata). African Wild Life 10: 36.

Anonymous. 1993. [The radiated tortoise Asterochelys radiata]. Akvariumist (Akvariumy i Terrariumy) 5: 1.

Ancey, N. 2012. Aménagement d’un terrarium tropical pour juvéniles de tortue radiée malgache, Astrochelys radiata. Cheloniens 25: 32-37.

Auffenberg, W. 1963. Testudo hypselonota Bourret referred to Geochelone radiata (Shaw). Journal of the Bombay Natural History Society 60: 462-466.

Auffenberg, W. 1978. Courtship and breeding behavior in Geochelone radiata (Testudines: Testudinidae). Herpetologica 34(3): 277-287.

Bandy, R.E. 1970. In quest of Geochelone radiata. International Turtle and Tortoise Society Journal 4(1): 19-23, 27.

Bartlett, R.D. 1991. Life with Matty and Maddie: my years with radiated tortoises. Tropical Fish Hobbyist 39(9): 128-129, 131-134, 136.

Behler, J.L.; Iaderosa, J. 1990. A review of the captive breeding program for the radiated tortoise at the New York Zoological Society's Wildlife Survival Center. pp. 160-162. In: Beaman, K.R.; Caporaso, F.; McKeown, S. & Graff, M.D. (eds.). Proceedings of the First International Symposium on Turtles and Tortoises: conservation and captive husbandry, Chapman University, August 9-12, 1990. California Turtle and Tortoise Club, Van Nuys. 171 pp.

Bidmon, H.J. 2011. Beleuchtungsfehler bei der Haltung tropischer Landschildkröten. Schildkröten im Fokus 8(1): 31-35.

Bloxam, Q.M.C. 1988. An investigation of temperature and activity rhythms in the radiated tortoise Geochelone radiata. Dodo 25: 30-36.

Boullay, S. 1995. Repatriation of radiated tortoises, Geochelone radiata, from Reunion Island to Madagascar. Chelonian Conservation and Biology 1(4): 319-320.

Bourret, R. 1941. Notes herpétologiques sur l'Indochine francaise. XXI. reptiles et batraciens recus au laboratoire des sciences naturelles de l'université au cours de l'année 1940. Description de deux espèces nouvelles. Direction de l'Instruction Publique (Hanoi). 16 pp.

Burchfield, P.M. 1975. Hatching the radiated tortoise Testudo radiata at Brownsville Zoo. International Zoo Yearbook 15: 90-92.

Burchfield, P.M.; Doucette, C.S.; Beimler, T.F. 1980. Captive management of the radiated tortoise Geochelone radiata at Gladys Porter Zoo, Brownsville. International Zoo Yearbook 20: 1-6.

Caccone, A.; Amato, G.; Gratry, O.C.; Behler, J.L.; Powell, J.R. 1999. A molecular phylogeny of four endangered Madagascar tortoises based on mtDNA sequences. Molecular Phylogenetics and Evolution 12(1): 1-9.

Castellano, C.M.; Hudson, R. 2013. On the road again: Radiated Tortoise conservation in southern Madagascar. Turtle Survival Alliance 2013: 30.

Coutard, C.; Aupulatta, A. 2006. Note sur l'elevage et la reproduction de la Tortue rayonnee, Astrochelys radiata (Shaw, 1802). Manouria 9(32): 11-20.

Currylow, A.F.T.; Ranaivoharivelo, M.; Louis, E.E. 2014. Natural history notes: Astrochelys radiata (Radiated Tortoise). Nest predation. Herpetological Review 45(1): 116-117.

Deeming, D.C. 1988. Eggshell structure of lizards of two sub-families of the Gekkonidae. Herpetological Journal 1(6): 230-234.

Doody, J.S.; Castellano, C.M.; Rakotondrainy, R.; Ronto, W.M.; Rakotondriamanga, T.M.; Duchene, J.J.; Randria, Z. 2011. Aggregated drinking behavior of Radiated Tortoises (Astrochelys radiata) in Arid southwestern Madagascar. Chelonian Conservation and Biology 10(1): 145-146.

Dowling, H.G. 1961. Vanishing giants and enduring dwarfs - the tortoises. Animal Kingdom 64: 66-75.

Durrell, L.; Groombridge, B.; Tonge, S.; Bloxam, Q. 1989. Geochelone radiata radiated tortoise, Sokake. Occasional Papers of the IUCN Species Survival Commission 5: 96-98.

Eglis, A. 1960. Hardy rayed tortoise from Brooklyn. Herpetologica 16: 28.

Fife, R. 2006. Care and breeding of the Radiated Tortoise, Geochelone radiata. pp. 505-510. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Fuhrmann-Bacci, P.; Valenduc, C. 2014. [Two experiences of husbandry of Astrochelys radiata in the Centre Region and Auvergne]. (In French). Cheloniens 36: 30-34.

Goodman, S.M.; Pidgeon, M.; O'Connor, S. 1994. Mass mortality of Madagascar radiated tortoise caused by road construction. Oryx 28(2): 115-118.

Griffiths, C. 2008. The wrong tortoises in the right place: ecosystem restoration in Mauritius using proxy alien species. TSA Newsletter 2008 (August): 46-47.

Griffiths, C.J.; Jones, C.G.; Hansen, D.M.; Puttoo, M.; Tatayah, R.V.; Mueller, C.B.; Harris, S. 2010. The use of extant non-indigenous tortoises as a restoration tool to replace extinct ecosystem engineers. Restoration Ecology 18(1): 1-7.

Griffiths, C.J.; Zuël, N.; Tatayah, V.; Jones, C.G.; Griffiths, O.; Harris, S. 2012. The welfare implications of using exotic tortoises as ecological replacements. PLoS ONE 7(6): e39395, 1-10.

Hammer, J. 2006. Die Strahlenschildkröte (Geochelone radiata) auf Madagaskar - Start einer Populationsuntersuchung. Schildkröten im Fokus 3(3): 27-28.

Hammer, J. 2007. Die Strahlenschildkröte Madagaskars eine gepanzerte Schönheit. Reptilia (D) 12(65): 45-50.

Hammer, J. 2011. Madagaskars Südwesten - Heimat einer gestreiften Exotin. Marginata 29: 8-15.

Hammer, J.M. 2015. Reproduction as a function of living conditions: the breeding biology of the Radiated Tortoise (Astrochelys radiata) under natural and captive conditions in Southwest Madagascar. Journal of Herpetology 49(4): 633-640.

Hammer, J.M.; Ramilijaona, O. 2009. Population study on Astrochelys radiata (Shaw, 1802) in the Tsimanampetsotsa National Park, southwest Madagascar. Salamandra 45(4): 219-232.

Heinle, N.; Bidmon, H.J. 2006. Raising and breeding the Madagascar Radiated Tortoise, Geochelone radiata (Shaw, 1802): several years of maintenance and successful reproduction after adapting egg incubation temperatures to the climatic conditions of their natural habitat. pp. 484-504. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Honegger, R.E.; Schmidt, C.R. 1964. Herpetologisches aus dem Züricher Zoo. 1. Beiträge zur Haltung un Zucht verschiedener Reptilien. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 17: 339-342.

Huber, A. 2001. Schildkröten in Madagaskar. Radiata 10(2): 20-25.

Hudson, R.; Ogle, M. 2009. Building partnerships to save Madagascar's imperiled tortoises. TSA Newsletter 2009 (August): 5-7.

Iaderosa, J.; Behler, J.L.; Holmstrom, W. 1990. Captive management and reproduction of the radiated tortoise (Geochelone radiata) at the New York Zoological Society's Wildlife Survival Center. AAZPA Regional Conference Proceedings 1990: 377-381.

Iaderosa, J.; Behler, J.L.; Holmstrom, W. 1990. Radiated tortoise (Geochelone radiata) egg deposition and incubation data from the New York Zoological Society's Wildlife Survival Center. AAZPA Annual Conference Proceedings 1990: 101-106.

Juvik, J.O. 1975. The radiated tortoise of Madagascar. Oryx 13(2): 145-148.

Lehmann, K. 1980. Zum Alter von Testudo radiata in Gefangenschaft (Reptilia: Testudines: Testudinidae). Salamandra 16(1): 61.

Lemagnen, E. 2008. L'alimentation des tortues sur l'ile de La Reunion et de l'Astrochelys radiata (Shaw, 1802) en particulier. Manouria 11(41): 11-16.

Leuteritz, T.E.J. 2003. Natural history notes: Geochelone radiata (Radiated Tortoise). Nocturnal activity. Herpetological Review 34(3): 240.

Leuteritz, T.E.J. 2003. Observations on diet and drinking behaviour of Radiated Tortoises (Geochelone radiata) in southwest Madagascar. African Journal of Herpetology 52(2): 127-130.

Leuteritz, T.E.J.; Ekbia, H.R. 2008. Not all roads lead to resilience: a complex systems approach to the comparative analysis of tortoises in arid ecosystems. Ecology and Society 13(1): Unpaginated.

Leuteritz, T.E.J.; Lamb, T.; Limberaza, J.C. 2005. Distribution, status, and conservation of Radiated Tortoises (Geochelone radiata) in Madagascar. Biological Conservation 124(4): 451-461.

Leuteritz, T.E.J.; Ravolanaivo, R. 2003. Distribution, status, and biology of the Radiated Tortoise, Geochelone radiata, in southwest Madagascar. Linnaeus fund research report. Chelonian Conservation and Biology 4(3): 730-732.

Leuteritz, T.E.J.; Ravolanaivo, R. 2005. Reproductive ecology and egg production of the Radiated Tortoise (Geochelone radiata) in southern Madagascar. African Zoology 40(2): 233-242.

Maehn, M. 2000. Zum Vorkommen von Geochelone radiata (Shaw, 1802) un Pyxis arachnoides Bell, 1827 am Cap Ste. Marie, süd-Madagascar. pp. 45-49. In: Artner, H.; Meier, E. (eds.). Schildkröten. Grosses Schildkrötensymposium. 5. Jahrestagung der Schildkrötenfreunde Österreich. Salzburg, 16. bis 19. Jänner 1997. Natur und Tier-Verlag, Münster. 181 pp.

Mahn, M. 1998. Madagaskar. 2. Teil zur Biologie und Ökologie der Strahlenschildkröte (Geochelone radiata, Shaw 1802) und der Spinnenschildkröte (Pyxis arachnoides, Bell 1827). Schildkröte 1(2): 42-43.

Mahn, M. 1998. Madagaskar. Bemerkungen zur Biologie und Ökologie der Strahlenschildkröte (Geochelone radiata Shaw 1802) und der Spinnenschildkröte (Pyxis arachnoides, Bell 1827). Schildkröte 1(1): 48-51.

Mallinson, J.J.C. 1985. Radiated tortoise at the Jersey Wildlife Park. IZN (International Zoo News) 32(3): 37.

Nussbaum, R.A.; Raxworthy, C.J. 2001. Commentary on conservation of "Sokatra," the Radiated Tortoise (Geochelone radiata) of Madagascar. Amphibian & Reptile Conservation 2(1): 6-14.

O'Brien, S.; Emahalala, E.R.; Beard, V.; Rakotondrainy, R.M.; Reid, A.; Raharisoa, V.; Coulson, T. 2003. Decline of the Madagascar Radiated Tortoise Geochelone radiata due to overexploitation. Oryx 37(3): 338-343.

Paquette, S.R.; Ferguson, B.H.; Lapointe, F.J.; Louis, E.E. 2009. Conservation genetics of the Radiated Tortoise (Astrochelys radiata) population from Andohahela National Park, Southeast Madagascar, with a discussion on the conservation of this declining species. Chelonian Conservation and Biology 8(1): 84-93.

Paquette, S.R.; Lapointe, F.J. 2007. The use of shell morphometrics for the management of the endangered Malagasy Radiated Tortoise (Geochelone radiata). Biological Conservation 134(1): 31-39.

Paquette, S.R.; Louis, E.E.; Lapointe, F.J. 2010. Microsatellite analyses provide evidence of male-biased dispersal in the Radiated Tortoise Astrochelys radiata (Chelonia: Testudinidae). Journal of Heredity 101(4): 403-412.

Pawlowski, S. 2018. Eingeschleppte Reptilien auf den zentralen Seychelleninseln und deren mögliche Bedrohung für die endemische Herpetofauna. Sauria (Berlin) 40(3): 29-39.

Peters, U. 1969. Some observations on the captive breeding of the Madagascar tortoise Testudo radiata at Sydney Zoo. International Zoo Yearbook 9: 29.

Pointet, A. 1998. Notes sur la reproduction de la tortue rayonnee de Madagascar: Astrochelys radiata (Shaw, 1802). Manouria 1(1): 22-24.

Proot, M. 2008. Reflexion sur l'alimentation en captivite de la Tortue radiee, Astrochelys radiata (Shaw 1804). Cheloniens 10: 41-47.

Proot, M. 2009. Exemple d'amenagement d'un enclos pour les Tortues radiees Astrochelys radiata. Cheloniens 16: 50-57.

Prutz, A. 2010. Die Aufzucht von Strahlenschildkröten, Astrochelys radiata, und der Umbau einer Sauna. Schildkröten im Fokus 7(2): 3-10.

Rafalowski, G. 2006. Symbiose der Strahlenschildkröte mit... wem? Emys 13(4): ii-xv.

Randriamahazo, H.; Currylow, A.; Hudson, R.; Castellano, C.M. 2014. Mobilizing TSA's confiscation to reintroduction strategy for Radiated Tortoises. Turtle Survival 2014: 34-37.

Raphael, B.L.; Behler, J.L.; Castellano, C.; Sweeney, N.P.; Cook, R.A. 2001. Health assessments of free-ranging Radiated Tortoises (Geochelone radiata) in Madagascar. pp. 188-191. In: Baer, C.K.; Willette, M.M. (eds.). Proceedings. American Association of Zoo Veterinarians American Association of Wildlife Veterinarians Association of Reptilian and Amphibian Veterinarians National Association of Zoo and Wildlife Veterinarians Joint Conference Orlando, Florida September 18-23, 2001. American Association of Zoo Veterinarians, place of publication not given. 411 pp.

Rasoma, R.V.J.; Raselimanana, A.P.; Ratovonamana, Y.R.; Ganzhorn, J.U. 2013. Habitat use and diet of Astrochelys radiata in the subarid zone of southern Madagascar. Chelonian Conservation and Biology 12(1): 56-69.

Razafindrakoto, L.A. 2006. Programme "Sokake" a Madagascar. Chelonii 4: 139-142.

Reason, B. 1996. Radiated tortoise (Geochelone radiata). Rephiberary 223: 14.

Rioux Paquette, S.; Behncke, S.M.; O'Brien, S.H.; Brenneman, R.A.; Louis, E.E.; Lapointe, F.J. 2007. Riverbeds demarcate distinct conservation units of the Radiated Tortoise (Geochelone radiata) in southern Madagascar. Conservation Genetics 8(4): 797-807.

Roltberty, S. 2014. [The Radiated Tortoise of Madagascar Astrochelys radiata (Shaw, 1802)]. (In French). Cheloniens 36: 20-29.

Schmidt, A. 1959. Testudo radiata Shaw, die madagassische Strahlenschildkröte. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 12: 116-117.

Schweizer, H. 1965. Ei-Zeitigung, Aufzucht und Entwicklung einer Strahlenschildkröte (Testudo radiata Shaw). Salamandra 1: 67-73.

Seccareccia, V.J.; Lloyd, M.L.; Martin, J. 1997. Coagulation parameters of clinically normal radiated (Geochelone radiata) and leopard (Geochelone pardalis babcocki) tortoises. Proceedings of the Annual Conference of the Association of Reptilian & Amphibian Veterinarians 4: 89-91.

Silva, M.C. de; Amarasinghe, A.A.T.; Silva, A. de; Karunarathna, D.M.S.S. 2011. Mugger crocodile (Crocodylus palustris Lesson, 1831) preys on a radiated tortoise in Sri Lanka. Taprobanica 3(1): 38-41.

Vinke, T.; Vinke, S. 2005. Die Strahlenschildkröte, eine faszinierende Schönheit. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 58(9): 30-34.

Vinke, T.; Vinke, S. 2005. Geochelone radiata. Die Strahlenschildkröte, eine faszinierende Schönheit. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 58(7): 64-68.

Walker, R.C.J. 2011. The conservation issues facing the critically endangered Madagascar radiated tortoise (Astrochelys radiata). Testudo 7(3): 39-52.

Walker, R.C.J.; Rafeliarisoa, T.H. 2012. Distribution of Radiated Tortoise (Astrochelys radiata) bush meat poaching effort. Chelonian Conservation and Biology 11(2): 223-226.

Walker, R.C.J.; Rafeliarisoa, T.H.; Currylow, A.F.T.; Rakotoniaina, J.C.; Louis, E.E. 2014. Short term monitoring reveals the rapid decline of southern Madagascar's critically endangered tortoise species. Herpetological Journal 24(3): 193-196.

Wermuth, H. 1956. Die Strahlenschildkröte, Testudo radiata Shaw. Aquarien und Terrarien (Leipzig) 3: 4.

Wermuth, H. 1956. Versuch der Deutung einiger bisher übersehener Schildkröten-Namen. Zoologische Beiträge 2: 399-423.

Wermuth, H. 1962. Testudo hyposelonota Bourret. Eine revalidierte Schildkröten-Art aus Indochina. Zoologische Garten N.F. (Leipzig) 26: 228-238.

Wermuth, H. 1965. Zum Status von Testudo hypselonota Bourret. Israel Journal of Zoology 14: 277-285.

Wirth, M.; Fritz, P. 2011. [Care and breeding of the Madagascan radiated tortoise. 4.]. (In German). Marginata 32: 35-47.

Wirth, M.; Fritz, P. 2011. Haltung und Nachzucht der Madagassischen Strahlen-schildkröte Teil 1. Marginata 29: 24-30.

Wirth, M.; Fritz, P. 2011. Haltung und Nachzucht der Madagassischen Strahlenschildkröte. Teil 2. Marginata 30: 42-51.

Wirth, M.; Fritz, P. 2011. Haltung und Nachzucht der Madagassischen Strahlenschildkröte. Teil 3. Marginata 31: 35-43.

Wirth, M.; Fritz, P. 2012. [Care and breeding of Madagascan radiated tortoises. Part 6]. (In German). Marginata 34: 36-46.

Wirth, M.; Fritz, P. 2012. [Care and breeding of the Madagascan radiated tortoise. Part 5]. (In German). Marginata 33: 35-43.

Woods-Ballard, A.J.; Fanning, E. 2004. A note on the hunting of Radiated Tortoises in the Beheloka region, Southwest Madagascar. Turtle and Tortoise Newsletter 7: 25-26.

Woods-Ballard, A.J.; Fanning, E.; Rix, C.E.; Walker, R.C.J. 2004. Radiated Tortoises found on the protected Island of Nosy Ve, southwest Madagascar. Turtle and Tortoise Newsletter 7: 26-27.

Zovickian, W.H. 1973. Captive reproduction of the radiated tortoise. HISS News Journal 1(4): 115-118.

Zovickian, W.H. 1973. Reproduction of the radiated. International Turtle and Tortoise Society Journal 7(2): 26-29, 35.

Zwartepoorte, H. 2011. Wiederholte Nachzucht der Madagassischen Strahlenschildkröte (Astrochelys radiata) im Zoo Rotterdam. Marginata 31: 26-30.

Astrochelys yniphora

Anonymous. 1993. Welterstnachzucht der stark bedrohten Madagaskar-Schienenschildkröte im Exotarium des Frankfurter Zoos. Herpetofauna (Weinstadt) 15(86): 17.

Andrianandrasana, T.H. 2000. Survey of the habitat of the ploughshare tortoise Geochelone yniphora in the Baly Bay region, Madagascar. Dodo 36: 92.

Angel, F. 1932. Sur un examplaire de Testudo hyniphora Vaillant, de Madagascar. Bulletin de la Société Zoologique de France 56: 383-385.

Bour, R. 2008. La Tortue a soc de Madagascar: passe, present... et futur incertain. Le type de Testudo yniphora Vaillant, 1885. Selection bibliographique. Manouria 11(41): 26-38.

Bour, R.; Rage, J.C. 1989. A propos de Astrochelys yniphora (=Testudo hyniphora), tortue terrestre de Madagascar, et d'un manuscrit ancien de M. Mourgue. Bulletin de la Societe Herpetologique de France 50: 43-45.

Bourou, R.; Bekarany, E.; Tiandray, H.; Ratsimbazafy, J.; Durbin, J.; Razandrimamilafiniarivo, O.C. 2006. Le succes de l'essai de lacher de la tortue a soc, Geochelone yniphora a Baie de Baly, Madagascar. Chelonii 4: 133-137.

Bourou, R.; Tiandray, H.; Razandrimamilafiniarivo, O.C.; Bekarany, E.; Durbin, J. 2001. Comparative reproduction in wild and captive female ploughshare tortoises Geochelone yniphora. Dodo 37: 70-79.

Burke, R.L. 1990. Conservation of the world's rarest tortoise. Conservation Biology 4(2): 122-123.

Caccone, A.; Amato, G.; Gratry, O.C.; Behler, J.L.; Powell, J.R. 1999. A molecular phylogeny of four endangered Madagascar tortoises based on mtDNA sequences. Molecular Phylogenetics and Evolution 12(1): 1-9.

Clausen, A.; Pedrono, M. 2018. Twilight of the Angonoka: Biology and Conservation of the World’s Rarest Tortoise. Natural History Publications, Borneo. 195 pp.

Curl, D. 1986. The rarest tortoise on earth. Oryx 20(1): 35-39.

Curl, D.A.; Scoones, I.C.; Guy, M.K.; Rakotoarisoa, G. 1985. The Madagascar tortoise Geochelone yniphora: current status and distribution. Biological Conservation 34(1): 35-54.

Curl, D.L. 1986. A recovery programme for the Madagascan plowshare tortoise Astrochelys (=Geochelone) yniphora. Dodo 23: 68-79.

Currylow, A.; Walde, A.D.; Filazaha, F.; Mandimbihasina, A.; Woolaver, L. 2015. Ploughshare Tortoise (Astrochelys yniphora) natural entrapment: rarest tortoise nearly becomes slightly rarer. Herpetology Notes 8: 485-487.

Currylow, A.F.T. 2014. No refuge at the end of the earth: Beating hustlers, poverty and disease to save Ploughshares. The Tortoise 1(3): 28-37.

Currylow, A.F.T.; Mandimbihasina, A.; Gibbons, P.; Bekarany, E.; Stanford, C.B.; Louis, E.E.; Crocker, D.E. 2017. Comparative ecophysiology of a critically endangered (CR) ectotherm: Implications for conservation management. PLoS ONE 12(8): e0182004

Durbin, J.; Rajafetra, V.; Reid, D.; Razandrizanakanirina, D. 1996. Local people and Project Angonoka - conservation of the ploughshare tortoise in north-western Madagascar. Oryx 30(2): 113-120.

Durrell, L. 1994. A is for angonoka: the ploughshare tortoise project and the ABCs of species conservation. Testudo 4(1): 10-16.

Durrell, L.; Groombridge, B.; Tonge, S.; Bloxam, Q. 1989. Geochelone yniphora - ploughshare tortoise, plowshare tortoise, angulated tortoise, Argonoka. Occasional Papers of the IUCN Species Survival Commission 5: 99-102.

Durrell, L.; Rakotonindrina, R.; Reid, D.; Durbin, J. 1994. The recovery of the angonoka (Geochelone yniphora) - an integrated approach to species conservation. pp. 384-393. In: Olney, P.J.S.; Mace, G.M. & Feistner, A.T.C. (eds.). Creative conservation: interactive management of wild and captive animals. Chapman & Hall, London, Glasgow, etc. 517 pp.

Hoogmoed, M.S. 1985. Two reptiles among the 12 most threatened animals of the world. Amphibia-Reptilia 6: 101-103.

Huber, A. 2001. Schildkröten in Madagaskar. Radiata 10(2): 20-25.

Hudson, R.; Ogle, M. 2009. Building partnerships to save Madagascar's imperiled tortoises. TSA Newsletter 2009 (August): 5-7.

Juvik, J.O.; Andrianarivo, A.J.; Blanc, C.P. 1981. The ecology and status of Geochelone yniphora: a critically endangered tortoise in northwestern Madagascar. Biological Conservation 19(4): 297-316.

Juvik, J.O.; Meier, D.E.; McKeown, S. 1990. Captive husbandry and conservation of the Madagascar ploughshare tortoise Geochelone yniphora. pp. 127-137. In: Beaman, K.R.; Caporaso, F.; McKeown, S. & Graff, M.D. (eds.). Proceedings of the First International Symposium on Turtles and Tortoises: conservation and captive husbandry, Chapman University, August 9-12, 1990. California Turtle and Tortoise Club, Van Nuys. 171 pp.

Kuchling, G.; Lopez, F.J. 2000. Endoscopic sexing of juvenile captive-bred ploughshare tortoises Geochelone yniphora at Ampijoroa, Madagascar. Dodo 36: 94-95.

McKeown, S.; Juvik, J.O.; Meier, D.E. 1982. Observations on the reproductive biology of the land tortoises Geochelone emys and Geochelone yniphora in the Honolulu Zoo. Zoo Biology 1(3): 223-235.

Metrailler, S. 1995. La tortue terrestre la plus rare du monde: Astrochelys (Geochelone) yniphora. CITS Bulletin 4: 30-37.

Montalti, A. 2010. Projet "Angonoka". (Nom malgache de la tortue a soc, Astrochelys yniphora). La genese d'un beau projet d'aide a la sauvegarde d'une tortue rare. Cheloniens 20: 35-43.

Mourgue, M. 1931. Note sur Testudo hyniphora Vaillant, de Madagascar. Bull. Soc. linn. Lyon 1017: 130-132.

Mourgue, M. 1989. Observations sur deux cas simultanes de triorchidie chez Testudo yniphora Vaillant (chersine en voie d'extinction) de Madagascar. Bulletin de la Societe Herpetologique de France 50: 46-48.

O'Brien, S.; Robert, B.; Tiandray, H. 2005. Consequences of violating the recapture duration assumption of mark-recapture models: a test using simulated and empirical data from an endangered tortoise population. Journal of Applied Ecology 42(6): 1096-1104.

O'Brien, S.; Robert, B.; Tiandray, H. 2005. Hatch size, somatic growth rate and size-dependent survival in the endangered ploughshare tortoise. Biological Conservation 126(2): 141-145.

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