Bibliography of the genus Brookesia / RepFocus

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Taxonomy of the genus Brookesia

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Bibliography of the genus
Brookesia (Short-nosed Stub-tailed Chameleons)

(Reptilia: Sauria: Chamaeleonidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.

Brookesia in general

Angel, F. 1929. Contribution a l'étude systematique des Lezards appartenant aux genres Uroplatus et Brookesia. Memoires Academie Malgache 9: 1-63.

Angel, F. 1933. Sur un genre malagache nouveau de la famille des Chamaeleontides. Bulletin du Muséum d'Histoire Naturelle (Paris) (Ser. 2) 6: 443-446.

Böhme, W. 1974. Ein seltenes Zwergchamäleon aus Madagascar. Salamandra 10(2): 80-82.

Brady, L. 1995. UEA Madagascar expedition 1993. Threatened chameleons. British Ecological Society Bulletin 26(1): 38-43.

Brygoo, E.R. 1978. Reptiles sauriens Chamaeleonidae. Genre Brookesia et complement pour le genre Chamaeleo. Faune de Madagascar 47: 1-173.

Carpenter, A.I.; Robson, O. 2005. A review of the endemic chameleon genus Brookesia from Madagascar, and the rationale for its listing on CITES Appendix II. Oryx 39(4): 375-380.

Girard, F. 1995. Notes about the egg incubation of the genus Brookesia (Malagasy terrestrial chameleons). Herptile 20(3): 136-137.

Glaw, F.; Köhler, J.; Townsend, T.M.; Vences, M. 2012. Rivaling the world's smallest reptiles: discovery of miniaturized and microendemic new species of leaf chameleons (Brookesia) from northern Madagascar. PLoS ONE 7(2): e31314, 1-24.

Herrel, A.; Meyers, J.J.; Aerts, P.; Nishikawa, K.C. 2000. The mechanics of prey prehension in chameleons. Journal of Experimental Biology 203(21): 3255-3263.

Hildenhagen, T. 2007. Zur Geschichte und Etymologie des Gattungsnamens Brookesia Gray, 1865. Sekretär 7(2): 9-15.

Hillenius, D. 1986. The relationship of Brookesia, Rhampholeon and Chamaeleo (Chamaeleonidae, Reptilia). Bijdragen tot de Dierkunde (Amsterdam) 56(1): 29-38.

Hillenius, D. 1988. The skull of Chamaeleo nasutus adds more information to the relationship of Chamaeleo with Rhampholeon and Brookesia (Chamaeleonidae, Reptilia). Bijdragen tot de Dierkunde (Amsterdam) 58(1): 7-11.

Hofmann, T. 2012. A notable record of the genus Brookesia (Gray, 1865) from the extreme South of Madagascar. Sauria (Berlin) 34(1): 51-53.

Jenkins, R.K.B.; Brady, L.D.; Bisoa, M.; Rabearivony, J.; Griffiths, R.A. 2002. Forest disturbance and river proximity influence chameleon abundance in Madagascar. Biological Conservation 109(3): 407-415.

Klaver, C.J.J. 1979. A review of Brookesia systematics with special reference to lung morphology (Reptilia: Sauria: Chamaeleonidae). Bonner Zoologische Beiträge 30(1-2): 162-175.

Lutzmann, N. 2004. Females carrying males in chameleon courtship. Reptilia (GB) 35: 34-36.

Necas, P.; Schmidt, W. 2004. Geheimnisvolle Mini-Drachen. Die Erd- und Stummelschwanzchamäleons der Gattungen Brookesia und Rhampholeon. Reptilia (D) 9(48): 18-27.

Necas, P.; Schmidt, W. 2004. Mysterious mini-dragons: the stump-tailed chameleons Brookesia and Rhampholeon. Reptilia (GB) 35: 10-21.

Necas, P.; Schmidt, W. 2004. Stummelschwanzchamäleons: Miniatürdrachen des Regenwaldes: die Gattungen Brookesia und Rhampholeon. Edition Chimaira, Frankfurt am Main. 255 pp.

Randrianantoandro, J.C.; Randrianavelona, R.; Andriantsimanarilafy, R.R.; Fideline, H.E.; Rakotondravony, D.; Jenkins, R.K.B. 2007. Roost site characteristics of sympatric dwarf chameleons (genus Brookesia) from western Madagascar. Amphibia-Reptilia 28(4): 577-581.

Raxworthy, C.J. 1991. Field observations on some dwarf chameleons (Brookesia spp.) from rainforest areas of Madagascar, with the description of a new species. Journal of Zoology (London) 224(1): 11-25.

Raxworthy, C.J.; Nussbaum, R.A. 1995. Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar. Journal of Zoology (London) 235(4): 525-558.

Rieppel, O.; Crumly, C. 1997. Paedomorphosis and skull structure in Malagasy chamaeleons (Reptilia: Chamaeleoninae). Journal of Zoology (London) 243(2): 351-380.

Schimmenti, G.; Jesu, R. 1996. Brookesia exarmata sp. nov. (Reptilia, Chamaeleonidae): a new dwarf chameleon from the limestone outcrops of western Madagascar. Italian Journal of Zoology (Modena) 63(2): 193-197.

Schimmenti, G.; Jesu, R. 1997. Some significant reptile discoveries from the Antsingy Forest ("Tsingy de Bemaraha" massif, western Madagascar). pp. 317-329. In: Böhme, W.; Bischoff, W. & Ziegler, T. (eds.). Herpetologia Bonnensis. Proceedings of the 8th Ordinary General Meeting of the Societas Europaea Herpetologica, 23-27 August 1995, Bonn, Germany. Societas Europaea Herpetologica, Bonn. 414 pp.

Schmidt, W. 1997. Some annotations concerning the biology of diverse species of dwarf chamaeleons belonging to the genus Brookesia. pp. 213-216. In: Wyk, J.H. van (ed.). Proceedings of the FitzSimons Commemorative Symposium (South African lizards: 50 years of progress) and Third H.A.A. Symposium on African herpetology: held at the Transvaal Museum, Pretoria, South Africa, 11-15 October 1993. Herpetological Association of Africa, Matieland. 227 pp.

Schmidt, W. 2000. Über die Erd-, Blatt- oder Stummelschwanzchamäleons der Gattungen Brookesia und Rhampholeon. Draco 1(1): 63-67.

Townsend, T.M.; Vieites, D.R.; Glaw, F.; Vences, M. 2009. Testing species-level diversification hypotheses in Madagascar: the case of microendemic Brookesia leaf chameleons. Systematic Biology 58(6): 641-656.

Brookesia antakarana

Raxworthy, C.J.; Nussbaum, R.A. 1995. Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar. Journal of Zoology (London) 235(4): 525-558.

Scherz, M.D.; Glaw, F.; Rakotoarison, A.; Wagler, M.; Vences, M. 2018. Polymorphism and synonymy of Brookesia antakarana and B. ambreensis, leaf chameleons from Montagne d’Ambre in north Madagascar. Salamandra 54(4): 259-268.

Brookesia bekolosy

Brookesia bonsi

Ramanantsoa, G.A. 1979. Description de deux nouvelles espèces de Brookesia (Reptilia, Squamata, Chamaeleonidae): B. legendrei et B. bonsi. Bulletin du Museum National d'Histoire Naturelle Section A Zoologie Biologie et Ecologie Animales 3: 685-693.

Brookesia brunoi

Crottini, A.; Miralles, A.; Glaw, F.; James, H.D.; Lima, A.; Vences, M. 2012. Description of a new pygmy chameleon (Chamaeleonidae: Brookesia) from central Madagascar. Zootaxa 3490: 63-74.

Brookesia brygooi

Randrianantoandro, C.J.; Razafimahatratra, B.; Soazandry, M.; Ratsimbazafy, J.; Jenkins, R.K.B. 2010. Habitat use by chameleons in a deciduous forest in western Madagascar. Amphibia-Reptilia 31(1): 27-35.

Randrianantoandro, J.C.; Randrianavelona, R.; Andriantsimanarilafy, R.R.; Fideline, H.E.; Rakotondravony, D.; Jenkins, R.K.B. 2007. Roost site characteristics of sympatric dwarf chameleons (genus Brookesia) from western Madagascar. Amphibia-Reptilia 28(4): 577-581.

Randrianantoandro, J.C.; Randrianavelona, R.; Andriantsimanarilafy, R.R.; Hantalalaina, E.F.; Rakotondravony, D.; Randrianasolo, M.; Ravelomanantsao, H.L.; Jenkins, R.K.B. 2008. Identifying important areas for the conservation of dwarf chameleons (Brookesia spp.) in Tsingy de Bemaraha National Park, western Madagascar. Oryx 42(4): 578-583.

Raxworthy, C.J.; Nussbaum, R.A. 1995. Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar. Journal of Zoology (London) 235(4): 525-558.

Brookesia confidens

Brookesia decaryi

Razafimahatratra, B.; Mori, A.; Hasegawa, M. 2008. Sleeping site pattern and sleeping behavior of Brookesia decaryi (Chamaeleonidae) in Ampijoroa Dry Forest, northwestern Madagascar. Current Herpetology 27(2): 93-99.

Brookesia dentata

Brygoo, E.R.; Domergue, C.A. 1974. Notes sur les Brookesia de Madagascar. 9. Observations sur B. tuberculata Mocquard, 1894, B. ramanantsoai sp. nov. et B. peyrierasi nom. nov. (Reptilia, Squamata, Chamaeleontidae). Bulletin du Muséum National d'Histoire Naturelle (Zoologie) (Paris) 189: 1769-1782.

Glaw, F.; Vences, M.; Ziegler, T.; Böhme, W.; Kohler, J. 1999. Specific distinctness and biogeography of the dwarf chameleons Brookesia minima, B. peyrierasi and B. tuberculata (Reptilia: Chamaeleonidae): evidence from hemipenial and external morphology. Journal of Zoology (London) 247(2): 225-238.

Raxworthy, C.J.; Nussbaum, R.A. 1995. Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar. Journal of Zoology (London) 235(4): 525-558.

Brookesia desperata

Brookesia ebenaui

Boettger, O. 1880. Diagnoses reptilium at batrachoirum novorum a Carlo Ebenau in insula Nossi-Be Madagascariensi. Zoologischer Anzeiger 4: 279-283.

Brygoo, E.R.; Domergue, C.A. 1970. Notes sur les Brookesia (Caméléonidés) de Madagascar. Description d'une espèce nouvelle B. antoetrae n.sp., et des hémipénis de B. stumpffi et B. ebenaui. Remarques sur la répartition de B. stumpffi. Bulletin du Muséum National d'Histoire Naturelle (Paris) 42: 830-839.

Ramanantsoa, G.A. 1979. Description de deux nouvelles espèces de Brookesia (Reptilia, Squamata, Chamaeleonidae): B. legendrei et B. bonsi. Bulletin du Museum National d'Histoire Naturelle Section A Zoologie Biologie et Ecologie Animales 3: 685-693.

Raxworthy, C.J. 1991. Field observations on some dwarf chameleons (Brookesia spp.) from rainforest areas of Madagascar, with the description of a new species. Journal of Zoology (London) 224(1): 11-25.

Raxworthy, C.J.; Nussbaum, R.A. 1995. Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar. Journal of Zoology (London) 235(4): 525-558.

Schmidt, W. 1998. Anmerkungen über die Lebenserwartung von Chamäleons. Salamandra 34(1): 75-76.

Brookesia exarmata

Randrianantoandro, J.C.; Randrianavelona, R.; Andriantsimanarilafy, R.R.; Fideline, H.E.; Rakotondravony, D.; Jenkins, R.K.B. 2007. Roost site characteristics of sympatric dwarf chameleons (genus Brookesia) from western Madagascar. Amphibia-Reptilia 28(4): 577-581.

Randrianantoandro, J.C.; Randrianavelona, R.; Andriantsimanarilafy, R.R.; Hantalalaina, E.F.; Rakotondravony, D.; Randrianasolo, M.; Ravelomanantsao, H.L.; Jenkins, R.K.B. 2008. Identifying important areas for the conservation of dwarf chameleons (Brookesia spp.) in Tsingy de Bemaraha National Park, western Madagascar. Oryx 42(4): 578-583.

Schimmenti, G.; Jesu, R. 1996. Brookesia exarmata sp. nov. (Reptilia, Chamaeleonidae): a new dwarf chameleon from the limestone outcrops of western Madagascar. Italian Journal of Zoology (Modena) 63(2): 193-197.

Brookesia karchei

Brygoo, E.R.; Blanc, C.P.; Domergue, C.A. 1970. Notes sur les Brookesia de Madagascar: 3. Brookesia karchei n. sp. du Massif du Marojezy. Ann. Fac. Sci. Univ. Madagascar 7: 267-271.

Brookesia lambertoni

Brookesia lineata

Brookesia micra

Glaw, F.; Köhler, J.; Townsend, T.M.; Vences, M. 2012. Rivaling the world's smallest reptiles: discovery of miniaturized and microendemic new species of leaf chameleons (Brookesia) from northern Madagascar. PLoS ONE 7(2): e31314, 1-24.

Villeneuve, A.R. 2017. Habitat selection and population density of the world's smallest chameleon, Brookesia micra, on Nosy Hara, Madagascar. Herpetological Conservation and Biology 12(2): 334-341

Brookesia minima

Glaw, F.; Vences, M.; Ziegler, T.; Böhme, W.; Kohler, J. 1999. Specific distinctness and biogeography of the dwarf chameleons Brookesia minima, B. peyrierasi and B. tuberculata (Reptilia: Chamaeleonidae): evidence from hemipenial and external morphology. Journal of Zoology (London) 247(2): 225-238.

Raxworthy, C.J. 1991. Field observations on some dwarf chameleons (Brookesia spp.) from rainforest areas of Madagascar, with the description of a new species. Journal of Zoology (London) 224(1): 11-25.

Raxworthy, C.J.; Nussbaum, R.A. 1995. Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar. Journal of Zoology (London) 235(4): 525-558.

Schimmenti, G.; Jesu, R. 1996. Brookesia exarmata sp. nov. (Reptilia, Chamaeleonidae): a new dwarf chameleon from the limestone outcrops of western Madagascar. Italian Journal of Zoology (Modena) 63(2): 193-197.

Schmidt, W.; Simon, H. 1988. Brookesia minima Boettger. Sauria (Berlin) 10(4): 33-36.

Schmidt, W.; Simon, H. 1988. Die kleinste bekannte Chamäleonart der Welt: Brookesia minima. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 41(5): 90.

Brookesia perarmata

Angel, F. 1933. Sur un genre malagache nouveau de la famille des Chamaeleontides. Bulletin du Muséum d'Histoire Naturelle (Paris) (Ser. 2) 6: 443-446.

Randrianantoandro, J.C.; Randrianavelona, R.; Andriantsimanarilafy, R.R.; Fideline, H.E.; Rakotondravony, D.; Jenkins, R.K.B. 2007. Roost site characteristics of sympatric dwarf chameleons (genus Brookesia) from western Madagascar. Amphibia-Reptilia 28(4): 577-581.

Randrianantoandro, J.C.; Randrianavelona, R.; Andriantsimanarilafy, R.R.; Hantalalaina, E.F.; Rakotondravony, D.; Randrianasolo, M.; Ravelomanantsao, H.L.; Jenkins, R.K.B. 2008. Identifying important areas for the conservation of dwarf chameleons (Brookesia spp.) in Tsingy de Bemaraha National Park, western Madagascar. Oryx 42(4): 578-583.

Schimmenti, G.; Jesu, R. 1997. Some significant reptile discoveries from the Antsingy Forest ("Tsingy de Bemaraha" massif, western Madagascar). pp. 317-329. In: Böhme, W.; Bischoff, W. & Ziegler, T. (eds.). Herpetologia Bonnensis. Proceedings of the 8th Ordinary General Meeting of the Societas Europaea Herpetologica, 23-27 August 1995, Bonn, Germany. Societas Europaea Herpetologica, Bonn. 414 pp.

Brookesia peyrierasi

Brygoo, E.R.; Domergue, C.A. 1974. Notes sur les Brookesia de Madagascar. 9. Observations sur B. tuberculata Mocquard, 1894, B. ramanantsoai sp. nov. et B. peyrierasi nom. nov. (Reptilia, Squamata, Chamaeleontidae). Bulletin du Muséum National d'Histoire Naturelle (Zoologie) (Paris) 189: 1769-1782.

Glaw, F.; Vences, M.; Ziegler, T.; Böhme, W.; Kohler, J. 1999. Specific distinctness and biogeography of the dwarf chameleons Brookesia minima, B. peyrierasi and B. tuberculata (Reptilia: Chamaeleonidae): evidence from hemipenial and external morphology. Journal of Zoology (London) 247(2): 225-238.

Raxworthy, C.J. 1991. Field observations on some dwarf chameleons (Brookesia spp.) from rainforest areas of Madagascar, with the description of a new species. Journal of Zoology (London) 224(1): 11-25.

Schimmenti, G.; Jesu, R. 1996. Brookesia exarmata sp. nov. (Reptilia, Chamaeleonidae): a new dwarf chameleon from the limestone outcrops of western Madagascar. Italian Journal of Zoology (Modena) 63(2): 193-197.

Brookesia ramanantsoai

Brookesia stumpffi

Brygoo, E.R.; Domergue, C.A. 1970. Notes sur les Brookesia (Caméléonidés) de Madagascar. Description d'une espèce nouvelle B. antoetrae n.sp., et des hémipénis de B. stumpffi et B. ebenaui. Remarques sur la répartition de B. stumpffi. Bulletin du Muséum National d'Histoire Naturelle (Paris) 42: 830-839.

Lowin, A.J. 2012. Chameleon species composition and density estimates of three unprotected dry deciduous forests between Montagne d'Ambre Parc National and Ankarana Reserve Speciale in northern Madagascar. Herpetology Notes 5: 107-113.

Mertens, R. 1951. Brookesia stumpffi ein madagassischen Zwergehamaleon in Gefangenschaft. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 4: 329-330.

Raxworthy, C.J. 1991. Field observations on some dwarf chameleons (Brookesia spp.) from rainforest areas of Madagascar, with the description of a new species. Journal of Zoology (London) 224(1): 11-25.

Schmidt, H.; Henkel, F.W.; Böhme, W. 1989. Zur Haltung und Fortpflanzungsbiologie von Brookesia stumpffi Boettger, 1894. (Sauria: Chamaeleonidae). Salamandra 25(1): 14-20.

Schmidt, W. 1986. Brookesia stumpffi Boettger. Sauria (Berlin) 8(2): 41-42.

Brookesia superciliaris

Boistel, R.; Herrel, A.; Daghfous, G.; Libourel, P.A.; Boller, E.; Tafforeau, P.; Bels, V. 2010. Assisted walking in Malagasy dwarf chamaeleons. Biology Letters 6(6): 740-743.

Kaloloha, A.; Misandeau, C.; Gehring, P.S. 2011. Notes on the diversity and natural history of the snake fauna of Ambodiriana-Manompana, a protected rainforest site in north-eastern Madagascar. Herpetology Notes 4: 397-402.

Lippe, R. 1993. Erfahrungen mit dem Augenbrauen-Chamäleon. Aquarium (Bornheim) 291: 37-38.

Rabearivony, J.; Brady, L.D.; Jenkins, R.K.B.; Ravoahangimalala, O.R. 2007. Habitat use and abundance of a low-altitude chameleon assemblage in eastern Madagascar. Herpetological Journal 17(4): 247-254.

Ratsoavina, F.M.; Gehring, P.S.; Renaivoarisoa, F.J.; Rafeliarisoa, T.H.; Crottini, A.; Louis, E.E.; Vences, M. 2010. Molecular phylogeography of a widespread Malagasy leaf chameleon species, Brookesia superciliaris. Zootaxa 2554: 62-64.

Raxworthy, C.J. 1991. Field observations on some dwarf chameleons (Brookesia spp.) from rainforest areas of Madagascar, with the description of a new species. Journal of Zoology (London) 224(1): 11-25.

Brookesia therezieni

Brygoo, E.R.; Domergue, C.A. 1970. Notes sur les Brookesia de Madagascar. 5. Description de deux espèces nouvelles: B. lambertoni n. sp. et B. therezieni n. sp. (Chamaelonidae). Bulletin du Muséum National d'Histoire Naturelle (Paris) 41 [1969]: 1091-1096.

Foley, D.H. 2003. Field observations on the movement patterns and roosting behavior of the Malagasy chameleon Brookesia therezieni (Squamata: Chamaeleonidae). Herpetological Natural History 9(1): 85-88.

Brookesia thieli

Bartlett, R.D. 1993. Agile anoles and clambering chameleons. Tropical Fish Hobbyist 42(3): 148-156, 158, 160-162.

Boistel, R.; Herrel, A.; Daghfous, G.; Libourel, P.A.; Boller, E.; Tafforeau, P.; Bels, V. 2010. Assisted walking in Malagasy dwarf chamaeleons. Biology Letters 6(6): 740-743.

Brady, L. 1995. UEA Madagascar expedition 1993. Threatened chameleons. British Ecological Society Bulletin 26(1): 38-43.

Brygoo, E.R.; Domergue, C.A. 1969. Notes sur les Brookesia de Madagascar: 2. Un Brookesia des forets orientales de Madagascar, B. thieli n. sp. (Chamaeleonides). Bulletin du Muséum National d'Histoire Naturelle (Paris) (Ser. 2) 41(6): 1103-1109.

Brygoo, E.R.; Domergue, C.A. 1970. Notes sur les Brookesia (Caméléonidés) de Madagascar. Description d'une espèce nouvelle B. antoetrae n.sp., et des hémipénis de B. stumpffi et B. ebenaui. Remarques sur la répartition de B. stumpffi. Bulletin du Muséum National d'Histoire Naturelle (Paris) 42: 830-839.

Flamme, A. 1996. Vorstellung der in der Zuchtgemeinschaft Chamaeleonidae gezüchteten Chamäleonarten. Teil 4. Brookesia thieli Brygoo & Domergue, 1969. Sauria (Berlin) 18(1): 41-46.

Girard, F. 1995. Maintien en captivite du cameleon Brookesia thieli Brygoo et Domergue, 1969. Revue Francaise d'Aquariologie Herpetologie 22(1-2): 59-60.

Jenkins, R.K.B.; Brady, L.D.; Huston, K.; Kauffmann, J.; Rabearivony, J.; Raveloson, G.; Rowcliffe, J.M. 1999. The population status of chameleons within Ranomafana National Park, Madagascar, and recommendations for future monitoring. Oryx 33(1): 38-46.

Lutzmann, N.; Flanagan, A. 2004. Field observations, captive care and breeding of the stump-tailed chameleon Brookesia thieli. Reptilia (GB) 35: 28-33.

Raxworthy, C.J.; Nussbaum, R.A. 1995. Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar. Journal of Zoology (London) 235(4): 525-558.

Brookesia tristis

Brookesia tuberculata

Brygoo, E.R.; Domergue, C.A. 1974. Notes sur les Brookesia de Madagascar. 9. Observations sur B. tuberculata Mocquard, 1894, B. ramanantsoai sp. nov. et B. peyrierasi nom. nov. (Reptilia, Squamata, Chamaeleontidae). Bulletin du Muséum National d'Histoire Naturelle (Zoologie) (Paris) 189: 1769-1782.

Glaw, F.; Vences, M.; Ziegler, T.; Böhme, W.; Kohler, J. 1999. Specific distinctness and biogeography of the dwarf chameleons Brookesia minima, B. peyrierasi and B. tuberculata (Reptilia: Chamaeleonidae): evidence from hemipenial and external morphology. Journal of Zoology (London) 247(2): 225-238.

Schimmenti, G.; Jesu, R. 1996. Brookesia exarmata sp. nov. (Reptilia, Chamaeleonidae): a new dwarf chameleon from the limestone outcrops of western Madagascar. Italian Journal of Zoology (Modena) 63(2): 193-197.

Brookesia vadoni

Brygoo, E.R.; Domergue, C.A. 1968. Notes sur les Brookesia de Madagascar: 1. Description d'un nouveau Brookesia de Madagascar: B. vadoni n. sp. (Chamaeleontide). Bulletin du Muséum National d'Histoire Naturelle (Paris) (Ser. 2) 40(4): 677-682.

Brookesia valeriae

Raxworthy, C.J. 1991. Field observations on some dwarf chameleons (Brookesia spp.) from rainforest areas of Madagascar, with the description of a new species. Journal of Zoology (London) 224(1): 11-25.