Bibliography of the genus Cordylus / RepFocus

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Taxonomy of the genus Cordylus

Biodiversity of the family Cordylidae

Bibliography of the genus
Cordylus (Girdled Lizards)

(Reptilia: Sauria: Cordylidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.

Cordylus in general

Badenhorst, N.C.; Mouton, P. le F.N.; Wyk, J.H. van 1992. Climates associated with melanistic cordylid populations (Reptilia: Sauria) in the south-western Cape, South Africa. Journal of the Herpetological Association of Africa 40: 70.

Broadley, D.G. 1971. A reassessment of the northern forms currently assigned to the Cordylus cordylus group. Journal of the Herpetological Association of Africa 7: 20-22.

Broadley, D.G.; Branch, W.R. 2002. A review of the small east African Cordylus (Sauria: Cordylidae), with the description of a new species. African Journal of Herpetology 51(1): 9-34.

Daniels, S.R.; Mouton, R. le F.N.; Toit, D.A. du 2004. Molecular data suggest that melanistic ectotherms at the south-western tip of Africa are the products of Miocene climatic events: evidence from cordylid lizards. Journal of Zoology (London) 263(4): 373-383.

Fürbringer, M. 1922. Das Zungenbein der Wirbeltiere insbesondere der Reptilien und Vögel. Nachgelassene Untersuchungen über systematische Phylogenie mit besonderer Berücksichtigung der Wurzel der Säugetiere. [Herausgegeben von H. Braus]. Abh. Heidelberger Akad. Wiss. (Abt. B) (Berlin) 11: 164 pp.

Lawrence, R.F. 1937. The girdle-tailed lizard and its mites. Cape Naturalist (Cape Town) 1(4): 107-115.

Oelofsen, B.W.; Mouton, P. le F.N. 1989. Identification keys and a natural classification: the genus Cordylus as a pointing case. Journal of the Herpetological Association of Africa 36(1): 79.

Schwier, H. 2007. Gürtelschweife - die Gattung Cordylus. Reptilia (D) 12(65): 16-23.

Stejneger, L.H. 1936. Types of the amphibian and reptilian genera proposed by Laurenti in 1768. Copeia 1936(3): 133-141.

Switak, K.H. 1995. Die Gürtelschweife der Gattung Cordylus (2). TI-Magazin 122: 39-42.

Visser, J.D. 1971. Cordylus cordylus and its races in the Cape Province - A general discussion. Journal of the Herpetological Association of Africa 7: 18-20.

Wever, E.G. 1970. The lizard ear: Cordylus, Platysaurus and Gerrhosaurus. Journal of Morphology 130: 37-55.

Cordylus angolensis

Greenbaum, E.; Stanley, E.L.; Kusamba, C.; Moninga, W.M.; Goldberg, S.R.; Bursey, C.R. 2012. A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of south-eastern Democratic Republic of the Congo. African Journal of Herpetology 61(1): 14-39.

Cordylus beraduccii

Broadley, D.G.; Branch, W.R. 2002. A review of the small east African Cordylus (Sauria: Cordylidae), with the description of a new species. African Journal of Herpetology 51(1): 9-34.

Cordylus cordylus

Bates, M.F. 2007. First records of the cape girdled lizard, Cordylus cordylus (Linnaeus, 1758), in Lesotho. Navorsinge van die Nasionale Museum (Bloemfontein) 23(6): 185-194.

Brody, L.S.; Mouton, P. le F.N.; Grant, W.S. 1993. Electrophoretic analysis of the Cordylus cordylus species group in the southwestern Cape, South Africa. Amphibia-Reptilia 14(1): 19-33.

Broeckhoven, C.; Diedericks, G.; Mouton, P. le F.N. 2015. What doesn't kill you might make you stronger: functional basis for variation in body armour. Journal of Animal Ecology 84(5): 1213-1221.

Burrage, B.R. 1974. Population structure in Agama atra and Cordylus cordylus cordylus in the vicinity of De Kelders, Cape Province. Annals of the South African Museum (Cape Town) 66(1): 1-23.

Clusella-Trullas, S.; Botes, A. 2008. Faecal analysis suggests generalist diets in three species of Western Cape cordylids. African Zoology 43(1): 125-130.

Clusella-Trullas, S.; Wyk, J.H. van; Spotila, J.R. 2009. Thermal benefits of melanism in cordylid lizards: a theoretical and field test. Ecology (Washington, D.C.) 90(8): 2297-2312.

Cooper, W.E.; Steele, L.J. 1999. Lingually mediated discriminations among prey chemicals and control stimuli in cordyliform lizards: presence in a gerrhosaurid and absence in two cordylids. Herpetologica 55(3): 361-368.

Cooper, W.E.; Whiting, M.J.; Wyk, J.H. van 1997. Foraging modes of cordyliform lizards. South African Journal of Zoology 32(1): 9-13.

Cooper, W.E.; Wyk, J.H. van 1994. Absence of prey chemical discrimination by tongue-flicking in an ambush-foraging lizard having actively foraging ancestors. Ethology 97(4): 317-328.

Cooper, W.E.; Wyk, J.H. van; Mouton, P. le F.N. 1995. Strike-induced chemosensory searching is absent in the cordylid lizard, Cordylus cordylus, a scleroglossan ambush forager. Chemoecology 5-6(1): 47-52.

Cooper, W.E.; Wyk, J.H. van; Mouton, P. le F.N. 1996. Pheromonal detection and sex discrimination of conspecific substrate deposits by the rock-dwelling cordylid lizard Cordylus cordylus. Copeia 1996(4): 839-845.

Cooper, W.E.; Wyk, J.H. van; Mouton, P. le F.N. 1999. Incomplete protective refuges: selection and associated defences by a lizard, Cordylus cordylus (Squamata: Cordylidae). Ethology 105(8): 687-700.

Cordes, I.G.; Mouton, P. le F.N.; Wyk, J.H. van 1995. Sexual dimorphism in two girdled lizard species, Cordylus niger and Cordylus cordylus. South African Journal of Zoology 30(4): 187-196.

Cronje, A.F. 1989. Melanistic Cordylus cordylus populations as relict cold-adapted populations. Journal of the Herpetological Association of Africa 36(1): 80.

Diedericks, G.; Daniels, S.R. 2014. Ain't no mountain high enough, ain't no valley low enough? Phylogeography of the rupicolous Cape Girdled Lizard (Cordylus cordylus) reveals a generalist pattern. Molecular Phylogenetics and Evolution 71: 234-248.

Dzialowski, E.M.; O'Connor, M.P. 2001. Physiological control of warming and cooling during simulated shuttling and basking in lizards. Physiological and Biochemical Zoology 74(5): 679-693.

FitzSimons, V. 1930. Descriptions of new South African Reptilia and Batrachia, with Distribution Records of Allied Species in the Transvaal Museum Collection. Annals of the Transvaal Museum 14(1): 20-48.

Heerden, A.J. van; Mouton, P. le F.N.; Wyk, J.H. van 1992. Description of a contact zone between Cordylus cordylus and Cordylus niger. Journal of the Herpetological Association of Africa 40: 26.

Hewitt, J. 1932. Some new Species and Subspecies of South African Batrachians and Lizards. Annals of the Natal Museum (Pietermaritzburg) 7(1): 105-128.

Kilchenmann, W. 1977. Gürtelechsen. Aquaria (St. Gallen) 24(5): 87-90.

Kutschera, U. 1976. Aufzucht und Pflege des Zwerg-Gürtelschweifes, Cordylus cordylus. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 29(8): 282-283.

Mouton, P. le F.N. 1985. Comments on the Cordylus cordylus complex in the southwestern Cape. Journal of the Herpetological Association of Africa 31: 16-18.

Mouton, P. le F.N. 1987. Phenotypic variation among populations of Cordylus cordylus (Reptilia: Cordylidae) in the south-western Cape Province. South African Journal of Zoology 22(2): 119-129.

Mouton, P. le F.N.; Oelofsen, B.W. 1988. A model explaining patterns of geographic character variation in Cordylus cordylus (Reptilia; Cordylidae) in the south western Cape, South Africa. South African Journal of Zoology 23(1): 20-31.

Mouton, P. le F.N.; Wyk, J.H. van 1989. Cordylus minor: a valid species of South African lizard (Reptilia: Cordylidae). South African Journal of Zoology 24(4): 322-328.

Mouton, P. le F.N.; Wyk, J.H. van 1990. Taxonomic status of the melanistic forms of the Cordylus cordylus complex (Reptilia: Cordylidae) in the south-western Cape, South Africa. South African Journal of Zoology 25(1): 31-38.

Phelps, T. 2009. Cordylus cordylus (Linnaeus, 1758). Cape Girdled Lizard. African Herp News 48: 18-19.

Toit, D.A. du; Mouton, R. le F.N.; Flemming, A.F.; Niekerk, A. van 2005. Climate and the presence of generation glands in female girdled lizards: A case study of the cordylus-niger-oelofseni complex. Journal of Herpetology 39(3): 384-388.

Trullas, S.C.; Terblanche, J.S.; Wyk, J.H. van; Spotila, J.R. 2007. Low repeatability of preferred body temperature in four species of cordylid lizards: Temporal variation and implications for adaptive significance. Evolutionary Ecology 21(1): 63-79.

Wirminghaus, J.O. 1990. Observations on the behaviour of the Cape girdled lizard Cordylus cordylus (Reptilia: Cordylidae). Journal of Natural History 24(6): 1617-1627.

Cordylus imkeae

Cooper, W.E.; Whiting, M.J.; Wyk, J.H. van 1997. Foraging modes of cordyliform lizards. South African Journal of Zoology 32(1): 9-13.

Loehr, V.J.T. 2010. Der Rooiberg Gürtelschweif, Cordylus imkeae Mouton & van Wyk, 1994: Freilandbeobachtungen, Pflege und Zucht. Sauria (Berlin) 32(4): 53-60.

Mouton, P. le F.N.; Wyk, J.H. van 1994. Taxonomic status of geographical isolates in the Cordylus minor complex (Reptilia: Cordylidae): a description of three new species. Journal of the Herpetological Association of Africa 43: 6-18.

Cordylus jonesii

Boulenger, G.A. 1891. Description of a new lizard of the genus Zonurus from the Transvaal. Annals and Magazine of Natural History (Ser. 6) (London) 7: 417.

Branch, W.R.; Braack, H.H. 1990. Another reptile translocation to a national park. Koedoe 33(1): 112.

FitzSimons, V. 1930. Descriptions of new South African Reptilia and Batrachia, with Distribution Records of Allied Species in the Transvaal Museum Collection. Annals of the Transvaal Museum 14(1): 20-48.

Fritz, P. 2012. Haltung und Vermehrung des Limpopo-Gürtelschweifs Cordylus jonesii (Boulenger, 1891) im Terrarium [Husbandry and propagation of the Limpopo Spiny-tailed Lizard, Cordylus jonesii (Boulenger, 1891) in the terrarium]. Sauria (Berlin) 34(3): 43-50.

Girard, F. 2002. Cordylus tropidosternum jonesi Boulenger, 1891. Tropical Girdled Lizard. Longevity. African Herp News 34: 25.

Krabbe-Paulduro, U.; Paulduro, E. 1987. Cordylus jonesii (Boulenger). Sauria (Berlin) 9(4): 93-96.

Tasman, K.R. 1957. Less known lizards. Proceedings Trans. Rhod. Scient. Assoc. 45: 27-45.

Wheeler, P.E. 1986. Thermal acclimation of metabolism and preferred body temperature in lizards. Journal of Thermal Biology 11(3): 161-166.

Cordylus machadoi

Stanley, E.L.; Ceríaco, L.M.P.; Bandeira, S.; Valério, H.; Bates, M.F.; Branch, W.R. 2016. A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert. Zootaxa 4061(3): 201-226.

Cordylus macropholis

Bauwens, D.; Castilla, A.M.; Mouton, P. le F.N. 1999. Field body temperatures, activity levels and opportunities for thermoregulation in an extreme microhabitat specialist, the girdled lizard (Cordylus macropholis). Journal of Zoology (London) 249(1): 11-18.

Broeckhoven, C.; Diedericks, G.; Mouton, P. le F.N. 2015. What doesn't kill you might make you stronger: functional basis for variation in body armour. Journal of Animal Ecology 84(5): 1213-1221.

Fras, P. le; Mouton, N. 2011. Aggregation behaviour of lizards in the arid western regions of South Africa. African Journal of Herpetology 60(2): 155-170.

McLachlan, G.R. 1979. Distribution of Cordylus macropholis. Journal of the Herpetological Association of Africa 20: 5.

McLachlan, G.R.; Spence, J.M. 1966. Notes on Cordylus macropholis, Boulenger. Annals of the Cape Provincial Museums (Grahamstown) 5: 139-143.

Mouton, P. le F.N.; Flemming, A.F.; Searby, C.A. 1998. Active generation glands present in neonates of some cordylid lizards: a case study of Cordylus macropholis (Sauria: Cordylidae). Journal of Morphology 235(3): 177-182.

Mouton, R. le F.N.; Flemming, A.F.; Nieuwoudt, C.J. 2000. Sexual dimorphism and sex ratio in a terrestrial girdled lizard, Cordylus macropholis. Journal of Herpetology 34(3): 379-386.

Nieuwoudt, C.J.; Mouton, R. le F.N.; Flemming, A.F. 2003. Adult male aggression towards juvenile males in the large-scaled girdled lizard, Cordylus macropholis. African Journal of Herpetology 52(1): 53-59.

Nieuwoudt, C.J.; Mouton, R. le F.N.; Flemming, A.F. 2003. Aggregation behaviour and movement patterns in the large-scaled girdled lizard, Cordylus macropholis. Amphibia-Reptilia 24(3): 345-357.

Nieuwoudt, C.J.; Mouton, R. le F.N.; Flemming, A.F. 2003. Sex ratio, group composition and male spacing in the large-scaled girdled lizard, Cordylus macropholis. Journal of Herpetology 37(3): 577-580.

Power, J.H. 1933. Herpetological Notes and Records. I. Transactions Royal Society of South Africa 21(3): 211-220.

Visagie, L.; Mouton, R. le F.N.; Bauwens, D. 2005. Experimental analysis of grouping behaviour in cordylid lizards. Herpetological Journal 15(2): 91-96.

Visser, J.D. 1979. The distribution of Cordylus macropholis. Journal of the Herpetological Association of Africa 21: 8.

Cordylus marunguensis

Cordylus mclachlani

Mouton, P. le F.N. 1986. Description of a new species of Cordylus Laurenti (Reptilia: Cordylidae) from the south-western Cape, South Africa. South African Journal of Zoology 21(4): 319-324.

Mouton, P. le F.N.; Visser, J.D.; Wyk, J.H. van 1992. Morphological variation in the girdled lizard Cordylus mclachlani Mouton 1986. South African Journal of Zoology 27(1): 35-37.

Cordylus meculae

Branch, W.R.; Rödel, M.O.; Marais, J. 2005. A new species of rupicolous Cordylus Laurenti 1768 (Sauria: Cordylidae) from northern Mozambique. African Journal of Herpetology 54(2): 131-138.

Cordylus minor

Mouton, P. le F.N.; Wyk, J.H. van 1989. A revision of the taxonomic status of Cordylus cordylus minor FitzSimons, 1943 (Reptilia: Cordylidae). Journal of the Herpetological Association of Africa 36(1): 78.

Mouton, P. le F.N.; Wyk, J.H. van 1989. Cordylus minor: a valid species of South African lizard (Reptilia: Cordylidae). South African Journal of Zoology 24(4): 322-328.

Mouton, P. le F.N.; Wyk, J.H. van 1994. Taxonomic status of geographical isolates in the Cordylus minor complex (Reptilia: Cordylidae): a description of three new species. Journal of the Herpetological Association of Africa 43: 6-18.

Schmitz, S. 1973. Das Echsenporträt, der Zwerggürtelschweif. Aquarien Magazin (Stuttgart) 7(10): 429.

Cordylus namakuiyus

Cordylus niger

Busack, S.D. 1959. Methods of eating and general behavior in some Old World saurians in captivity. Herpetologica 15(4): 234.

Bustard, R. 1955. Observations on the birth of two species of lizard in the vivarium. British Journal of Herpetology 2: 6-9.

Bustard, R. 1960. South Africa girdle-tailed lizards. Aquarist and Pondkeeper 25: 32-33.

Clusella-Trullas, S.; Botes, A. 2008. Faecal analysis suggests generalist diets in three species of Western Cape cordylids. African Zoology 43(1): 125-130.

Clusella-Trullas, S.; Wyk, J.H. van; Spotila, J.R. 2009. Thermal benefits of melanism in cordylid lizards: a theoretical and field test. Ecology (Washington, D.C.) 90(8): 2297-2312.

Cooper, W.E.; Whiting, M.J. 2007. Effects of risk on flight initiation distance and escape tactics in two southern African lizard species. Acta Zoologica Sinica 53(3): 446-453.

Cooper, W.E.; Whiting, M.J.; Wyk, J.H. van 1997. Foraging modes of cordyliform lizards. South African Journal of Zoology 32(1): 9-13.

Cordes, I.G.; Mouton, P. le F.N. 1995. The cryptic significance of melanism in cordylid lizards: a case study of Cordylus niger and C. polyzonus. Journal of the Herpetological Association of Africa 44(1): 16-21.

Cordes, I.G.; Mouton, P. le F.N. 1996. The conservation status of the Saldanha-Langebaan lizard fauna. Koedoe 39(1): 71-83.

Cordes, I.G.; Mouton, P. le F.N.; Wyk, J.H. van 1995. Sexual dimorphism in two girdled lizard species, Cordylus niger and Cordylus cordylus. South African Journal of Zoology 30(4): 187-196.

Dzialowski, E.M.; O'Connor, M.P. 2001. Physiological control of warming and cooling during simulated shuttling and basking in lizards. Physiological and Biochemical Zoology 74(5): 679-693.

FitzSimons, V. 1930. Descriptions of new South African Reptilia and Batrachia, with Distribution Records of Allied Species in the Transvaal Museum Collection. Annals of the Transvaal Museum 14(1): 20-48.

Heerden, A.J. van; Mouton, P. le F.N.; Wyk, J.H. van 1992. Description of a contact zone between Cordylus cordylus and Cordylus niger. Journal of the Herpetological Association of Africa 40: 26.

Mouton, P. le F.N.; Wyk, J.H. van 1990. Taxonomic status of the melanistic forms of the Cordylus cordylus complex (Reptilia: Cordylidae) in the south-western Cape, South Africa. South African Journal of Zoology 25(1): 31-38.

Toit, D.A. du; Mouton, R. le F.N.; Flemming, A.F.; Niekerk, A. van 2005. Climate and the presence of generation glands in female girdled lizards: A case study of the cordylus-niger-oelofseni complex. Journal of Herpetology 39(3): 384-388.

Trullas, S.C.; Terblanche, J.S.; Wyk, J.H. van; Spotila, J.R. 2007. Low repeatability of preferred body temperature in four species of cordylid lizards: Temporal variation and implications for adaptive significance. Evolutionary Ecology 21(1): 63-79.

Cordylus nyikae

Broadley, D.G.; Mouton, R. le F.N. 2000. A new species of rupicolous Cordylus Laurenti from Malawi (Sauria: Cordylidae). African Journal of Herpetology 49(2): 169-172.

Cordylus oelofseni

Clusella-Trullas, S.; Botes, A. 2008. Faecal analysis suggests generalist diets in three species of Western Cape cordylids. African Zoology 43(1): 125-130.

Clusella-Trullas, S.; Wyk, J.H. van; Spotila, J.R. 2009. Thermal benefits of melanism in cordylid lizards: a theoretical and field test. Ecology (Washington, D.C.) 90(8): 2297-2312.

Costandius, E.; Mouton, R. le F.N.; Boucher, C. 2006. Conservation status of the dwarf crag lizard, Pseudocordylus nebulosus, from the Hottentots Holland Mountains, South Africa. South African Journal of Wildlife Research 36(2): 123-132.

Dzialowski, E.M.; O'Connor, M.P. 2001. Physiological control of warming and cooling during simulated shuttling and basking in lizards. Physiological and Biochemical Zoology 74(5): 679-693.

Mouton, P. le F.N.; Wyk, J.H. van 1990. Taxonomic status of the melanistic forms of the Cordylus cordylus complex (Reptilia: Cordylidae) in the south-western Cape, South Africa. South African Journal of Zoology 25(1): 31-38.

Toit, D.A. du; Mouton, R. le F.N.; Flemming, A.F.; Niekerk, A. van 2005. Climate and the presence of generation glands in female girdled lizards: A case study of the cordylus-niger-oelofseni complex. Journal of Herpetology 39(3): 384-388.

Trullas, S.C.; Terblanche, J.S.; Wyk, J.H. van; Spotila, J.R. 2007. Low repeatability of preferred body temperature in four species of cordylid lizards: Temporal variation and implications for adaptive significance. Evolutionary Ecology 21(1): 63-79.

Cordylus rhodesianus

Broadley, D.G. 1962. The herpetofauna of the Inyanga National Park. Journal of the Herpetological Association of Rhodesia 17/18: 8-10.

Hewitt, J. 1933. Descriptions of some new reptiles and a frog from Rhodesia. Occasional Papers of the Rhodesian Museum 1933(2): 45-50.

Tasman, K.R. 1957. Less known lizards. Proceedings Trans. Rhod. Scient. Assoc. 45: 27-45.

Cordylus rivae

Broadley, D.G.; Branch, W.R. 2002. A review of the small east African Cordylus (Sauria: Cordylidae), with the description of a new species. African Journal of Herpetology 51(1): 9-34.

Nielsen, S.V.; Colston, T.J. 2014. The phylogenetic position of Ethiopia's sole endemic and biogeographically enigmatic cordylid lizard, Cordylus rivae (Squamata: Cordylidae), and a discussion of its conservation status. African Journal of Herpetology 63(2): 166-176.

Cordylus tropidosternum

Collin, A. 2008. Terrarium pour cordyle tropicale, Cordylus tropidosternum (Cope, 1869). Basiliscus 2: 64-70.

Cott, H.B. 1934. The Zoological Society's Expedition to the Zambesi, 1927. 5. On a Collection of Lizards, mainly from Portuguese East Africa, with Descriptions of new Species of Zonurus, Monopeltis, and Chirindia. Proceedings of the Zoological Society of London 1934: 145-173.

Dijk, P. van; Manley, G.A.; Gallo, L. 1998. Correlated amplitude fluctuations of spontaneous otoacoustic emissions in six lizard species. Journal of the Acoustical Society of America 104(3)(1): 1559-1564.

Fritz, P. 2007. Cordylus tropidosternum care and breeding of the tropical girdled lizard. Reptilia (GB) 52: 20-25.

Fritz, P. 2007. Zwerggürtelschweife: Haltung und Vermehrung von Cordylus tropidosternum (Cope, 1869). Reptilia (D) 12(65): 24-31.

Greenbaum, E.; Stanley, E.L.; Kusamba, C.; Moninga, W.M.; Goldberg, S.R.; Bursey, C.R. 2012. A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of south-eastern Democratic Republic of the Congo. African Journal of Herpetology 61(1): 14-39.

Klerks, M.; Struijk, R.; Loehr, V.J.T. 2002. Die Haltung des Zwerggürtelschweifs Cordylus tropidosternum tropidosternum (Cope, 1869). Elaphe 10(2): 31-43.

Manley, G.A. 2009. Spontaneous otoacoustic emissions in lizards: a comparison of the skink-like lizard families Cordylidae and Gerrhosauridae. Hearing Research 255(1-2): 58-66.

Struijk, R. 2002. Cordylus tropidosternum tropidosternum Cope 1869. Tropical Girdled Lizard. Reproduction. African Herp News 34: 25-27.

Cordylus vittifer

Bates, M.F.; Broadley, D.G. 2012. Geographical distribution: Cordylus vittifer Reichenow, 1887. Common Girdled Lizard. Mozambique. African Herp News 56: 34-35.

FitzSimons, V. 1956. Reptiles and Amphibians of the Wonderboom Nature Reserve. Fauna and Flora (Pretoria) 7: 89-106.

Jacobsen, N.H.G. 1972. Some notes on the biology and behaviour of the Transvaal girdled lizard (Cordylus vittifer) in captivity. Journal of the Herpetological Association of Africa 9: 35-37.

Jacobsen, N.H.G.; Pienaar, U. de V. 1984. New reptile records from the Kruger National Park. Koedoe 26: 135-144.