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Taxonomy of the genus Cuora Biodiversity of the family Geoemydidae

Bibliography of the genus
Cuora (Asian Box Turtles)

(Reptilia: Testudines: Geoemydidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Cuora in general

Artner, H. 1998. Nomenklatur aktuell: Bestimmungsschlüssel für die Gattung Cuora. Emys 5(3): 30-32.

Cook, T. 2008. Box Turtles: A Complete Guide to Terrapene and Cuora. T.F.H. Publications, Neptune City, New Jersey. 128 pp.

Das, I. 1988. Turtles of Manas Tiger Reserve. Hamadryad 13(1): 27-28.

Ernst, C.H.; Laemmerzahl, A.F.; Lovich, J.E. 2008. A morphological review of the Cuora flavomarginata complex (Testudines: Geoemydidae). Proceedings of the Biological Society of Washington 121(3): 391-397.

Ernst, C.H.; Lovich, J.E. 2015. Interanal seam loss in Asian turtles of the Cuora flavomarginata complex (Testudines, Geoemydidae). Herpetological Bulletin 132: 1-4.

Feng, Z.J.; Zou, S.C.; Chen, C.F.; Zhao, Y.Y. 2004. [General review of study on Cuora in China, with comments on the conservation of Cuora aurocapitata]. (In Chinese, English summary). Sichuan Journal of Zoology 23(3): 243-248.

Fritz, U.; Obst, F.J. 1997. Zum taxonomischen Status von Cuora galbinifrons serrata Iverson & McCord, 1992 und Pyxidea mouhotii (Gray, 1862) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen (Dresden) 49(2): 261-279.

Honda, M.; Yasukawa, Y.; Hirayama, R.; Ota, H. 2002. Phylogenetic relationships of the Asian box turtles of the genus Cuora sensu lato (Reptilia: Bataguridae) inferred from mitochondrial DNA sequences. Zoological Science (Tokyo) 19(11): 1305-1312.

McCord, W.P.; Joseph-Ouni, M. 2002. Chelonian illustrations. 2. Asian box turtles. Reptilia (GB) 22: 30-33.

Parham, J.F.; Simison, W.B.; Kozak, K.H.; Feldman, C.R.; Shi, H. 2001. New Chinese turtles: endangered or invalid? A reassessment of two species using mitochondrial DNA, allozyme electrophoresis and known-locality specimens. Animal Conservation 4(4): 357-367.

Pauler, I. 1980. Die Schildkrötengattung Cuora. Herpetofauna (Weinstadt) 2(6): 15-18.

Pfau, B.; Jasser-Häger, I. 2006. Feeding box turtles. Radiata 15(4): 33-39.

Praschag, P. 2009. Golden coin box turtle genetics and implications for captive management in the U.S. TSA Newsletter 2009 (August): 22-23.

Price, R. 1993. An introduction to the Asian box turtles. Tropical Fish Hobbyist 41(9): 128-130, 132, 134-137.

Schilde, M. 2005. Bemerkungen zur Systematik der Gattung Cuora Gray, 1855 (sensu lato). Radiata 14(2): 4-9.

Schilde, M. 2005. Notes on the systematic of the genus Cuora Gray, 1855 (sensu lato). Radiata 14(2): 4-9.

Shi, H.; Parham, J.F.; Simison, W.B.; Wang, J.; Gong, S.; Fu, B. 2005. A report on the hybridization between two species of threatened Asian box turtles (Testudines: Cuora) in the wild on Hainan Island (China) with comments on the origin of 'serrata'-like turtles. Amphibia-Reptilia 26(3): 377-381.

Spinks, P.Q.; Shaffer, H.B. 2007. Conservation phylogenetics of the Asian box turtles (Geoemydidae, Cuora): mitochondrial introgression, numts, and inferences from multiple nuclear loci. Conservation Genetics 8(3): 641-657.

Spinks, P.Q.; Thomson, R.C.; Zhang, Y.P.; Che, J.; Wu, Y.; Shaffer, H.B. 2012. Species boundaries and phylogenetic relationships in the critically endangered Asian box turtle genus Cuora. Molecular Phylogenetics and Evolution 63(3): 656-667.

Struijk, R.; Woldring, L.A. 2005. Das Cuora-Zuchtprogramm - Ein Überblick. Radiata 14(2): 64-73.

Struijk, R.; Woldring, L.A. 2005. Cuora studbooks, and overview. Radiata 14(2): 64-73.

Tiedemann, R.; Schneider, A.R.R.; Havenstein, K.; Blanck, T.; Meier, E.; Raffel, M.; Zwartepoorte, H.; Plath, M. 2014. New microsatellite markers allow high-resolution taxon delimitation in critically endangered Asian Box Turtles, genus Cuora. Salamandra 50(3): 139-146.

Wahl, S.; McCormack, T. 2013. Suchhunde für die Schildkrötenforschung. Marginata 37: 36-40.

Webb, R.G. 1995. The date of publication of Gray's Catalogue of shield Reptiles. Chelonian Conservation and Biology 1(4): 322-323.

Zhou, T. 2007. [A survey of captive population dynamics for six endemic Chinese box turtle species (Testudines: Bataguridae: Cuora)]. (In Chinese). Sichuan Journal of Zoology 26(2): 448-450, 451.

Cuora amboinensis

Alviola, P.L.; Cosico, E.A.; Eres, E.G. 2003. Determination of relative age, sex and sex ratio of the Malayan Box Turtle (Coura amboinensis Daudin) in Pinaglubayan River, Polillo Island, Quezon, Philippines. Asia Life Sciences 12(2): 111-121.

Appleyard, D.C. 1976. Care of Malaysian box turtles (Cuora amboinensis). British Herpetological Society Newsletter 15: 15.

Artner, H. 1998. Nomenklatur aktuell: Cuora amboinensis und Cuora galbinifrons. Emys 5(4): 34-35.

Bastard, R. 1962. Lake turtle and amboina box tortoise. Aquarist and Pondkeeper 27: 151, 164.

Beil, J. 2005. Observations on keeping and reproduction of Cuora amboinensis kamaroma Rummler & Fritz, 1991. Radiata 14(2): 61-63.

Biswas, S. 1984. Some notes on the reptiles of the Andaman and Nicobar Islands. Journal of the Bombay Natural History Society 81(2): 476-481.

Brongersma, L.D. 1947. Zoological notes from Port Dickson. 1. Amphibians and reptiles. Zoologische Mededelingen (Leiden) 27: 300-308.

Dachsel, M. 1954. Cuora amboinensis Daudin, die Malayische Dosenschildkröte. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 7: 214-215.

Das, K.C.; Gupta, A. 2017. An ecological note on the new record of Cuora amboinensis (Riche in Daudin, 1801) (Reptilia: Testudines: Geoemydidae) in northeastern India. Journal of Threatened Taxa 9(7): 10459-10462.

Dathe, F. 2001. Pflege und Vermehrung von Amboina-Scharnierschildkröten Cuora amboinensis (Daudin, 1802) im Tierpark Berlin-Friedrichsfelde. Milu 10(4): 443-452.

Daudin, F.M. 1801. Histoire Naturelle, Générale et Particulière des Reptiles. Vol. 2. F. Dufart, Paris. 432 pp.

David, P.; Kumar, M.V.R. 1998. Short notes on Malayan box turtle (Cuora amboinensis) in Great Nicobar Biosphere Reserve. Cobra (Chennai) 31: 20-21.

Debry, A. 2007. Elevage et reproduction en captivite de la Tortue-boite d'Amboine, Cuora amboinensis kamaroma Rummler & Fritz, 1991. Manouria 10(35): 33-36.

Ernst, C.H.; Laemmerzahl, A.F.; Lovich, J.E. 2011. Does the "kamaroma"-plastron pattern morph occur in both Philippine subspecies of the turtle Cuora amboinensis? Proceedings of the Biological Society of Washington 124(4): 259-269.

Ernst, C.H.; Laemmerzahl, A.F.; Lovich, J.E. 2016. A morphological review of subspecies of the Asian Box Turtle, Cuora amboinensis (Testudines, Geomydidae). Proceedings of the Biological Society of Washington 129(1): 144-156.

Ernst, C.H.; Lovich, J.E.; Laemmerzahl, A.F.; Sekscienski, S. 1997. A comparison of plastral scute lengths among members of the box turtle genera Cuora and Terrapene. Chelonian Conservation and Biology 2(4): 603-607.

Farkas, B.; Sasvári, L. 1992. On freshwater turtles (Testudines) new to China. pp. 151-157. In: Korsos, Z. & Kiss, I. (eds.): Proceedings of the 6th Ordinary General Meeting of the Societas Europaea Herpetologica, 19-23 August 1991, Budapest, Hungary. Hungarian Natural History Museum, Budapest. 531 pp.

Fidenci, P. 2009. Geographic distribution: Cuora amboinensis (Malayan Box Turtle). Herpetological Review 40(4): 449.

Fidenci, P. 2010. Geographic distribution: Cuora amboinensis kamaroma (Malayan Box Turtle). Herpetological Review 41(1): 105.

Fidenci, P.; Castillo, R. 2010. Some data on the distribution, conservation status and protection of freshwater turtles in the Palawan Island group, Philippines. Testudo 7(2): 76-87.

Filella, E. 1999. Cuora amboinensis, the southeast Asia box turtle and its subspecies. Reptilia (GB) 9: 65-69.

Fritz, U.; Freitag, H. 2008. Geoemydid turtles of the Sulu Archipelago, Philippines, and a historical record of Cyclemys dentata (Gray, 1831) for Siasi Island. Herpetology Notes 1: 11-12.

Fritz, U.; Mendau, D. 2002. Ein Gattungsbastard zweier südostasiatischer Schildkröten: Cuora amboinensis kamaroma Rummler & Fritz, 1991 x Mauremys annamensis (Siebenrock, 1903). Salamandra 38(3): 129-134.

Furrer, J. 1971. Cuora amboinensis - ein angenehmer Pflegling. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 24: 60-61.

Galgon, F.; Fritz, U. 2002. Captive bred hybrids between Chinemys reevesii (Gray, 1831) and Cuora amboinensis kamaroma Rummler & Fritz, 1991. (Testudines: Geoemydidae). Herpetozoa 15(3-4): 137-148.

Gaulke, M. 1995. On the distribution of emydid turtles and the anuran genus Microhyla in the Philippines. Asiatic Herpetological Research 6: 49-52.

Gaulke, M. 2001. Die Herpetofauna von Sibaliw (Panay), einem der letzten Tieflandregenwaldgebiete der West-Visayas, Philippinen. I. Amphibien und Anmerkungen zu einer Schildkrötenart. Herpetofauna (Weinstadt) 23(130): 5-14.

Gaulke, M.; Altenbach, A.V. 2006. Die Sumpfschildkrötenfauna der Philippinen. Marginata 10: 48-53.

Gaulke, M.; Fritz, U. 1998. Distribution patterns of batagurid turtles in the Philippines. (Testudines: Bataguridae: Cuora, Cyclemys, Heosemys.). Herpetozoa 11(1-2): 3-12.

Grychta, U. 1988. Cuora amboinensis (Daudin 1802) - Die Amboina - Scharnierschildkröte ein Bericht über Haltung, Paarungsverhalten, Eiablage und gelungener Nachzucht. Sauria (Berlin) 10(4): 27-29.

Guinguene, D. 2009. Reproduction et elevage de la Tortue boite asiatique Cuora amboinensis. Cheloniens 16: 36-43.

Guo, C.; Wang, M.; Nie, L. 1998. A cytogenetic study on three species of turtle. (In Chinese, English summary). Acta Hydrobiologica Sinica 22(1): 17-25.

Helm, W. 2002. Meine Erfahrungen zu Haltung, Zucht und Aufzucht von Cuora amboinensis kamaroma Rummler & Fritz, 1991. Radiata 11(2): 23-29.

Inskeep, R. 1984. A note on the captive breeding of the box turtle Cuora amboinensis (Daudin, 1802). British Journal of Herpetology 6(10): 383-384.

Ives, I.E.; Platt, S.G.; Tasirin, J.S.; Hunowu, I.; Siwu, S.; Rainwater, T.R. 2008. Field surveys, natural history observations, and comments on the exploitation and conservation of Indotestudo forstenii, Leucocephalon yuwonoi, and Cuora amboinensis in Sulawesi, Indonesia. Chelonian Conservation and Biology 7(2): 240-248.

Kelechian, F. 2009. Histoire et succes dans la maintenance et la reproduction de la Tortue-boite Asiatique Cuora amboinensis kamaroma. Cheloniens 16: 33-35.

Kendrick, R.C.; Ades, G.W.J. 2009. Taxonomic and morphometric analysis of a trade confiscation of turtle shells from Java, Indonesia. TurtleLog 4: 1-4.

Leavitt, D.D. 1967. Cuora amboinensis. International Turtle and Tortoise Society Journal 1(3): 15.

Mudde, P. 1982. [Cuora amboinensis the Amboina box turtle]. (In Dutch). Lacerta 40(10-11): 261-263.

Mudde, P.M. 1981. [The Malayan box turtle (Cuora amboinensis)]. (In Dutch). Lacerta 39(6-7): 65-67.

Muddle, P. 1999. Breeding the amboina box turtle Cuora amboinensis kamaroma. Reptilia (GB) 9: 60-64.

Natchev, N.; Heiss, E.; Lemell, P.; Stratev, D.; Weisgram, J. 2009. Analysis of prey capture and food transport kinematics in two Asian box turtles, Cuora amboinensis and Cuora flavomarginata (Chelonia, Geoemydidae), with emphasis on terrestrial feeding patterns. Zoology (Jena) 112(2): 113-127.

Nickl, S. 2010. Erfahrungen bei der Pflege und Vermehrung der Amboina-Scharnierschildkröte, Cuora amboinensis (Riche in Daudin, 1801). Radiata 19(3): 2-11.

Platt, S.G.; Lee, R.J.; Klemens, M.W. 2001. Notes on the distribution, life history, and exploitation of turtles in Sulawesi, Indonesia, with emphasis on Indotestudo forstenii and Leucocephalon yuwonoi. Chelonian Conservation and Biology 4(1): 154-159.

Rummler, H.J.; Fritz, U. 1991. Geographische Variabilität der Amboina-Scharnierschildkröte Cuora amboinensis (Daudin, 1802) mit Beschreibung einer neuen Unterart, C. a. kamaroma subsp. nov. Salamandra 27(1-2): 17-45.

Saxena, A. 1994. Captive breeding of the Malayan box turtle (Cuora amboinensis) from the Nicobar Islands. Hamadryad 19: 93-94.

Schoppe, S. 2008. Science in CITES: the biology and ecology of the Southeast Asian box turtle and its uses and trade in Malaysia. TRAFFIC Southeast Asia, Petaling Jaya. 56 pp.

Schoppe, S. 2010. Die kleinste südostasiatische Scharnierschildkröte (Cuora amboinensis). Marginata 26: 40-45.

Schoppe, S.; Das, I. 2011. Cuora amboinensis (Riche in Daudin 1801) - Southeast Asian Box Turtle. Chelonian Research Monographs 5: 053.1-053.13.

Singh, R.R.; Singh, K.J. 2012. Box turtles in and adjacent to Loktak Lake, Manipur - India. Taprobanica 4(2): 103-104.

Som, S.; Cottet, M. 2016. Rescue and relocation programme of turtles and tortoises and Elongated Tortoise monitoring programme in the Nam Theun 2 Reservoir (Laos). Hydroecologie Appliquee 19: 383-406.

Vogrinc, P.N.; Ryan, J.R.M. 2016. Natural history notes: Cuora amboinensis (Southeast Asian Box Turtle). Diet. Herpetological Review 47(1): 121-122.

Wangyal, J.T.; Wangchuk, D.; Das, I. 2012. First report of turtles from the Himalayan Kingdom of Bhutan. Chelonian Conservation and Biology 11(2): 268-272.

Wermuth, H. 1956. Versuch der Deutung einiger bisher übersehener Schildkröten-Namen. Zoologische Beiträge 2: 399-423.

Cuora aurocapitata

Artner, H. 2004. Haltung und Nachzucht von Pan's Scharnierschildkröte Cuora pani pani Song, 1984 und der Goldkopf-Scharnierschildkröte Cuora pani aurocapitata Luo & Zong, 1988. Emys 11(1): 4-21.

Artner, H. 2006. Elevage et reproduction de la tortue-boite de Pan, Cuora pani pani Song, 1984 et de la tortue-boite a tete jaune, Cuora pani aurocapitata Luo & Zong, 1988. Manouria 9(32): 27-39.

Artner, H. 2006. Keeping and breeding of Pan's box turtle Cuora pani pani Song, 1984 and Cuora pani aurocapitata Luo & Zong, 1988. pp. 52-68. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Blanck, T.; Kremser, T. 2007. Bemerkungen zur Verbreitung von Cuora pani aurocapitata sowie neue Fundortnachweise. 1. Marginata 13: 52-55.

Blanck, T.; Kremser, T. 2007. Bemerkungen zur Verbreitung von Cuora pani aurocapitata sowie neue Fundortnachweise. 2. Marginata 14: 50-52.

Blanck, T.; Ting, Z.; Protiva, T.; Crow, P.; Tiedemann, R. 2017. New subspecies of Cuora cyclornata (Blanck, McCord & Le, 2006), Cuora trifasciata (Bell, 1825) and Cuora aurocapitata (Luo & Zong, 1988). Sichuan Journal of Zoology 36(4): 368-385.

Bruin, R.W.F. de; Zwartepoorte, H. 1994. Captive management and breeding of Cuora aurocapitata (Testudines: Emydidae). Herpetological Review 25(2): 58-59.

Feng, Z.J.; Wang, J.; Liu, X.; Zhou, X.; Lu, A.P.; Zhao, Y.Y.; Zou, S.C. 2006. [Research and protection of golden-headed box turtle (Cuora aurocapitata)]. (In Chinese, English summary). Sichuan Journal of Zoology 25(2): 326-328.

Feng, Z.J.; Zou, S.C.; Chen, C.F.; Zhao, Y.Y. 2004. [General review of study on Cuora in China, with comments on the conservation of Cuora aurocapitata]. (In Chinese, English summary). Sichuan Journal of Zoology 23(3): 243-248.

Liu, H.Q.; Zhou, S.B.; Wang, Y.; Wu, X.; Chen, M.L. 2007. [Vegetation diversity of wild Cuora aurocapitata habitats]. (In Chinese, English summary). Shengtaixue Zazhi 26(12): 1996-2002.

Lu, W.; Hou, M.I. 2003. [Description of captive breeding of Cuora aurocapitata for 15 years]. (In Chinese, English summary). Sichuan Journal of Zoology 22(1): 22-23, 48.

Struijk, R.P.J.H. 2018. Herpetoculture notes: Cuora aurocapitata (Yellow-headed Box Turtle). Reproduction. Herpetological Review 49(4): 697-698.

Tang, X.S.; Cheng, B.G.; Ouyang, L.W. 2005. [Investigation on distribution and existence current status of golden-headed box turtle]. (In Chinese, English summary). Chinese Journal of Zoology 40(6): 99-102.

Wan, Q.; Li, F.; Shen, B.P.; Wang, B.; Xu, J.J. 2010. [Investigation on the wild living environment and current resources status of golden-headed box turtle]. (In Chinese). Changjiang Liuyu Ziyuan Yu Huanjing 19(11): 1262-1269.

Wang, Y.Y.; Wu, X.; Gu, C.M.; Peng, Y.; Qian, X.J.; Liu, H.Q. 2007. [Study on survey and conservation of golden-headed box turtle's habitats. pp. 235-244]. (In Chinese). In: Ji, X. Herpetologica Sinica 11. China Scientific Book Services, Beijing. 283 pp.

Zhang, Y.Y.; Nie, L.W.; Huang, Y.; Pu, Y.G.; Zhang, L. 2009. The mitochondrial DNA control region comparison studies of four hinged turtles and its phylogentic significance of the genus Cuora sensu lato (Testudinata: Geoemydidae). Genes & Genomics 31(5): 349-359.

Zou, S.; Chen, C.; Yang, K. 1996. The Cuora aurocapitata and it's present critical situation. Chinese Journal of Zoology 31(3): 11-12.

Cuora bourreti

Fiebig, J.; Lehr, E. 2000. Haltung und Erstnachzucht der Bourret-Scharnierschildkröte Cuora galbinifrons bourreti Obst & Reimann, 1994, mit Anmerkungen zum Bedrohungsstatus. Salamandra 36(3): 147-156.

Fritz, U.; Petzold, A.; Auer, M. 2006. Osteology in the Cuora galbinifrons complex suggests conspecifity of C. bourreti and C. galbinifrons, with notes on shell osteology and phalangeal formulae within the Geoemydidae. Amphibia-Reptilia 27(2): 195-205.

Heuberger, W.; Heuberger, K. 2008. Erfolgreiche Haltung und mehrfache Nachzucht von CistoClemmys (galbinifrons) bouretti, der Hinterindischen Scharnierschildkröte. Reptilia (D) 13(72): 64-71.

Janssen, M.; Janssen, J. 2008. Beobachtungen und Erfahrungen bei Haltung und Nachzucht der Gelbkopf-Scharnierschildkröte Cuora galbinifrons bourreti. Marginata 17: 22-27.

Jarvis, P.; Augustine, L. 2018. Herpetoculture: Cuora bourreti (Bourret's Box Turtle). Brumation, oviposition and incubation. Herpetological Review 49(3): 486-487.

Obst, F.J.; Reimann, M. 1994. Bemerkenswerte Variabilität bei Cuora galbinifrons Bourret, 1939, mit Beschreibung einer neuen geographischen Unterart: Cuora galbinifrons bourreti subsp. nov. (Reptilia: Testudines: Cryptodira: Bataguridae). Zoologische Abhandlungen (Dresden) 48(1): 125-138.

Struijk, R.P.J.H.; Blanck, T.E.G. 2015. Herpetoculture notes: Cuora mouhotii mouhotii (Northern Keeled Box Turtle) and C. bouretti (Bourret's Box Turtle). Captive hybridization. Herpetological Review 46(2): 206.

Stuart, B.L.; Hallam, C.D.; Sayavong, S.; Nanthavong, C.; Sayaleng, S.; Vongsa, O.; Robichaud, W.G. 2011. Two additions to the turtle fauna of Laos. Chelonian Conservation and Biology 10(1): 113-116.

Cuora cyclornata

Blanck, T.; McCord, W.P.; Le, M. 2006. On the variability of Cuora trifasciata (Bell, 1825). Chimaira, Frankfurt a.M. 155 pp.

Blanck, T.; Ting, Z.; Protiva, T.; Crow, P.; Tiedemann, R. 2017. New subspecies of Cuora cyclornata (Blanck, McCord & Le, 2006), Cuora trifasciata (Bell, 1825) and Cuora aurocapitata (Luo & Zong, 1988). Sichuan Journal of Zoology 36(4): 368-385.

Philippen, H.D. 2008. Meiers Dreistreifen-Scharnierschildkröte Cuora cyclornata meieri Blanck, McCord & Le, 2006. Marginata 18: 31-34.

Cuora evelynae

Ernst, C.H.; Lovich, J.E. 1990. A new species of Cuora (Reptilia: Testudines: Emydidae) from the Ryukyu Islands. Proceedings of the Biological Society of Washington 103(1): 26-34.

Ernst, C.H.; Lovich, J.E.; Laemmerzahl, A.F.; Sekscienski, S. 1997. A comparison of plastral scute lengths among members of the box turtle genera Cuora and Terrapene. Chelonian Conservation and Biology 2(4): 603-607.

Hennen, U. 2004. Die Gelbrand-Scharnierschildkröte der Ryukyu-Inseln - Cuora flavomarginata evelynae. Schildkröten im Fokus 1(1): 3-11.

Cuora flavomarginata

Anonymous. 2013. [The morphological differences analysis of Cuora flavomarginata in Central China and southern China]. (In Chinese). Xinyang Shifan Xueyuan Xuebao Ziran Kexue Ban 26(2): 240-244, 295.

Bartlett, R.D.; Bartlett, P.P. 2018. Herpetoculture: Cuora flavomarginata (Yellow-margined Box Turtle). Longevity and macrocephaly. Herpetological Review 49(3): 487-488.

Becker, H. 1996. Bemerkungen zur Haltung und Nachzucht von Cuora flavomarginata flavomarginata (Gray, 1863). Salamandra 32(2): 65-72.

Becker, H. 1999. Erganzende Bemerkungen zur Haltung und zur Nachzucht von Cuora flavomarginata (Gray, 1863). Elaphe 7(3): 2-10.

Buskirk, J.R. 1989. New locality records for Chinese non-marine chelonians. Chinese Herpetological Research 2(2): 65-68.

Chen, T.H.; Lue, K.Y. 2002. Growth patterns of the yellow-margined box turtle (Cuora flavomarginata) in northern Taiwan. Journal of Herpetology 36(2): 201-208.

Chen, T.H.; Lue, K.Y. 2008. Thermal preference of the yellow-margined box turtle (Cuora flavomarginata) (Testudines: Geoemydidae) inhabiting a mesic lowland forest, northern Taiwan. Amphibia-Reptilia 29(4): 513-522.

Chen, T.Y.; Lee, Y.T.; Chi, C.H. 2011. Observation of reproductive cycle of female yellow-margined box turtle (Cuora flavomarginata) using radiography and ultrasonography. Zoo Biology 30(6): 689-698.

Coutard, C. 2007. Note sur l'elevage et la reproduction de la Tortue-boite a bords jaunes, CistoClemmys flavomarginata (Gray, 1863). Manouria 10(34): 16-26.

Egretzberger, G. 2005. Experiences made with the husbandry of the Yellow-margined Box Turtle, Cuora flavomarginata flavomarginata (Gray, 1863). Radiata 14(2): 52-54.

Ernst, C.H.; Lovich, J.E.; Laemmerzahl, A.F.; Sekscienski, S. 1997. A comparison of plastral scute lengths among members of the box turtle genera Cuora and Terrapene. Chelonian Conservation and Biology 2(4): 603-607.

Esser, B.; Nickl, S. 2014. Beobachtungen im Habitat der Gelbrand-Scharnierschildkröte Coura flavomarginata (Gray, 1863) auf Taiwan un daraus resultierende Folgerungen für die Haltung in menschlicher Obhut. Radiata 23(3): 4-19.

Fong, J.J.; Parham, J.F.; Fu, J. 2002. A reassessment of the distribution of Cuora flavomarginata Gray 1863 on mainland China. Russian Journal of Herpetology 9(1): 9-14.

Fritzsche, D.; Marlies, F. 2005. Experiences with the Yellow-margined Box Turtle Cuora flavomarginata. Radiata 14(2): 50-51.

Gray, J.E. 1863. Observations on the box tortoises, with the descriptions of three new Asiatic species. Proceedings of the Zoological Society of London 1863: 173-179.

Hennen, U. 2000. Die Gelbrand-Scharnierschildkröte Cuora flavomarginata - eine interessante aus China. Radiata 9(3): 3-15.

Hennen, U. 2014. Die Gelbrand-Scharnierschildkröte (Cuora flavomarginata). Teil 1: Biologie und Haltung. Marginata 44: ?

Hennen, U. 2015. Die Gelbrand-Scharnierschildkröte (Cuora flavomarginata). Teil 2: Nachzucht im Terrarium und Gefährdung in der Natur Marginata 45: ?

Hsu, H.F. 1930. Preliminary note on a new variety of Cyclemys flavomarginata from China. Contr. Biol. Lab. Sci. Soc. China Nanking 6(1): 1-7.

Huang, B. 2008. Population characteristics to morphometric biology of Cistolemmys [CistoClemmys] flavomarginata Gray in Dabieshan region (II). Sichuan Journal of Zoology 27(5): 873-875.

Huang, B.; Feng, Z.G. 2011. [Morphological characteristics and subpopulations differentiation analysis of the Cistoclemmys flavomarginata]. (In Chinese). Xinyang Shifan Xueyuan Xuebao Ziran Kexue Ban 24(4) (Suppl. General Serial 105): 490-494.

Huang, B.; Jing, K.Y. 2010. [Research on characteristic of lay eggs and accumulated hatching temperature of Cistoclemmys flavomarginata in a domestication state]. (In Chinese). Xinyang Shifan Xueyuan Xuebao Ziran Kexue Ban 23(1) (General Serial no. 98): 79-83.

Huang, B.; Yang, J.H.; Zhao, W.P. 2004. [The breeding biology of CistoClemmys flavomarginata]. (In Chinese, English summary). Chinese Journal of Zoology 39(4): 40-44.

Huang, B.; Yang, Y.L.; Wang, R.P. 2012. [Studies on growth laws of juvenile Cistoclemmys flavomarginata]. (In Chinese). Xinyang Shifan Xueyuan Xuebao Ziran Kexue Ban 25(1)(General Serial no. 106): 58-63.

Klerks, M. 2006. Keeping and breeding the Chinese box turtle, Cuora flavomarginata (Gray, 1863). pp. 13-17. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Landzettel, F. 1999. Erfahrungen bei der Haltung und Vermehrung der Gelbrandscharnierschildkröte, Cuora flavomarginata. Elaphe 7(4): 13-16.

Lee, D. 2005. Le Consortium sur la tortue asiatique. Manouria 8(28): 17-23.

Lin, Y.F.; Wu, S.; Lin, T.E.; Mao, J.J.; Chen, T.H. 2010. Population status and distribution of the endangered Yellow-margined Box Turtle Cuora flavomarginata in Taiwan. Oryx 44(4): 581-587.

Lu, J. & al. 1999. Study on reproductive technic [technique] of Cistoclemmys flavomarginata. (In Chinese). Sichuan Journal of Zoology 18(2): 96-97.

Lue, K.Y.; Chen, T.H. 1999. Activity, movement patterns, and home range of the yellow-margined box turtle (Cuora flavomarginata) in northern Taiwan. Journal of Herpetology 33(4): 590-600.

McCoy, C.J.; Richmond, N.D. 1967. The identity of the Chinese box turtle, Terrapene culturalia. Copeia 1966: 886.

Natchev, N.; Heiss, E.; Lemell, P.; Stratev, D.; Weisgram, J. 2009. Analysis of prey capture and food transport kinematics in two Asian box turtles, Cuora amboinensis and Cuora flavomarginata (Chelonia, Geoemydidae), with emphasis on terrestrial feeding patterns. Zoology (Jena) 112(2): 113-127.

Nuss, C. 2007. Unerwarteter Nachzuchterfolg bei der Gelbrand-Scharnierschildkröte (Cuora flavomarginata). Radiata 16(2): 29-32.

Okada, S. 1891. Catalogue of vertebrated animals of Japan. Kinkodo, Tokyo, 128 pp.

Ota, H.; Yasukawa, Y.; Fu, J.; Chen, T.H. 2009. Cuora flavomarginata (Gray 1863) - Yellow-margined Box Turtle Chelonian Research Monographs 5: 035.1-035.10.

Pawlowski, S. 2005. Bericht zur erfolgreichen Haltung und Nachzucht der Gelbrand-Scharnierschildkröte, Cuora flavomarginata. Radiata 14(2): 55-59.

Pawlowski, S. 2005. Report on the successful maintenance and breeding of the Yellow-margined Box Turtle, Cuora flavomarginata. Radiata 14(2): 55-59.

Pawlowski, S. 2007. Haltungstipps und Haltungsfehler bei der Pflege terrestrischer Sumpfschildkröten am Beispiel der Gelbrand-Scharnierschildkröte Cuora flavomarginata spp. (Gray, 1863). Elaphe 15(4): 43-52.

Pawlowski, S.; Strmac, M. 2003. Bemerkungen zu Nachzucht der Gelbrandscharnierschildkröte, Cuora flavomarginata, im Zimmerterrarium. Elaphe 11(2): 27-32.

Pawlowski, S.; Strmac, M. 2006. Eine erfolgreiche Überwinterungsstrategie für Gelbrand-Scharnierschildkröten, Cuora flavomarginata ssp. (Gray, 1863). Sauria (Berlin) 28(1): 11-17.

Romero Gomez, M.; Valverde, J. 2009. Haltung und Nachzucht von CistoClemmys flavomarginata, der Gelbrand-Scharnierschildkröte. Reptilia (D) 13(74): 34-39.

Schaffer, G.; Felsner, H. 2000. Haltung und Navchzucht der Gelbrand-Scharnierschildkröte Cuora flavomarginata. pp. 111-116. In: Artner, H.; Meier, E. (eds.). Schildkröten. Grosses Schildkrötensymposium. 5. Jahrestagung der Schildkrötenfreunde Österreich. Salzburg, 16. bis 19. Jänner 1997. Natur und Tier-Verlag, Münster. 181 pp.

Tanaka, S.; Sato, F. 1983. Brief observation of the mating behaviour of the box turtle Cuora flavomarginata flavomarginata in nature. Biological Magazine Okinawa 21: 75-76.

Thevot, F. 2010. L'elevage de Cistoclemmys flavomarginata et Cuora mouhotii. Cheloniens 19: 35-40.

Yasukawa, Y.; Ota, H. 1999. Geographic variation and biogeography of the geoemydine turtles (Testudines: Bataguridae) of the Ryukyu Archipelago, Japan. pp. 271-297. In: Ota, H. (ed.). Tropical island herpetofauna: origin, current diversity, and conservation. Developments in Animal and Veterinary Sciences 29: 1-353.

Zhang, H. 1986. Study on the breeding of Cuora flavomarginate. Chinese Journal of Zoology 21(1): 11-13.

Zhang, Y.Y.; Nie, L.W.; Huang, Y.; Pu, Y.G.; Zhang, L. 2009. The mitochondrial DNA control region comparison studies of four hinged turtles and its phylogentic significance of the genus Cuora sensu lato (Testudinata: Geoemydidae). Genes & Genomics 31(5): 349-359.

Zhou, G.T.; Pan, F.L.; Wu, Z.L. 2003. [The comparison of growth performance among three species turtle juveniles: Chinemys reevesii, Mauremys mutica and CistoClemmys flavomarginatas]. (In Chinese, English summary). Fisheries Science (Liaoning) 22(1): 32-33.

Zwartepoorte, H. 1986. Keeping and breeding Cuora flavomarginata. Lacerta 44(5): 82-85.

Cuora galbinifrons

Anonymous. 1975. Three new species of reptiles from Hainan Island, Guangdong Province. Acta Zoologica Sinica 21(4): 379-384.

Anonymous. 1995. Another mystery turtle. Tropical Fish Hobbyist 43(6): 138-139.

Anonymous. 2014. [Simulation of Cuora galbinifrons nest predation]. (In Chinese). Shengtaixue Zazhi 33(6): 1629-1633.

Artner, H. 1998. Nomenklatur aktuell: Cuora amboinensis und Cuora galbinifrons. Emys 5(4): 34-35.

Augustine, L.; Mendyk, R.W. 2012. Natural history notes: Cuora galbinifrons galbinifrons (Indochinese Box Turtle). Post-copulatory sniffing behavior. Herpetological Review 43(2): 264-265.

Becker, H. 2013. Zwillingsanlage bei Cuora galbinifrons (Bourret, 1939). Radiata 22(2): 23-25.

Becker, H.; Philippen, H.D. 2008. Bemerkungen zur Innerartlichen Varibilität von Cuora galbinifrons. Marginata 17: 10-15.

Borek, A.; Miller, P.; Yoshimi, D.; Pramuk, J. 2018. Husbandry of the Indochinese Box Turtle (Cuora galbinifrons: Geoemydidae) at Woodland Park Zoo. Herpetological Review 49(2): 264-270.

Bour, R. 2005. The type specimens of Cuora galbifrons Bourret, 1939. Emys 12(1): 33-36.

Bourret, R. 1940. Notes herpétologiques sur l'Indochine francaise. XVIII. reptiles et batraciens recus au Laboratoire des Sciences Naturelle de l'Université au cours de l'année 1939. Descriptions de quatre espèces et d'une variété nouvelles. Bulletin General de l'Instruction Publique (Hanoi) 18(4) (annexe): 5-39.

Bruin, R. de 2006. Breeding the Indochinese Box Turtle, Cuora galbinifrons galbinifrons. pp. 69-76. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Buskirk, J.R. 1989. New locality records for Chinese non-marine chelonians. Chinese Herpetological Research 2(2): 65-68.

Ernst, C.H.; Lovich, J.E.; Laemmerzahl, A.F.; Sekscienski, S. 1997. A comparison of plastral scute lengths among members of the box turtle genera Cuora and Terrapene. Chelonian Conservation and Biology 2(4): 603-607.

Fritz, U.; Petzold, A.; Auer, M. 2006. Osteology in the Cuora galbinifrons complex suggests conspecifity of C. bourreti and C. galbinifrons, with notes on shell osteology and phalangeal formulae within the Geoemydidae. Amphibia-Reptilia 27(2): 195-205.

Fritz, U.; Ziegler, T.; Herrmann, H.W.; Lehr, E. 2002. Intergradation between subspecies of Cuora galbinifrons Bourret, 1939 and Pyxidea mouhotii (Gray, 1862) in southern North Vietnam (Reptilia: Testudines: Geoemydidae). Faunistische Abhandlungen (Dresden) 23(1): 59-74.

Fritzsche, D.; Marlies, F. 2005. Experiences with the Indochinese Box Turtle Cuora galbinifrons Bourret, 1939. Radiata 14(2): 48-49.

Götz, M. 2004. A brief report on the long-term rehabilitation of wild-caught Cuora galbinifrons. Testudo 6(1): 37-40.

Hiller, A. 2005. Beobachtungen bei der Haltung von Cuora galbinifrons galbinifrons Bourret, 1939. Radiata 14(2): 44-47.

Hiller, A. 2005. Notes on keeping Cuora galbinifrons galbinifrons Bourret, 1939. Radiata 14(2): 44-47.

Hiller, A.; Hiller, I. 2008. Beobachtungen an der Hinterindischen Scharnierschildkröte, Cuora galbinifrons galbinifrons. I. Marginata 17: 16-21.

Hiller, A.; Hiller, I. 2008. Beobachtungen an der Hinterindischen Scharnierschildkröte, Cuora galbinifrons galbinifrons. II. Marginata 18: 39-42.

Hu, S.; Zhao, E. 1982. Three new species of reptiles from Hainan Island, Guangdong Province. Smithsonian Herpetological Information Service 53: 1-9. (Translation of Acta Zoologica Sinica 24(4) [1975]: 379-384).

Hu, S.; Zhao, E.M. 1975. [Three new species of reptiles from Hainan Island, Guangdong Province (In Chinese). Acta Zoologica Sinica 24(4): 379-384.

Iverson, J.B.; McCord, W.P. 1992. A new subspecies of Cuora galbinifrons (Testudines: Batagurinae) from Hainan Island, China. Proceedings of the Biological Society of Washington 105(3): 433-439.

Lehr, E.; Fritz, U.; Obst, F.J. 1998. Die Unterarten von Cuora galbinifrons Bourret, 1939 (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen (Dresden) 50(1): 77-97.

Natchev, N.; Lemell, P.; Heiss, E.; Beisser, C.; Weisgram, J. 2010. Aquatic feeding in a terrestrial turtle: a functional-morphological study of the feeding apparatus in the Indochinese box turtle Cuora galbinifrons (Testudines, Geoemydidae). Zoomorphology (Berlin) 129(2): 111-119.

Nguyen, Q.T.; Doan, V.K.; Bain, R. 2005. Geographic distribution: Cuora galbinifrons (Indochinese Box Turtle). Herpetological Review 36(3): 335.

Obst, F.J.; Reimann, M. 1994. Bemerkenswerte Variabilität bei Cuora galbinifrons Bourret, 1939, mit Beschreibung einer neuen geographischen Unterart: Cuora galbinifrons bourreti subsp. nov. (Reptilia: Testudines: Cryptodira: Bataguridae). Zoologische Abhandlungen (Dresden) 48(1): 125-138.

Petzold, H.G. 1963. Über einige Schildkröten aus Nord-Vietnam im Tierpark Berlin. Senckenbergiana Biologica 44(1): 1-20.

Petzold, H.G. 1965. Cuora galbinifrons und andere südostasiatische Schildkröten im Tierpark Berlin. 1. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 18: 87-91.

Petzold, H.G. 1965. Cuora galbinifrons und andere südostasiatische Schildkröten im Tierpark Berlin. 2. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 18: 119-121.

Price, R. 1992. Vocalization by Cuora galbinifrons (Chinese flower box turtle). Bulletin of the Chicago Herpetological Society 27(7): 150.

Price, R. 1995. Interactive behavior in Cuora galbinifrons (Chinese flower box turtle). Bulletin of the Chicago Herpetological Society 30(9): 192-193.

Shi, H.; Parham, J.F.; Simison, W.B.; Wang, J.; Gong, S.; Fu, B. 2005. A report on the hybridization between two species of threatened Asian box turtles (Testudines: Cuora) in the wild on Hainan Island (China) with comments on the origin of 'serrata'-like turtles. Amphibia-Reptilia 26(3): 377-381.

Som, S.; Cottet, M. 2016. Rescue and relocation programme of turtles and tortoises and Elongated Tortoise monitoring programme in the Nam Theun 2 Reservoir (Laos). Hydroecologie Appliquee 19: 383-406.

Stuart, B.L.; Parham, J.F. 2004. Molecular phylogeny of the critically endangered Indochinese Box Turtle (Cuora galbinifrons). Molecular Phylogenetics and Evolution 31(1): 164-177.

Tscherepach, P. 1969. Ein sensationelles Bilddokument: Zwillingsbildung auch bei Landschildkröten. Aquarien Terrarien 16: 133.

Wang, J.; Shi, H.; Xue, C.Q.; Wang, L.; Zhao, E.M. 2011. [Population densities of Cuora galbinifrons at Diaoluoshan Nature Reserve, Hainan Island, China]. (In Chinese). Sichuan Journal of Zoology 30(3): 471-474.

Zhang, Y.Y.; Nie, L.W.; Huang, Y.; Pu, Y.G.; Zhang, L. 2009. The mitochondrial DNA control region comparison studies of four hinged turtles and its phylogentic significance of the genus Cuora sensu lato (Testudinata: Geoemydidae). Genes & Genomics 31(5): 349-359.

Cuora mccordi

Artner, H. 2006. Captive breeding of McCord's box turtle Cuora mccordi Ernst, 1988. pp. 77-90. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Ernst, C.H. 1988. Cuora mccordi, a new Chinese box turtle from Guangxi Province. Proceedings of the Biological Society of Washington 101(2): 466-470.

Hertwig, S. 2005. Husbandry and breeding of Cuora mccordi Ernst, 1988. Radiata 14(2): 37--37.

Parham, J.F.; Stuart, B.L.; Bour, R.; Fritz, U. 2004. Evolutionary distinctiveness of the extinct Yunnan box turtle (Cuora yunnanensis) revealed by DNA from an old museum specimen. Proceedings of the Royal Society Biological Sciences Series B 271: S391-S394. (Suppl 6).

Raffel, M.; Zwartepoorte, H. 2009. European turtle breeding reports 2008/2009. TSA Newsletter 2009 (August): 28-29.

Schroller, V. 2005. Husbandry and breeding of McCord's Hinged Turtle, Cuora mccordi Ernst, 1988. Radiata 14(2): 39-40.

Zhou, T.; Blanck, T.; McCord, W.P.; Li, P.P. 2008. Tracking Cuora mccordi Ernst, 1988: the first record of its natural habitat: a re-description; with data on captive populations and its vulnerability. Hamadryad 32(1): 46-58.

Cuora mouhotii

Anonymous. 1995. Another mystery turtle. Tropical Fish Hobbyist 43(6): 138-139.

Auer, M. 2011. Beobachtungen zu Vorkommen and Handel von Schildkröten in Nordlaos. Sauria (Berlin) 33(1): 3-11.

Becker, H. 2010. Bemerkungen zur Zucht von Pyxidea mouhoutii mouhotii und Pyxidea mouhotii obsti, sowie Beobachtungen zu Plastronfärbungen der Jungtiere. Marginata 27: 48-55.

Becker, H. 2012. Bemerkungen zum Erreichen der Geschlechtsreife einer weiblichen Indischen Dornschildkröte Pyxidea mouhotii mouhotii (Gray, 1862). Testudo (SIGS) 21(4): 18-24.

Choudhury, A. 1993. Keeled box turtle in Karbi Anglong - a new locality record. Journal of the Bombay Natural History Society 90(3): 517.

Choudhury, A. 1996. The keeled box turtle Pyxidea mouhotii Gray - a new record for Manipur. Journal of the Bombay Natural History Society 93(3): 590-591.

Choudhury, A. 1996. The keeled box turtle Pyxidea mouhotii Gray on the north bank of the Brahmaputra - a new record. Journal of the Bombay Natural History Society 93(1): 97.

Choudhury, A. 1998. Pyxidea mouhotii (Gray) in southern Assam and Mizoram. Journal of the Bombay Natural History Society 95(3): 511-512.

Choudhury, A. 2001. Further records of Pyxidea mouhotii (Gray) from the north bank of R. Brahmaputra, Assam. Journal of the Bombay Natural History Society 98(1): 116-117.

Choudhury, A. 2001. Some chelonian records from Mizoram. Journal of the Bombay Natural History Society 98(2): 184-190.

Choudhury, A. 2005. An interesting specimen of Pyxidea mouhotii (Gray) from Manipur in northeastern India. Reptile RAP 7: 11.

Das, I. 1987. Distribution of the heeled box turtle Pyxidea mouhotii (Gray). Journal of the Bombay Natural History Society 84(1): 221-222.

Das, I.; McCormack, T.E.M.; Dijk, P.P. van; Hoang, H.V.; Struijk, R.P.J.H. 2016. Cuora mouhotii (Gray 1862) – Keeled Box Turtle. Chelonian Research Monographs 5(99): 1-12.

Farkas, B.; Sasvári, L. 1992. On freshwater turtles (Testudines) new to China. pp. 151-157. In: Korsos, Z. & Kiss, I. (eds.): Proceedings of the 6th Ordinary General Meeting of the Societas Europaea Herpetologica, 19-23 August 1991, Budapest, Hungary. Hungarian Natural History Museum, Budapest. 531 pp.

Fritz, U.; Andreas, B.; Lehr, E. 1998. Eine neue Unterart der Dreikiel-Scharnierschildkröte, Pyxidea mouhotii (Gray, 1862) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen (Dresden) 50(1): 33-43.

Fritz, U.; Obst, F.J. 1997. Zum taxonomischen Status von Cuora galbinifrons serrata Iverson & McCord, 1992 und Pyxidea mouhotii (Gray, 1862) (Reptilia: Testudines: Bataguridae). Zoologische Abhandlungen (Dresden) 49(2): 261-279.

Fritz, U.; Ziegler, T.; Herrmann, H.W.; Lehr, E. 2002. Intergradation between subspecies of Cuora galbinifrons Bourret, 1939 and Pyxidea mouhotii (Gray, 1862) in southern North Vietnam (Reptilia: Testudines: Geoemydidae). Faunistische Abhandlungen (Dresden) 23(1): 59-74.

Hong, M.L.; Shi, H.; Fu, L.R.; Gong, S.; Fong, J.J.; Parham, J.F. 2008. Scientific refutation of traditional Chinese medicine claims about turtles. Applied Herpetology 5(2): 173-187.

Iverson, J.B.; McCord, W.P. 1992. A new subspecies of Cuora galbinifrons (Testudines: Batagurinae) from Hainan Island, China. Proceedings of the Biological Society of Washington 105(3): 433-439.

Ly, T.; Hoang, H.D.; Stuart, B.L. 2013. Occurrence of the endangered Keeled Box Turtle, Cuora mouhotii, in southern Vietnam. Chelonian Conservation and Biology 12(1): 184-187.

Petzold, H.G. 1963. Über einige Schildkröten aus Nord-Vietnam im Tierpark Berlin. Senckenbergiana Biologica 44(1): 1-20.

Platt, S.G.; Platt, K.; Myo, K.M.; Moe, K.; Soe, M.M.; Naing, T.Z.; Lin, N.; Rainwater, T.R. 2013. Noteworthy records of chelonians from the Chindwin River Basin and Naga Hills of western Myanmar. Herpetological Conservation and Biology 8(2): 335-350.

Pointet, A. 1997. La tortue a trois carenes, Pyxidea mouhotii (Gray 1862). CITS Bulletin 9: 26-40.

Sachsse, W. 1973. Pyxidea mouhotii, eine landbewohnende Emydide Südostasiens (Testudines). Salamandra 9(2): 49-53.

Sachu, K.; Purkayastha, J.; Bhattacharjee, R. 2017. Geographic distribution: Cuora mouhotii (Keeled Box Turtle). Herpetological Review 48(1): 122.

Shi, H.; Parham, J.F.; Simison, W.B.; Wang, J.; Gong, S.; Fu, B. 2005. A report on the hybridization between two species of threatened Asian box turtles (Testudines: Cuora) in the wild on Hainan Island (China) with comments on the origin of 'serrata'-like turtles. Amphibia-Reptilia 26(3): 377-381.

Singh, R.R.; Singh, K.J. 2012. Box turtles in and adjacent to Loktak Lake, Manipur - India. Taprobanica 4(2): 103-104.

Struijk, R.; Blanck, T.E.G. 2016. A likely new natural hybrid form of “Cuora serrata” (Cuora picturata x Cuora mouhotii obsti) and its finding in the wild in Phu Yen province, Vietnam. Herpetology Notes 9: 73-80.

Struijk, R.P.J.H.; Blanck, T.E.G. 2015. Herpetoculture notes: Cuora mouhotii mouhotii (Northern Keeled Box Turtle) and C. bouretti (Bourret's Box Turtle). Captive hybridization. Herpetological Review 46(2): 206.

Struijk, R.P.J.H.; McCormack, T.E.M.; Nguyen, T.T.; Pham, T.V.; Stumpel, J.B.G.; WANG, J.; Auer, M. 2016. Zusammenfassung Intergradation zwischen Cuora mouhotii mouhotii (Gray, 1862) und Cuora mouhotii obsti Fritz, Andreas & Lehr, 1998 mit Bemerkungen zur geografischen Verbreitung der Art und ihrer phänotypischen Variation. Sauria (Berlin) 38(1): 31-47.

Thevot, F. 2010. L'elevage de Cistoclemmys flavomarginata et Cuora mouhotii. Cheloniens 19: 35-40.

Wang, J.; Gong, S.P.; Shi, H.T.; Liu, Y.X.; Zhao, E.M. 2011. Reproduction and nesting of the endangered Keeled Box Turtle (Cuora mouhotii) on Hainan Island, China. Chelonian Conservation and Biology 10(2): 159-164.

Wangyal, J.T.; Wangchuk, D.; Das, I. 2012. First report of turtles from the Himalayan Kingdom of Bhutan. Chelonian Conservation and Biology 11(2): 268-272.

Zhang, L.; Nie, L.W.; Cao, C.H.; Zhang, Y. 2008. The complete mitochondrial genome of the Keeled box turtle Pyxidea mouhotii and phylogenetic analysis of major turtle groups. Journal of Genetics and Genomics 35(1): 33-40.

Zhang, Y.Y.; Nie, L.W.; Huang, Y.; Pu, Y.G.; Zhang, L. 2009. The mitochondrial DNA control region comparison studies of four hinged turtles and its phylogentic significance of the genus Cuora sensu lato (Testudinata: Geoemydidae). Genes & Genomics 31(5): 349-359.

Cuora pani

Artner, H. 1999. Erstnachzucht von Pan's Scharnierschildkröte Cuora pani Song, 1984 in menschlicher Obhut. Emys 6(5): 4-20.

Artner, H. 2004. Haltung und Nachzucht von Pan's Scharnierschildkröte Cuora pani pani Song, 1984 und der Goldkopf-Scharnierschildkröte Cuora pani aurocapitata Luo & Zong, 1988. Emys 11(1): 4-21.

Artner, H. 2006. Elevage et reproduction de la tortue-boite de Pan, Cuora pani pani Song, 1984 et de la tortue-boite a tete jaune, Cuora pani aurocapitata Luo & Zong, 1988. Manouria 9(32): 27-39.

Artner, H. 2006. Keeping and breeding of Pan's box turtle Cuora pani pani Song, 1984 and Cuora pani aurocapitata Luo & Zong, 1988. pp. 52-68. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Buskirk, J.R. 1989. New locality records for Chinese non-marine chelonians. Chinese Herpetological Research 2(2): 65-68.

Ernst, C.H. 1988. Redescriptions of two Chinese Cuora (Reptilia: Testudines: Emydidae). Proceedings of the Biological Society of Washington 101(1): 155-161.

Ernst, C.H.; McCord, W.P. 1987. Two new turtles from southeast Asia. Proceedings of the Biological Society of Washington 100(3): 624-628.

Hertwig, S. 2005. Husbandry and breeding of Cuora pani Song, 1984. Radiata 14(2): 43.

Huang, B.; Huang, Y. 2011. [A new record of distribution in Henan Province: Cuora pani Song]. (In Chinese). Xinyang Shifan Xueyuan Xuebao Ziran Kexue Ban 24(1) (General Serial no. 102): 71-72.

Li, D. 2000. [Cuora pani - first report from Sichuan Province]. (In Chinese). Sichuan Journal of Zoology 19(3): 156.

Parham, J.F.; Li, D. 1999. A new locality for Cuora pani Song 1984 with comments on its known range. Asiatic Herpetological Research 8: 111-113.

Song, M.T. 1984. A new species of the turtle genus Cuora (Testudoformes[Testudiformes]: Testudinidae). Acta Zootaxonomica Sinica 9(3): 330-332.

Song, M.T. 2001. A new species of the turtle genus Cuora (Testudoformes: Testudinidae). [Translation of: Acta Zootaxonomica Sinica 9(3) 1984: 330-332]. Asiatic Herpetological Research 9 [2000]: 142-144.

Theiler, M.R. 2005. Notes on the husbandry and breeding of Cuora pani Song, 1984. Radiata 14(2): 41-42.

Cuora picturata

Lehr, E.; Fritz, U.; Obst, F.J. 1998. Cuora galbinifrons picturata subsp. nov., eine neue Unterart der Hinterindischen Scharnierschildkröte. Herpetofauna (Weinstadt) 20(113): 5-11.

Ly, T.; Hoang, H.D.; Stuart, B.L. 2011. Market turtle mystery solved in Vietnam. Biological Conservation 144(5): 1767-1771.

Niessen, L. 2008. Haltung und Nachzucht der Südvietnamesischen Scharnierschildkröte, Cuora galbinifrons picturata. Marginata 17: 28-30.

Struijk, R.; Blanck, T.E.G. 2016. A likely new natural hybrid form of “Cuora serrata” (Cuora picturata x Cuora mouhotii obsti) and its finding in the wild in Phu Yen province, Vietnam. Herpetology Notes 9: 73-80.

Cuora trifasciata

Blanck, T.; McCord, W.P.; Le, M. 2006. On the variability of Cuora trifasciata (Bell, 1825). Chimaira, Frankfurt a.M. 155 pp.

Blanck, T.; Ting, Z.; Protiva, T.; Crow, P.; Tiedemann, R. 2017. New subspecies of Cuora cyclornata (Blanck, McCord & Le, 2006), Cuora trifasciata (Bell, 1825) and Cuora aurocapitata (Luo & Zong, 1988). Sichuan Journal of Zoology 36(4): 368-385.

Chen, C.S.; Wei, K.; Liu, K.; Geng, Y.; Xu, Z.H.; Gao, Y.L. 2011. Study on feeding effect of Cuora trifasciata by different diet in artificial conditions. Sichuan Journal of Zoology 30(4): 586-589.

Cheung, S.M. 2003. Chinese three-striped box turtle Cuora trifasciata in Hong Kong: possibly the last viable wild population in the world. Porcupine 28: 9.

Coutard, C.; Trani, M.; Tafani, J.S.; Cadi, A. 2004. Liste rouge TSA: Especes menacees elevees au centre A Cupulatta (1ere partie). Manouria 7(23): 37-44.

Dijk, P.P. van 2005. Die Dreistreifen-Scharnierschildkröte - ein Opfer des vermeintlichen Nutzens. Schildkröten im Fokus 2(1): 13-16.

Harding, J.H. 1989. Courtship behavior in the Chinese box turtle, Cuora trifasciata. Herpetological Review 20(4): 85.

Herz, M. 2005. An albino Cuora trifasciata (Bell, 1825). Radiata 14(2): 35-36.

Hong, M.L.; Shi, H.; Fu, L.R.; Gong, S.; Fong, J.J.; Parham, J.F. 2008. Scientific refutation of traditional Chinese medicine claims about turtles. Applied Herpetology 5(2): 173-187.

Iverson, J.B.; McCord, W.P. 1992. A new Chinese eyed turtle of the genus Sacalia (Batagurinae: Testudines). Proceedings of the Biological Society of Washington 105(3): 426-432.

Li, G.S.; Tang, D. 2000. [Cuora trifasciata, its artificial protection and reproduction]. (In Chinese, English summary). Sichuan Journal of Zoology 19(3): 149-150.

Li, W.; Zhang, X.C.; Zhao, J.; Shi, Y.; Zhu, X.P. 2015. Complete mitochondrial genome of Cuora trifasciata (Chinese Three-striped Box Turtle), and a comparative analysis with other box turtles. Gene (Amsterdam) 555(2): 169-177.

Lovich, J.E. 1989. Another exotic turtle record for Hawaii. Elepaio 49(12): 86-87.

McCord, W.P.; Iverson, J.B. 1992. A new species of Ocadia (Testudines: Bataguridae) from Hainan Island, China. Proceedings of the Biological Society of Washington 105(1): 13-18.

Meier, E. 2000. Eine methode zur Zucht aggressiver und stressempfindlicher Wasserschildkröten, exemplarisch dargestellt an der Moorschildkröte Clemmys muhlenbergii (Schöpff, 1801) und Dreistreifen-Scharnierschildkröte Cuora trifasciata (Bell, 1825). pp. 53-68. In: Artner, H.; Meier, E. (eds.). Schildkröten. Grosses Schildkrötensymposium. 5. Jahrestagung der Schildkrötenfreunde Österreich. Salzburg, 16. bis 19. Jänner 1997. Natur und Tier-Verlag, Münster. 181 pp.

Parham, J.F.; Shi, H. 2001. The discovery of Mauremys iversoni-like turtles at a turtle farm in Hainan Province, China: the counterfeit golden coin. Asiatic Herpetological Research 9 [2000]: 71-76.

Petzold, H.G. 1963. Über einige Schildkröten aus Nord-Vietnam im Tierpark Berlin. Senckenbergiana Biologica 44(1): 1-20.

Pritchard, P.C.H.; McCord, W.P. 1991. A new emydid turtle from China. Herpetologica 47(2): 139-147.

Raffel, M.; Zwartepoorte, H. 2009. European turtle breeding reports 2008/2009. TSA Newsletter 2009 (August): 28-29.

Shi, H. 2006. The fate of a wild-caught golden coin turtle (Cuora trifasciata) on Hainan Island, China. Turtle and Tortoise Newsletter 9: 15-17.

Spinks, P.Q.; Thomson, R.C.; Shaffer, H.B. 2009. A reassessment of Cuora cyclornata Blanck, McCord and Le, 2006 (Testudines, Geoemydidae) and a plea for taxonomic stability. Zootaxa 2018: 56-68.

Timmins, R.J.; Khounboline, K. 1999. Occurrence and trade of the golden turtle, Cuora trifasciata, in Laos. Chelonian Conservation and Biology 3(3): 441-447.

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Zhou, G. 1998. Relationship between activity of the Cuora trifasciata and intensity of illumination. (In Chinese, English summary). Acta Scientiarum Naturalium Universitatis Normalis Hunanensis 21(1): 93-96.

Zhou, T. 2000. [How to feed Cuora trifasciata]. (In Chinese). Sichuan Journal of Zoology 19(3): 184.

Zhu, X.P.; Chen, Y.L.; Liu, Y.H.; Wei, C.Q.; Lu, X.D. 2001. [Comparative study of the growth of Mauremys mutica, Cuora trifasciata and Chelydra serpentina]. (In Chinese, English summary). Journal of Fisheries of China 25(6): 507-511.

Zwartepoorte, H. 2008. The golden coin box turtle status in EAZA institutions. TSA Newsletter 2008 (August): 25.

Cuora yunnanensis

Blanck, T. 2005. Cuora yunnanensis (Boulenger, 1906), die Yunnan-Scharnierschildkröte. Nach 100 Jahren wiederentdeckt. Radiata 14(2): 10-33.

Ernst, C.H. 1988. Redescriptions of two Chinese Cuora (Reptilia: Testudines: Emydidae). Proceedings of the Biological Society of Washington 101(1): 155-161.

He, J.; Zhou, T.; Rao, D.Q.; Zhang, Y.P. 2007. Molecular identification and phylogenetic position of Cuora yunnanensis. Chinese Science Bulletin 52(23): 3305-3309.

Parham, J.F.; Stuart, B.L.; Bour, R.; Fritz, U. 2004. Evolutionary distinctiveness of the extinct Yunnan box turtle (Cuora yunnanensis) revealed by DNA from an old museum specimen. Proceedings of the Royal Society Biological Sciences Series B 271: S391-S394. (Suppl 6).

Zhou, T. 2005. [Discovery of a living male Yunnan box turtle, Cuora yunnanensis Boulerger, 1906]. (In Chinese, English summary). Sichuan Journal of Zoology 24(3): 345-346.

Zhou, T.; McCord, W.P.; Blanck, T.; Sun, X.F. 2008. Rediscovery and first breeding of the Yunnan box turtle, Cuora yunnanensis. Reptilia (GB) 59: 29-34.

Zhou, T.; Zhao, E.M. 2004. [On the occurrence of living Cuora yunnanensis since fifty-eight years, and its description]. (In Chinese, English summary). Sichuan Journal of Zoology 23(4): 325-326.

Cuora zhoui

Blanck, T. 2013. The turtle from nowhere: an Asian box turtle on the brink of extinction. The Tortoise 1(2): 64-69.

Lee, D.S. 2011. Ponte de la tortue-boite du guangxi, cuora zhoui, en captivité. Cheloniens 24: 4-7.

Lee, D.S.; MacGregor, L.C. 2008. Haltung und Nachzucht von Zhous Scharnierschildkröte, Curoa zhoui Zhao, 1990. Radiata 17(2): 19-27.

McCord, W.P.; Iverson, J.B. 1991. A new box turtle of the genus Cuora (Testudines: Emydidae) with taxonomic notes and a key to the species. Herpetologica 47(4): 407-420.

Meier, E. 2000. Eine Arche für die letzten ihrer Art. Die geplante Erhaltungszuchtstation im Allwetterzoo Münster. Reptilia (D) 5(22): 34-39.

Meier, E. 2005. Remarks on Zhou's Box Turtle, Cuora zhoui Zhao, 1990. Radiata 14(4): 34.

Parham, J.F.; Stuart, B.L.; Bour, R.; Fritz, U. 2004. Evolutionary distinctiveness of the extinct Yunnan box turtle (Cuora yunnanensis) revealed by DNA from an old museum specimen. Proceedings of the Royal Society Biological Sciences Series B 271: S391-S394. (Suppl 6).

Wang, Y. 2008. [Artificial propagation of Cuora zhoui]. (In Chinese). Sichuan Journal of Zoology 27(1): 25.

Zhou, T. 2006. [Observation of Cuora zhoui in captivity]. (In Chinese, English summary). Sichuan Journal of Zoology 25(2): 390-392.