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Taxonomy of the genus Geochelone Biodiversity of the family Testudinidae

Bibliography of the genus
Geochelone (South Asian Star Tortoises)

(Reptilia: Testudines: Testudinidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Geochelone in general

Crumly, C.R. 1982. A cladistic analysis of Geochelone using cranial osteology. Journal of Herpetology 16(3): 215-234.

Schleich, H.H. 1984. Brief abstract on wildlife observations on tortoises (Geochelone, Testudo) and their significance for the paleochelonology. Studia Geologica Salmanticensia Volumen Especial 1: 235-238.

Geochelone elegans

Ahamad, R.; Dharmaretnam, M.; Silva, A. de 2004. Preliminary observations on the star tortoise (Geochelone elegans) in Batticaloa district, Sri Lanka. Lyriocephalus 5(1-2): 135-139. (Special issue).

Alagar-Rajan, S.; Balasubramanian, P. 1992. Some food plants of the star tortoise Geochelone elegans at Point Calimere Wildlife Sanctuary, Tamil Nadu. Journal of the Bombay Natural History Society 88(2) [1991]: 290.

Anderson, J.A.; Minton, S.A. 1963. Two noteworthy herpetological records from the Thar Parkar Desert, West Pakistan. Herpetologica 19: 152.

Babu, N.B.; Stengel, C.J. 2011. What seizures can tell us about the Indian star tortoise trade. Traffic Bulletin 23(3): 113-116.

Bhupathy, S.; Silori, C.S.; Sunderraj, S.F.W. 1994. Additional locality records for two Indian tortoise species. Journal of the Bombay Natural History Society 91(1): 149-150.

Bianchini, M. 2011. Maintenance et reproduction de la tortue etoilee indienne Geochelone elegans. Cheloniens 21: 22-27.

Bidmon, H.J. 2002. Die Indische Sternschildkröte Geochelone elegans (Schöpff, 1795). Eine problematische tropische Landschildkröte. Mehrjährige Haltung und Vermehrung in Deutschland. 1. Aussehen, Verbreitung und Lokalrassen, Eingewöhnung, Vorstellung der Zuchtgruppe. Reptilia (D) 6(32): 56-64.

Bidmon, H.J. 2002. Die Indische Sternschildkröte Geochelone elegans (Schöpff, 1795). Eine problematische tropische Landschildkröte? Mehrjährige Haltung und Vermehrung in Deutschland. 2. Haltung. Reptilia (D) 7(33): 56-63.

Bidmon, H.J. 2002. Die Indische Sternschildkröte Geochelone elegans (Schöpff, 1795). Eine problematische tropische Landschildkröte? Mehrjährige Haltung und Vermehrung in Deutschland. 3. Nachzucht. Reptilia (D) 7(34): 56-62.

Bidmon, H.J. 2002. Die Indische Sternschildkröte Geochelone elegans (Schöpff, 1795). Eine problematische tropische Landschildkröte? Mehrjährige Haltung und Vermehrung in Deutschland. 4. Schlupf, Aufzucht und Krankheiten. Reptilia (D) 7(35): 56-63.

Bidmon, H.J. 2004. Laub und Tropische Landschildkröten: Eine Beobachtung. Schildkröten im Fokus 1(3): 11-16.

Bidmon, H.J. 2006. Geochelone elegans (Schöpff, 1795), the Indian star tortoise: is it really a "difficult" tropical tortoise? several years of indoor and outdoor husbandry and breeding in Germany. pp. 18-49. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Biswas, S.; Acharjyo, L.N. 1985. Observations on Geochelone elegans (Schöpff) in captivity, Orissa, India. Journal of the Bombay Natural History Society 81(3) [1984]: 707-708.

D'Cruze, N.; Mookerjee, A.; Vyas, R.; MacDonald, D.W.; Silva, A. de 2018. Geochelone elegans (Schoepff 1795) – Indian Star Tortoise, Star Tortoise. Chelonian Research Monographs 5(106): 1-13.

Dick, A. 1998. Geochelone elegans. Pflege und Nachzucht der indischen Sternschildkröte. Reptilia (D) 3(10): 48-50.

Dowling, H.G. 1961. Vanishing giants and enduring dwarfs - the tortoises. Animal Kingdom 64: 66-75.

Eglis, A. 1960. An aberrant tortoise from Ceylon. Herpetologica 16: 14.

Ekanayake, H.M.A.R.; Weerakoon, C.K.; Warusamanna, N.D.; Kularatna, W.J.J.; Nathanael, S.; Samarakoon, S.M.M.; Goonasekera, M.M.; Silva, A. de 2004. Preliminary observations on the status of captive star tortoises (Geochelone elegans) in Sri Lanka. Lyriocephalus 5(1-2): 65-72. (Special issue).

Fife, J.D. 2007. Star Tortoises - The natural History, Caprive care, and Breeding of Geochelone elegans and Geochelone platynota. Living Art Publishing, Ada, Oklahoma. 116 pp.

Frazier, J. 1989. Report on the chelonians of Gir Wildlife Sanctuary. Hamadryad 14(1): 3-5.

Frazier, J. 1989. Report on the star tortoises in the Sakkarbagh Zoo, Junagadh. Hamadryad 14(1): 5-8.

Frazier, J.G. 1990. Report on the star tortoise in the Sakkarbagh Zoo, Junagadh. Herpeton (Pune) 3: 7-8.

Gadhvi, I.R. 2001. Indian star tortoise (Geochelone elegans) feeding on slug. Zoos' Print Journal 16(7): 548.

Gaertner, K. 2001. Zur Haltung und Vermehrung der Indischen Sternschildkröte. Reptilia (D) 6(27): 64-68.

Gaur, A.; Reddy, A.; Annapoorni, S.; Satyarebala, B.; Shivaji, S. 2006. The origin of Indian star tortoises (Geochelone elegans) based on nuclear and mitochondrial DNA analysis: a story of rescue and repatriation. Conservation Genetics 7(2): 231-240.

Guyot, G.; Ashton, R. 2000. Elevage de la tortue rayonnee d'Inde, Geochelone elegans. Manouria 3(9): 12-19.

Ivanchev, I.E. 2012. [Indian Star Tortoise (Geochelone elegans) - several years of husbandry and breeding experience in Bulgaria]. (In German). Schildkröten im Fokus 9(3): 3-20.

Janssen, M.; Janssen, J. 2009. Beobachtungen und Erfahrungen bei der Haltung und Nachzucht der Indischen Sternschildkröte, Geochelone elegans (Schöpff, 1795). Radiata 18(4): 2-9.

Jayakar, S.D.; Spurway, H. 1964. Bimodality of laying-hatching times in Testudo elegans Schrepff (Chelonia). Nature (London) 204: 603.

Jayakar, S.D.; Spurway, H. 1966. Contribution to the biology of the Indian starred tortoise Testudo elegans Schöpff. Journal of the Bombay Natural History Society 63: 83-114.

Jayawardena, J. 1974. The starred tortoise (Testudo elegans). Loris 13(3): 136-137, 143.

Jayawardena, J. 1997. Record of a large Geochelone elegans (Schöpff, 1795) from Sri Lanka. Lyriocephalus 3(1): 34-35.

Jayawickrama, D.L.D.; Nathanael, E.S.; Silva, A. de 2010. Observations on Geochelone elegans (Reptilia: Testudinidae) (star tortoise) in the Mihintale Sanctuary, Sri Lanka. Lyriocephalus 7(1-2): 73-79.

Jayson, E.A. 1993. Starred tortoise Geochelone elegans (Schöpff) in Chinnar Wildlife Sanctuary, Kerala. Journal of the Bombay Natural History Society 90(1): 112.

Kirsche, W. 1976. Beitrag zur Biologie der Sternschildkröte (Testudo elegans Schöpff). Zoologische Garten (Jena) 46(1-2): 66-81.

Lafebre, S.; Martelli, P.; Murray, J. 2004. Management of confiscated Indian star tortoises at the Singapore Zoological Gardens. Turtle and Tortoise Newsletter 7: 11-12.

Matz, G.; Stemmler, O.; Vanderhaege, M. 1971. Sur quelques tortues etoilees. Aquarama 5(13): 27-31.

McCann, C. 1937. Sexual dimorphism in the star-shelled tortoise (Testudo elegans). Journal of the Bombay Natural History Society 39: 870-871.

Meyer, J.R.; Moore, C.W.; Page, C.D. 1990. Star tortoise (Geochelone elegans) captive propagation program at the Jacksonville Zoological Park. AAZPA Regional Conference Proceedings 1990: 382-389.

Moll, E.O. 1989. Geochelone elegans - Indian Star Tortoise. Occasional Papers of the IUCN Species Survival Commission 5: 113-114.

Narayanan, A. 2016. Variation in allometric shell growth and relationship with sexual size dimorphism in repatriated Indian Star Tortoises (Geochelone elegans) (Reptilia: Testudinidae). Current Science (Bangalore) 110(2): 206-213.

Nietzke, G. 1974. Testudo elegans - Sternschildkröte. Aquarium Aqua Terra 8(56): 75-76.

Noureen, U. 2014. The star of Pakistan. The Tortoise 1(3): 38-45.

Pawlowski, S. 2009. Im Habitat der Indischen Sternschildkröte, Geochelone elegans (Schöpff, 1975) auf Sri Lanka. Radiata 18(4): 12-22.

Peters, K. 2003. Kurze Bemerkungen zum Alter der Indischen Sternschildkröte Geochelone elegans (Schöpff, 1795). Sauria (Berlin) 25(4): 23.

Pillay, R.S.N. 1930. Notes on the star tortoise. Journal of the Bombay Natural History Society 34: 828.

Prakash, I. 1971. Testudo elegans in western Rajasthan. Journal of the Bombay Natural History Society 68: 273-274.

Rao, K.T.; Rao, M.V.S. 1991. Incubation and hatching of the Indian star tortoise Geochelone elegans in captivity. Journal of the Bombay Natural History Society 87(3) [1990]: 461-462.

Rao, R.J. 2005. Captive breeding of star tortoise (Geochelone elegans) in the Deer Park, NFCL, Kakinada, Andhra Pradesh. Reptile RAP 7: 7-9.

Reynolds, R.P. 1982. Programa de herpetologia. Estacion Cientifica Charles Darwin Informe Anual 1981: 51-55.

Reyst, A.T. 1948. Testudo elegans, Schöpff. Lacerta 6: 69-70.

Sanz, A.; Valverde, F.J. 1999. Keeping and breeding the Indian star tortoise. Reptilia (GB) 9: 43-49.

Saus, A. 2007. Haltung und Nachzucht der Sternschildkröte Geochelone elegans (Schöpff, 1794) aus Sri Lanka. Draco 8(2) (30): 50-55.

Saxena, M.M. 2003. A note on conditioning in starred tortoise Geochelone elegans (Schöpff) for yellow coloured food. Cobra (Chennai) 54: 23.

Scholdei, J. 2009. Erst beschlagnahmt, dann vermehrt: Sternschildkröten (Geochelone elegans) im Allwetterzoo Münster. Reptilia (D) 14(77): 59-62.

Scholdei, J. 2009. Zwillingsschlupf bei Sternschildkröten Geochelone elegans, im Allwetterzoo Münster. Sauria (Berlin) 31(4): 45-47.

Scholdei, J. 2011. Ein nicht alltäglicher Doppelschlupf von Zwillingen bei der Sternschildroete (Geochelone elegans) im Allwetterzoo Münster - weitere Daten über die Gewichts- und Wachstumsentwicklung. Marginata 29: 56-58.

Schwartz, S.R. 1968. Geochelone elegans. Bulletin of the Philadelphia Herpetological Society 16: 31-32.

Schwertner, P. 1979. Intelligenz und Lernfähigkeit bei Reptilien. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 32(7): 245-247.

Sekhar, A.C.; Gurunathan, N.; Anandhan, G. 2004. Star tortoise - a victim of the exotic pet trade. Tigerpaper (Bangkok) 31(1): 4-6.

Sharma, S.K. 2004. Forest fires and slow-moving reptiles. Cobra (Chennai) 55: 18.

Sharma, V.; Sharma, N.; Kishor, J.; Lal, J.B.; Dutt, U.; Kumar, S.K.; Meena, D. 2011. Occurrence, distribution and recent threats to Indian star tortoise Geochelone elegans (Schöpff, 1795) in the central Aravalli Mountain ranges, Rajasthan, India. Cobra (Chennai) 5(1): 20-24.

Shepherd, C.R.; Burgess, E.A.; Loo, M. 2004. Demand driven: the trade of Indian star tortoises Geochelone elegans in Peninsular Malaysia. TRAFFIC Southeast Asia, Petaling Jaya. 11 pp.

Silva, A. de 1995. The status of Geochelone elegans in North Western Province of Sri Lanka: preliminary findings. pp. 47-49. In: Devaux, B. (ed.). International Congress of Chelonian Conservation. France - Gonfaron - Tortoise Village - 6th to 10th of July 1995. Proceedings. Editions SOPTOM, Gonfaron, France. 344 pp.

Silva, A. de 2003. The biology and status of the star tortoise (Geochelone elegans Schöpff, 1795) in Sri Lanka. Published by the author, place of publication unknown. 100 pp.

Silva, A. de; Vamberger, M. 2016. Observations of Geochelone elegans (Schoepff, 1795) on an island off Kalpitiya Lagoon, Sri Lanka. Radiata (English edition) 25(1): 23-24.

Silva, A. de; Wijeratne, R.; Rodrigo, K.; Hettiarachchi, H.A.H.R.; Prasad, T. 2017. Record of possibly the world's largest star tortoise (Geochelone elegans) from Lunugamwehera National Park, Sri Lanka. Radiata (English Edition) 26(3): 30-34

Soman, P.W. 1991. Cannibalism in the star tortoise Geochelone elegans. Journal of the Bombay Natural History Society 88(1): 118.

Stemmler-Gyger, O. 1964. Zwei bemerkenswerte Exemplare der Sternschildkröte. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 17: 116-118.

Sunita, P.; Sekhar, P.S.R.; Subba Rao, M.V. 2008. Distribution status and breeding biology of Indian starred tortoise (Geochelone elegans) and flap-shell turtle (Lissemys punctata) in north coastal region of Andhra Pradesh. Journal of Nature Conservation 20(2): 245-256.

Tamilarasan, P. 1994. Studies on courtship and nesting behaviour of starred tortoise Geochelone elegans in captivity at Madras Snake Park. Cobra (Madras) 15: 10-12.

Tardent, P. 1972. Haltung und Zucht der Sternschildkröte, Testudo elegans. Salamandra 8(3-4): 165-175.

Vyas, R. 2005. Captive breeding of the Indian star tortoise (Geochelone elegans). Zoos' Print Journal 20(5): 1859-1864.

Vyas, R. 2006. The Indian star tortoise Geochelone elegans - Status in the protected areas of Gujarat and in Indian zoos. Zoos' Print Journal 21(4): 2220-2222.

Vyas, R. 2009. The status of north-western population of Star Tortoise (Geochelone elegans). ENVIS Bulletin Wildlife and Protected Areas 12(1): 11.

Vyas, R. 2011. Record size of Indian Star Tortoise: Geochelone elegans (Schöpff, 1795). Russian Journal of Herpetology 18(1): 47-50.

Vyas, R. 2012. Carapacial scute anomalies of star tortoise (Geochelone elegans) in Western India. Taprobanica 4(2): 105-107.

Vyas, R. 2014. Schildanomalien bei der Indischen Sternschildkröte, Geochelone elegans (Schoepff, 1795). Sauria (Berlin) 36(2): 37-41.

Vyas, R.; Parasharya, B.M. 2000. Distribution and status of the star tortoise (Geochelone elegans) in Gujarat State, India. Zoos' Print Journal 15(4): 239-242.

Vyas, R.; Thaker, M. 2017. Reptile Rap: Turtle Predation: Evidence of partial consumption of adult female Indian Star Tortoise (Geochelone elegans) by Asiatic Lion (Panthera leo persica): account of vertebrate predation on Testudines. Zoos' Print Magazine 32(12): 14-20.

Vyas, R.; Thaker, M. 2017. Turtle Predation: Evidence of partial consumption of adult female Indian Star Tortoise (Geochelone elegans) by Asiatic Lion (Panthera leo persica): account of vertebrate predation on Testudines. Zoos' Print 32(12): 14-20.

Walker, S. 2002. Asian turtle crisis. Global turtle conservation program. Hey! South Asian zoos! What are we doing? Zoos' Print Journal 17(9): 1-3.

Wermuta, H. 1960. Die Sternschildkröte Testudo elegans Schöpff. Aquarien und Terrarien (Leipzig) 7(10): 303-305.

Whelan, J.P.; Coakley, J. 1982. Life history notes: Geochelone elegans (Indian Star Tortoise). Fecundity. Herpetological Review 13(3): 97.

Whitaker, R. 1974. Feeding habits of the star tortoise - Geochelone elegans. Journal of the Bombay Natural History Society 71(1): 147-148.

Geochelone platynota

Anonymous. 2013. Searching for a home in the homeland of the Burmese Star Tortoise. The Tortoise 1(2): 26.

Charpentier, P.; Fuhrmann-Bacci, P. 2010. Essai d'installation et de maintenance d'une belle tortue terrestre etoilee de climat tropical humide: la tortue a dos plat ou etoilee de Birmanie Geochelone platynota (Blith, 1863). Cheloniens 17: 23-27.

Coutard, C.; Trani, M.; Tafani, J.S.; Cadi, A. 2004. Liste rouge TSA: Especes menacees elevees au centre A Cupulatta (1ere partie). Manouria 7(23): 37-44.

Elston, J.J. 2012. Coming out of their shells: enrichment for tortoises. Animal Keepers' Forum 39(7): 323-328.

Fife, J.D. 2007. Star Tortoises - The natural History, Caprive care, and Breeding of Geochelone elegans and Geochelone platynota. Living Art Publishing, Ada, Oklahoma. 116 pp.

Hoekstra, R.; Bidmon, H.J. 2006. The secret tortoise: Geochelone platynota, die Burmesische Stern-Schildkröte eine noch weit gehend unbekannte Landschildkröte. Schildkröten im Fokus 3(3): 3-22.

Jasser-Haeger, I.; Philippen, H.D. 2006. Burmesische Sternschildkröte Geochelone platynota (Blyth, 1863). Marginata 11: 31-34.

Moll, E.O. 1989. Geochelone platynota - Burmese star tortoise. Occasional Papers of the IUCN Species Survival Commission 5: 115.

Owens, T.C.; Scott, B. 2008. Significant firsts hatch at the San Diego Zoo while maintaining key programs. TSA Newsletter 2008 (August): 16.

Platt, S.G.; Khaing, S.T.; Win, K.K.; Kalyar 2001. A tortoise survey of Shwe Settaw Wildlife Sanctuary, Myanmar, with notes on the ecology of Geochelone platynota and Indotestudo elongata. Chelonian Conservation and Biology 4(1): 172-177.

Platt, S.G.; Platt, K.; Lay Lay Khaing; Thin Thin Yu; Shwe Htay Aung; New, S.S.; Me Me Soe; Khin Myo Myo; Tint Lwin; Win Ko Ko; Aung, S.H.N.; Rainwater, T.R. 2017. Back from the brink: Ex-situ conservation and recovery of the critically endangered Burmese Star Tortoise (Geochelone platynota) in Myanmar. Herpetological Review 48(3): 570-575.

Platt, S.G.; Platt, K.; Myo, K.M.; Moe, K.; Soe, M.M.; Naing, T.Z.; Lin, N.; Rainwater, T.R. 2013. Noteworthy records of chelonians from the Chindwin River Basin and Naga Hills of western Myanmar. Herpetological Conservation and Biology 8(2): 335-350.

Platt, S.G.; Thanda, S.; Win, K.K.; Platt, K.; Myo, K.M.; Rainwater, T.R.; Emmett, D. 2011. Geochelone platynota (Blyth 1863) - Burmese Star Tortoise, Kye Leik Chelonian Research Monographs 5: 057.1-057.9.

Platt, S.G.; Thanda, S.; Win, K.K.; Platt, K.; Myo, K.M.; Rainwater, T.R.; Emmett, D. 2011. Geochelone platynota (Blyth 1863) - Burmese Star Tortoise, Kye Leik. Chelonian Research Monographs 5: 057.1-057.9.

Platt, S.G.; Win, K.K.; Khaing, L.L.; Myo Myo Khin; Swe, T.; Lwin, T.; Rainwater, T.R. 2003. Population status and conservation of the critically endangered Burmese star tortoise Geochelone platynota in central Myanmar. Oryx 37(4): 464-471.

Platt, S.G.; Win, K.K.; Khaing, L.L.; Myo Myo Khin; Thanda, S.; Kalyar; Rainwater, T.R. 2004. Recent records and comments on the distribution and conservation status of Geochelone platynota (Blyth, 1863) in the dry zone of central Myanmar. Hamadryad 29(1): 128-131.

Soe, M.M.; Nwe, S.S.; Platt, S.G.; Rainwater, T.R. 2016. Natural history notes: Geochelone platynota (Burmese Star Tortoise). Scavenging. Herpetological Review 47(3): 444-445.