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Taxonomy of the genus Malacochersus Biodiversity of the family Testudinidae

Bibliography of the genus
Malacochersus (African Pancake Tortoise)

(Reptilia: Testudines: Testudinidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Malacochersus tornieri

Aoki, R. 1990. Life history notes: Malacochersus tornieri (Pancake Tortoise). Behavior. Herpetological Review 21(3): 60.

Bidmon, H.J. 2012. [Hamilton's rule, kin selection or even altruism in the pancake tortoise, Malacochersus tornieri? - an observation on social behaviour]. (In German). Schildkröten im Fokus 9(2): 3-26.

Boulenger, G.A. 1920. Une tortue extraordinaire: Testudo loveridgii, sp.n. C. R. Soc. Biol. (Paris) 170: 263-266.

Broadley, D.G. 1989. Malacochersus tornieri - Pancake tortoise; soft-shelled tortoise. Occasional Papers of the IUCN Species Survival Commission 5: 62-64.

Chansa, W.; Wagner, P. 2006. On the status of Malacochersus tornieri (Siebenrock, 1903) in Zambia. Salamandra 42(2-3): 187-190.

Darlington, A.F.; Davis, R.B. 1990. Reproduction in the Pancake Tortoise Malacochersus tornieri, in captive collections. Herpetological Review 21(1): 16-18.

Dathe, F. 2005. Pflege und Vermehrung der Spaltenschildkröte Malacochersus tornieri (Siebenrock, 1903), im Tierpark Berlin-Friedrichsfelde. Milu 11(5): 585-595.

Dickson, J. 1992. Maintenance and breeding the Pancake Tortoise at Bristol Zoo. IZN (International Zoo News) 39(3): 29-34.

Dowling, H.G. 1961. Vanishing giants and enduring dwarfs - the tortoises. Animal Kingdom 64: 66-75.

Dworschak, C. 2016. Die Haltung von Spaltenschildkröten Malacocersus torneri in Zimmerterrarium. Schildkröten im Fokus 13(2): 4-12.

Eglis, A. 1960. Notes on the soft-shelled tortoise Malacochersus tornieri (Siebenrock). Herpetologica 16: 12-14.

Eglis, A. 1964. Flat and fast - the pancake tortoise. Animal Kingdom 68: 107-110.

Ernst, C.H.; Laemmerzahl, A.F.; Holland, M.M.; Roscoe, R. 2002. Reproduction in four species of turtles at the National Zoological Park, Washington D.C. USA. Herpetological Bulletin 81: 17-21.

Ewert, M.A.; Hatcher, R.E.; Goode, J.M. 2004. Sex determination and ontogeny in Malacochersus tornieri, the pancake tortoise. Journal of Herpetology 38(2): 291-295.

Harmon, L.J. 2000. A translocation strategy for confiscated pancake tortoises. Chelonian Conservation and Biology 3(4): 738-743.

Honegger, R.E. 1971. Beitrag zur Fortpflanzungsbiologie einiger tropischer Reptilien. Freunde Kölner Zoo 4 [1970]: 175-179.

Hufer, H. 2006. Zur Haltung und Vermehrung der Spalten-Schildkröte Malacochersus tornieri (Siebenrock, 1903). Schildkröten im Fokus 3(1): 3-16.

Ireland, L.C.; Gans, C. 1972. The adaptive significance of the flexible shell of the tortoise Malacochersus tornieri. Animal Behaviour 20(4): 778-781.

IUCN Tortoise & Freshwater Turtle Specialist Group. 1996. Malacochersus tornieri. The IUCN Red List of Threatened Species 1996: e.T12696A3373951. http://dx.doi.org/10.2305/IUCN.UK.1996.RLTS.T12696A3373951.en.

Klemens, M.W.; Moll, D. 1995. An assessment of the effects of commercial exploitation on the Pancake Tortoise, Malacochersus tornieri, in Tanzania. Chelonian Conservation and Biology 1(3): 197-206.

Koch-Isenburg, L. 1977. Die ostafrikanische Spaltenschildkröte Malacochersus tornieri (Siebenrock). Naturwissenschaftlicher Verein Darmstadt Bericht N.F. 1976(1): 2-3.

Legrain, A.; Dumont, N. 2011. Maintenance et reproduction de la tortue à carapace souple malacochersus tornieri. Cheloniens 24: 24-29.

Malonza, P.K. 2004. Ecology and distribution of the pancake tortoise, Malacochersus tornieri in Kenya. Journal of East African Natural History 92(1-2): 81-96.

Mertens, R. 1942. Über das Verhalten der Spaltenschildkröte, Malacochersus tornieri (Siebenrock). Zoologische Garten N.F. (Leipzig) 14: 245-251.

Moll, D.; Klemens, M.W. 1996. Ecological characteristics of the Pancake Tortoise, Malacochersus tornieri, in Tanzania. Chelonian Conservation and Biology 2(1): 26-35.

Mwaya, R.T. 2006. Some insights of the pancake tortoise, Malacochersus tornieri: Testudinidae, ecology from Tarangire National Park, Tanzania. Chelonii 4: 115-126.

Mwaya, R.T. 2009. The floristic composition of the habitat of Malacochersus tornieri at a hill in Tarangire National Park, Tanzania. Salamandra 45(2): 115-118.

Mwaya, R.T. 2013. Ein Sammel-Stopp für Spaltenschildkröten – was bewirkt er? Marginata 37: 20-30.

Mwaya, R.T.; Moll, D.; Malonza, P.K.; Ngwava, J.M. 2018. Malacochersus tornieri (Siebenrock 1903) – Pancake Tortoise, Tornier’s Tortoise, Soft-shelled Tortoise, Crevice Tortoise, Kobe Ya Mawe/ Kobe Kama Chapati. Chelonian Research Monographs 5(107): 1-15.

Pauler, I. 1990. Zur Nachzucht der Spaltenschildkröte Malacochersus tornieri (Siebenrock, 1903). Herpetofauna (Weinstadt) 12(66): 6-10.

Philippen, D. 1989. Zucht von Spaltenschildkröte. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 42(9): 524.

Philippen, H.D. 2013. Eine außergewöhnliche Art: die Spaltenschildkröte (Malacochersus tornieri). Marginata 37: 12-19.

Pickering, D.C. 1990. Captive management of the Pancake Tortoise Malacochersus tornieri, at the Tulsa Zoological Park. AAZPA Regional Conference Proceedings 1990: 373-376.

Procter, J.B. 1922. A study of the remarkable tortoise, Testudo loveridgii Blgr., and the morphogeny of the chelonian carapace. Proceedings of the Zoological Society of London 1922: 483-526.

Riener, R. 1999. Haltung und Nachzucht der Spaltenschildkröte Malacochersus tornieri (Siebenrock, 1903). Emys 6(3): 4-19.

Rogner, M. 2001. Malacochersus tornieri Spaltenschildkröten. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 54(9): 12-15.

Rogner, M. 2012. Die Spaltenschildkröte - Malacochersus tornieri. Natur und Tier-Verlag, Münster. 61 pp.

Sanz, A.; Valverde, F.J. 2002. Captive maintenance of Malacochersus tornieri (Siebenrock, 1903). Reptilia (GB) 22: 49-53.

Schmidt, F. 2006. Some observations on pancake tortoises (Malacochersus tornieri) in Ruaha National Park, Tanzania. pp. 393-408. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Shaw, C.E. 1963. Notes on the eggs, incubation and young of some African reptiles. British Journal of Herpetology 3: 63-71.

Shaw, C.E. 1970. The hardy (and prolific) soft shelled tortoise. International Turtle and Tortoise Society Journal 4(1): 6-9, 30-31.

Siebenrock, F. 1903. Schildkröten des östlichen Hinterindien. Anz. Akad. Wiss. 40: 106-108.

Skelton, T.; Redrobe, S. 2002. Management guidelines for the welfare of zoo animals. The African pancake tortoise Malacochersus tornieri. Federation of Zoological Gardens of Great Britain and Ireland., London. 11 pp.

Stanford, C.B.; Tuma, M. 2013. Pancake breakfast: the Pancake Tortoise's biology is wonderfully bizarre, but also renders the species vulnerable to extinction. The Tortoise 1(2): 56-63.

Vinke, S.; Vinke, T. 1998. Die Spaltenschildkröte. Eine Spezialistin unter den Landschildkröten. Aquarium (Bornheim) 346: 50-54.

Vinke, T.; Vinke, S. 2000. Malacochersus tornieri (Siebenrock). Sauria (Berlin) 22(3): 495-498

Wettstein-Westersheimb, O. von 1924. Über Testudo tornieri Siebr. und Testudo loveridgii Blgr. Anatomischer Anzeiger 61: 201-208.

Wilke, H. 1984. Breeding the pancake tortoise Malacochersus tornieri at Frankfurt Zoo. International Zoo Yearbook 23: 137-139.