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Taxonomy of the genus Mesoclemmys Biodiversity of the family Chelidae

Bibliography of the genus
Mesoclemmys (Hump-backed Toad-headed Turtles)

(Reptilia: Testudines: Chelidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Mesoclemmys in general

Bour, R.; Zaher, H. 2005. A new species of MesoClemmys, from the open formations of northeastern Brazil (Chelonii, Chelidae). Papeis Avulsos de Zoologia (São Paulo) 45(24): 295-311.

McCord, W.P.; Joseph-Ouni, M.; Lamar, W.W. 2001. A taxonomic reevaluation of Phrynops (Testudines: Chelidae) with the description of two new genera and a new species of Batrachemys. Revista de Biologia Tropical 49(2): 715-764.

Mesoclemmys dahli

Forero-Medina, G.; Cardenas-Arevalo, G.; Castaño-Mora, O.V. 2011. Abundance, home range, and movement patterns of the endemic species Dahl's Toad-headed Turtle (Mesoclemmys dahli) in Cesar, Colombia. Chelonian Conservation and Biology 10(2): 228-236.

Forero-Medina, G.; Cardenas-Arevalo, G.; Castaño-Mora, O.V. 2012. Habitat modeling of Dahl's Toad-headed Turtle (Mesoclemmys dahli) in Colombia. Herpetological Conservation and Biology 7(3): 313-322.

Forero-Medina, G.; Castaño-Mora, O.V.; Cárdenas-Arevalo, G.; Medina-Rangel, G.F. 2013. Mesoclemmys dahli (Zangerl and Medem 1958) – Dahl's Toad-Headed Turtle, Carranchina, Tortuga Montañera. Chelonian Research Monographs 5(69): 1-8.

Medem, F. 1966. Contribuciones al conocimiento sobre la ecologia y distribucion geografica de Phrynops (Batrachemys) dahli; (Testudinata, Pleurodina, Chelidae). Caldasia (Bogota) 9: 467-489.

Medina-Rangel, G.F.; Forero-Medina, G. 2008. Geographic distribution: MesoClemmys dahli (Dahl's Toad-headed Turtle). Herpetological Review 39(3): 366.

Montes-Correa, A.C.; Saboya-Acosta, L.P.; Páez, V.P.; Vega, K.; Renjifo, J.M. 2014. [Distribution of tortoises and freshwater turtles of the Colombian Caribbean]. (In Spanish). Acta Biologica Colombiana 19(3): 341-350.

Ossa Velasquez, J.L. de la 1998. Phrynops dahli: a little-known turtle endemic to the Caribbean coast of Colombia. Reptilia (GB) 3: 47-50.

Ossa-Velasquez, J. de la 2000. Phrynops dahli. Eine wenig bekannte Schildkrötenart von der Karibikküste Kolumbiens. Reptilia (D) 5(21): 63-66.

Vargas-Ramírez, M.; Michels, J.; Castaño-Mora, O.V. 2012. Weak genetic divergence between the two South American toad-headed turtles Mesoclemmys dahli and M. zuliae (Testudines: Pleurodira: Chelidae). Amphibia-Reptilia 33(3-4): 373-385.

Zangerl, R.; Medem, F. 1958. A new-species of chelid turtle, Phrynops (Batrachemys) dahli, from Colombia. Bulletin of the Museum of Comparative Zoology at Harvard 119: 373-390.

Mesoclemmys gibba

Barreto, L.; Sousa Ribeiro, L.E. de; Nunes Ribeiro, A.B.; Rocha Azevedo, R.; Tavares, D.L.; Silva Abreu, J.M. da; Batista Cutrim, N. 2010. [Mapping of occurrence areas and conservation aspects of freshwater turtles in Maranhao State, Brazil]. (In Portuguese). Boletim do Laboratorio de Hidrobiologia 23(1): 49-56.

Bour, R. 2005. Type specimens of Emys gibba Schweigger, 1812, Platemys miliusii Dumeril & Bibron, 1835, and other related nominal species. Emys 12(5): 24-29.

Brito, E.S. de; Fagundes, C.K.; Strüssmann, C.; Pignati, M.T.; Vogt, R.C. 2012. Geographic distribution: Mesoclemmys gibba (Gibba Turtle). Brazil: Rondonia. Herpetological Review 43(3): 442.

Brito, E.S. de; Strüssmann, C.; Kawashita-Ribeiro, R.A.; Morais, D.H.; Ávila, R.W.; Campos, V.A. 2012. New records and distribution extensions of three species of Mesoclemmys Gray, 1863 (Testudines: Chelidae) in Mato Grosso state, Brazil, with observations on terrestrial movements. Check List 8(2): 294-297.

Deforce, E.A.; Deforce, C.D.; Lindeman, P.V. 2004. Natural history notes: Phrynops gibbus (Gibba Turtle). Trap-happy behavior. Herpetological Review 35(1): 55-56.

Elvey, C.; Newlon, K. 1998. Natural history notes: Eunectes murinus (Green Anaconda) and Phrynops giba [gibba] (gibba turtle). Predation. Herpetological Review 29(2): 103-104.

Ernst, C.H. 1981. Phrynops gibbus. Catalogue of American Amphibians and Reptiles 279: 1-2.

Goode, M. 1988. Reproduction and growth of the chelid turtle Phrynops (Mesoclemmys) gibbus at the Columbus Zoo. Herpetological Review 19(1): 11-13.

Grossmann, P. 1989. Beiträge zur Haltung und Nachzucht der Buckelschildkröte Phrynops (Mesoclemmys) gibbus (Schweigger 1812). Sauria (Berlin) 11(3): 11-15.

Hoogmoed, M.S.; Gruber, U. 1983. Spix and Wagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (the Netherlands). Spixiana Supplement 9: 319-415.

Kanberg, H. 1926. Mesoclemmys gibba Schw., eine seltene brasilianische Schlangenhalsschildkröte. Mitteilungen aus dem Zoologischen Museum in Berlin 12: 335-338.

Kanberg, H. 1937. Die Brasilianische Kielschildkröte (Mesoclemmys gibba Schw.). Aquarium (Berlin) 11: 159-160.

Keller, C.; Villamarín, F.; Bernhard, R.; Silva, D.F. da 2016. Checklist of chelonians from the upper Madeira River and the lower Madeira-Purus interfluvium (Brazilian Amazon), including a range expansion for Podocnemis sextuberculata Cornalia, 1849. Check List 12(4)(Article 1937): 1-32.

Lima Silveira, A. 2011. Geographic distribution: Mesoclemmys gibba. Herpetological Review 42(2): 239.

Medem, F. 1973. Beiträge zur Kenntnis über die Fortpflanzung der Buckel-Schildkröte, Phrynops (Mesoclemmys) gibbus. Salamandra 9(3-4): 91-98.

Metrailler, S. 1995. Notes sur l'elevage de Phrynops (Mesoclemmys) gibbus (Schweigger, 1812). CITS Bulletin 5: 33-40.

Métrailler, S. 2001. Elevage et reproduction de Phrynops gibbus (Schweigger, 1812). Manouria 4(10): 21-26.

Métrailler, S. 2006. Keeping and breeding of MesoClemmys gibba (Schweigger, 1812). pp. 338-341. In: Artner, H.; Farkas, B. & Loehr, V. (eds.). Turtles: proceedings: International Turtle & Tortoise Symposium, Vienna 2002. Edition Chimaira, Frankfurt am Main. 618 pp.

Mittermeier, R.A.; Rhodin, A.G.J.; Medem, F.; Soini, P.; Hoogmoed, M.S.; Carrillo de Espinoza, N. 1978. Distribution of the South American chelid turtle Phrynops gibbus, with observations on habitat and reproduction. Herpetologica 34(1): 94-100.

Molina, F.B.; Machado, F.A.; Zaher, H. 2012. Taxonomic validity of Mesoclemmys heliostemma (McCord, Joseph-Ouni & Lamar, 2001) (Testudines, Chelidae) inferred from morphological analysis. Zootaxa 3575: 63-77.

Oliveira-Ferronato, B. de; Ignacio Piña, C.; Cochachez Molina, F.; Espinoza, R.A.; Morales, V.R. 2013. Feeding habits of Amazonian freshwater turtles (Podocnemididae and Chelidae) from Peru. Chelonian Conservation and Biology 12(1): 119-126.

Oliveira-Ferronato, B. de; Molina, F.B.; Molina, F.C.; Espinosa, R.A.; Morales, V.R. 2011. New locality records for chelonians (Testudines: Chelidae, Podocnemididae, Testudinidae) from Departamento de Pasco, Peru. Herpetology Notes 4: 219-224.

Schneider, L.; Ferrara, C.R.; Vogt, R.C.; Santos-Junior, L.B. 2009. Geographic distribution: Phrynops gibbus (Gibba Turtle). Herpetological Review 40(2): 236.

Vogt, R.C.; Ferrara, C.R.; Schneider, L.; Santos Junior, L.B. 2009. Natural history notes: Brazilian Amazonian turtles. Habitat. Herpetological Review 40(2): 213.

Mesoclemmys nasuta

Bour, R. 1973. Contribution a la connaissance de Phrynops nasutus (Schweigger: 1812) et Phrynops tuberculatus (Luederwaldt: 1926). Description d'une nouvelle sous-espèce originaire de Paraguay, Phrynops tuberculatus Vanderhaegel (Testudinata - Pleurodira - Chelidae). Bulletin de la Société Zoologique de France 98(1): 175-190.

Fagundes, C.K.; Vogt, R.C.; Marco Junior, P. de 2016. Testing the efficiency of protected areas in the Amazon for conserving freshwater turtles. Diversity and Distributions 22(2): 123-135.

Lescure, J.; Fretey, J. 1975. Étude taxinomique de Phrynops (Batrachemys) nasutus (Schweigger) (Testudinata, Chelidae). Bulletin du Muséum National d'Histoire Naturelle (Zoologie) (Paris) 239: 1317-1328.

Medem, F. 1960. Informe sobre reptiles Colombianos. 4. El primer hallazgo de la tortuga Phrynops (Batrechemys) nasuta (Schweigger) en Colombia. Novedades Colombianas 1: 284-290.

Medem, P. 1960. (IV) Informe sobre reptiles Colombianos. El primer hallazgo de la Tortuga Phrynops (Batrachernys) nasuta (Schweigger) en Colombia. (V) Observaciones sobre la distribusion geografioa y ecologia de la Tortuga Phrynops geoffroana ssp. Novedades Colombianas 1(5): 284-300.

Mertens, R. 1967. Bemerkenswerte Süsswasserschildkröten aus Brasilien. Senckenbergiana Biologica 48: 71-82.

Mertens, R. 1970. Zur Kenntnis von Phrynops nasutus. Senckenbergiana Biologica 51: 17-20.

Metrailler, S. 1994. Observations de cinq specimens de Phrynops (Batrachemys) n. nasutus en Guyane Francaise. CITS Bulletin 3: 23-41.

Metrailler, S. 1995. Contribution a la connaissance de Phrynops (Batrachemys) n. nasutus en Guyane Francaise. CITS Bulletin 5: 25-30.

Rivas, G.A.; Barros, T.R.; Barros Molina, F. de; Trebbau, P.; Pritchard, P.C.H. 2015. The presence of Mesoclemmys raniceps and Mesoclemmys nasuta in Venezuela and comments on the type locality of Hydraspis maculata (Chelidae). Chelonian Conservation and Biology 14(1): 104-107.

Mesoclemmys perplexa

Bour, R.; Zaher, H. 2005. A new species of MesoClemmys, from the open formations of northeastern Brazil (Chelonii, Chelidae). Papeis Avulsos de Zoologia (São Paulo) 45(24): 295-311.

Campos, F.S.; Moraes, R.L. de; Amorim Pereira, C.S. 2011. New state record of Mesoclemmys perplexa Bour and Zaher, 2005 (Reptilia: Chelidae) in Brazil. Herpetology Notes 4: 263-264.

Loebmann, D. 2008. Geographic distribution: MesoClemmys perplexa. Herpetological Review 39(2): 236.

Mesoclemmys raniceps

Balensiefer, D.C.; Vogt, R.C. 2012. Geographic distribution: Mesoclemmys raniceps (Amazon Toad-headed Turtle). Herpetological Review 43(2): 303.

Brito, E.S. de; Strüssmann, C.; Kawashita-Ribeiro, R.A.; Morais, D.H.; Ávila, R.W.; Campos, V.A. 2012. New records and distribution extensions of three species of Mesoclemmys Gray, 1863 (Testudines: Chelidae) in Mato Grosso state, Brazil, with observations on terrestrial movements. Check List 8(2): 294-297.

Fachin-Teran, A.; Vogt, R.C.; Gomez, M. de F.; Soares, S. 1995. Alimentacion de tres especies de tortugas Chelidae en Costa Marques, Rondonia - Brasil. Boletin de Lima 16(91-96) [1994]: 409-416.

Gaibazzi, G. 2005. Elevage et reproduction de la platemyde de l'Amazone, Batrachemys raniceps (Gray, 1855). Manouria 8(27): 27-32.

Keller, C.; Villamarín, F.; Bernhard, R.; Silva, D.F. da 2016. Checklist of chelonians from the upper Madeira River and the lower Madeira-Purus interfluvium (Brazilian Amazon), including a range expansion for Podocnemis sextuberculata Cornalia, 1849. Check List 12(4)(Article 1937): 1-32.

Mertens, R. 1969. Eine neue Halswender-Schildkröte aus Peru. Senckenbergiana Biologica 50: 132.

McCord, W.P.; Joseph-Ouni, M.; Lamar, W.W. 2001. A taxonomic reevaluation of Phrynops (Testudines: Chelidae) with the description of two new genera and a new species of Batrachemys. Revista de Biologia Tropical 49(2): 715-764.

Molina, F.B.; Machado, F.A.; Zaher, H. 2012. Taxonomic validity of Mesoclemmys heliostemma (McCord, Joseph-Ouni & Lamar, 2001) (Testudines, Chelidae) inferred from morphological analysis. Zootaxa 3575: 63-77.

Moravec, J. 2017. First data on reproduction and hatchling morphology of Mesoclemmys heliostemma (McCord, Joseph-Ouni & Lamar, 2001). Herpetozoa 29(3-4): 205-208.

Morcatty, T.Q. 2015. Geographic distribution: Mesoclemmys raniceps (Black-lined Toad-headed Turtle). Herpetological Review 46(3): 382.

Morcatty, T.Q.; Cobra, I.V.D. 2015. Geographic distribution: Mesoclemmys heliostemma (Amazon Toad-headed Turtle). Herpetological Review 46(3): 381-382.

Oliveira-Ferronato, B. de; Molina, F.B.; Molina, F.; Espinosa, R.; Morales, V. 2010. Hatchling morphology of the Amazonian toad-headed turtle (Mesoclemmys raniceps (Gray 1855)) from Peruvian tropical Andes (Testudines: Chelidae). Herpetology Notes 3: 211-214.

Rivas, G.A.; Barros, T.R.; Barros Molina, F. de; Trebbau, P.; Pritchard, P.C.H. 2015. The presence of Mesoclemmys raniceps and Mesoclemmys nasuta in Venezuela and comments on the type locality of Hydraspis maculata (Chelidae). Chelonian Conservation and Biology 14(1): 104-107.

Schilde, M. 2008. Haltung und Erstnachzucht von Batrachemys raniceps (Gray, 1856), der Amazonas-Krötenkopfschildkröte. Radiata 17(3): 55-62.

Vasquez, M.R.O. 2003. Animales de Caza en la zona Reservada Allpahuayo-Mishana. Folia Amazonica 14(1): 7-16.

Vasquez, M.R.O. 2003. Una nueva localidad para la distribucion geografica de Batrachemys heliostemma (Reptilia, Chelidae) en la Amazonia Peruana. Folia Amazonica 14(1): 189-191.

Webb, R.G. 1995. The date of publication of Gray's Catalogue of shield Reptiles. Chelonian Conservation and Biology 1(4): 322-323.

Mesoclemmys tuberculata

Alves Pereira, A.M.; Vieira Brito, S.; Araujo Filho, J.A. de; Martins Teixeira, A.A.; Alves Teles, D.; Oliveira Santana, D.; Ferreira Lima, V.; Oliveira Almeida, W. de 2018. Diet and helminth parasites of freshwater turtles Mesoclemmys tuberculata, Phrynops geoffroanus (Pleurodira: Chelidae) and Kinosternon scorpioides (Criptodyra: Kinosternidae) in a semiarid region, northeast of Brazil. Acta Herpetologica 13(1): 21-32.

Barreto, L.; Sousa Ribeiro, L.E. de; Nunes Ribeiro, A.B.; Rocha Azevedo, R.; Tavares, D.L.; Silva Abreu, J.M. da; Batista Cutrim, N. 2010. [Mapping of occurrence areas and conservation aspects of freshwater turtles in Maranhao State, Brazil]. (In Portuguese). Boletim do Laboratorio de Hidrobiologia 23(1): 49-56.

Batistella, A.M.; Poetter, C.; Barreto, L.; Vogt, R.C. 2008. Geographic distribution: MesoClemmys tuberculata. Herpetological Review 39(1): 107-108.

Bour, R. 1973. Contribution a la connaissance de Phrynops nasutus (Schweigger: 1812) et Phrynops tuberculatus (Luederwaldt: 1926). Description d'une nouvelle sous-espèce originaire de Paraguay, Phrynops tuberculatus Vanderhaegel (Testudinata - Pleurodira - Chelidae). Bulletin de la Société Zoologique de France 98(1): 175-190.

Lima Silveira, A.; Valinhas e Valinhas, R. 2010. First record of Mesoclemmys tuberculata (Reptilia, Testudines, Chelidae) in the cerrado area of Minas Gerais state, southeastern Brazil. Biotemas 23(4): 157-161.

Loebmann, D.; Giacometti Mai, A.C.; Garcia, A.M. 2006. Reptilia, Chelidae, MesoClemmys tuberculata: geographical distribution extension. Check List 2(1): 32-33.

Mota Rodrigues, J.F.; Borges-Nojosa, D.M. 2012. Natural history notes: Mesoclemmys tuberculata (Tuberculate Toad-headed Turtle). Behavior. Herpetological Review 43(3): 475.

Oliveira Santana, D.; Marques, T.S.; Vieira, G.H.C.; Gomes Faria, R.; Oliveira Mesquita, D. 2015. Hatchling morphology of the Tuberculate Toadhead Turtle (Mesoclemmys tuberculata [Lüederwaldt, 1926]) from northeastern Brazil (Testudines: Chelidae). Herpetology Notes 8: 407-410.

Reis, P.M.; Fernandes Coelho, R.D.; Walker, F.M.; Barrios Ribeiro, L. 2014. Natural history notes: Mesoclemmys tuberculata (Tuberculate Toad-headed Turtle). Hatchlings. Herpetological Review 45(4): 691.

Santana, D.O.; Marques, T.S.; Vieira, G.H.C.; Moura, G.J.B.; Faria, R.G.; Mesquita, D.O. 2016. Mesoclemmys tuberculata (Luederwaldt 1926) – Tuberculate Toad-headed Turtle. Chelonian Research Monographs 5(97): 1-8.

Mesoclemmys vanderhaegei

Avila, R.W.; Souza, F.L.; Landgref-Filho, P.; Campos, A.L.C. 2006. Errata: Reptilia, Chelidae, Acanthochelys macrocephala: distribution, habitat, and geographical distribution map. Check List 2(2)

Avila, R.W.; Souza, F.L.; Landgref-Filho, P.; Campos, A.L.C. 2006. Reptilia, Chelidae, Acanthochelys macrocephala: distribution, habitat, and geographical distribution map. Check List 2(2): 35-37.

Baldo, D.; Krauczuk, E. 2000. Geographic distribution: Phrynops vanderhaegei (Vanderhaege's Toadhead Turtle). Herpetological Review 31(3): 184.

Baldo, D.; Martinez, P.; Boeris, J.M.; Giraudo, A.R. 2007. Reptilia, Chelonii, Chelidae, Phrynops geoffroanus Schweigger, 1812 and MesoClemmys vanderhaegei (Bour, 1973): distribution extension, new country record, and new province records in Argentina. Check List 3(4): 348-352.

Bour, R.; Pauler, I. 1987. Identite de Phrynops vanderhaegei Bour, 1973, et des especes affines (Reptilia-Chelonii-Chelidae). Bulletin du Museum d'Histoire Naturelle de Marseille 47: 3-23.

Brandao, R.A.; Zerbini, G.J.; Sebben, A.; Barros Molina, F. de 2002. Notes on distribution and habitats of Acanthochelys spixii and Phrynops vanderhaegei (Testudines, Chelidae) in Central Brazil. Boletin de la Asociacion Herpetologica Espanola 13(1-2): 11-15.

Brito, E.S. de; Strüssmann, C.; Baicere-Silva, C.M. 2009. Courtship behavior of MesoClemmys vanderhaegei (Bour, 1973) (Testudines: Chelidae) under natural conditions in the Brazilian Cerrado. Herpetology Notes 2: 67-72.

Brito, E.S. de; Strüssmann, C.; Ferreira Penha, J.M. 2009. Population structure of MesoClemmys vanderhaegei (Bour, 1973) (Testudines: Chelidae) in the Cerrado of Chapada dos Guimaraes, Mato Grosso, Brazil. Biota Neotropica 9(4): 245-248.

Brito, E.S. de; Strüssmann, C.; Kawashita-Ribeiro, R.A.; Morais, D.H.; Ávila, R.W.; Campos, V.A. 2012. New records and distribution extensions of three species of Mesoclemmys Gray, 1863 (Testudines: Chelidae) in Mato Grosso state, Brazil, with observations on terrestrial movements. Check List 8(2): 294-297.

Brito, E.S.; Souza, F.L.; Strüssmann, C. 2016. Feeding habits of Mesoclemmys vanderhaegei (Testudines: Chelidae). Acta Herpetologica 11(1): 1-13.

Cabrera, M.R.; Böhm, S. 2015. The identity of a controversial neotropical turtle hatchling (Reptilia: Testudines: Chelidae). Herpetological Review 46(4): 510-512.

Cassano, M.J. 2017. Restricción de la distribución de Mesoclemmys vanderhaegei (Bour, 1973) (Testudines: Chelidae) en Argentina. Cuadernos de Herpetologia 31(1): 25-27.

Etchepare, E.G. 2007. Geographic distribution: MesoClemmys vanderhaegei (NCN). Herpetological Review 38(4): 480.

Leandro Souza, F.; Martins, M.; Sawaya, R.J. 2000. A new record and observations of Vanderhaege's toad-headed turtle, Phrynops vanderhaegei (Testudines, Chelidae) in SE Brazil. Boletin de la Asociacion Herpetologica Espanola 11(2): 85-88.

Lima-Silveira, A. 2009. Geographic distribution: MesoClemmys vanderhaegei. Herpetological Review 40(2): 235-236.

Maffei, F.; Nascimento, B.T.M. do; Moya, G.M.; Donatelli, R.J. 2016. New distribution records of Mesoclemmys vanderhaegei (Testudines: Chelidae) from southeastern Brazil, including observations on reproduction. Journal of Threatened Taxa 8(11): 9322-9326.

Marques, T.S.; Böhm, S.; Brito, E.S. de; Cabrera, M.R.; Verdade, L.M. 2014. Mesoclemmys vanderhaegei (Bour 1973) – Vanderhaege's Toad-headed Turtle, Karumbé-hy. Chelonian Research Monographs 5(83): 1-8.

Marques, T.S.; Brito, E.S.; Lara, N.R.F.; Beloto, L.M.; Valadão, R.M.; Camargo, P.B. de; Verdade, L.M. 2018. The trophic niche of Mesoclemmys vanderhaegei (Testudines: Chelidae): evidence from stable isotopes. Zoologia 34: e19985.

Marques, T.S.; Lara, N.R.F.; Bassetti, L.A.B.; Ferronato, B.O.; Malvasio, A.; Verdade, L.M. 2013. Population structure of Mesoclemmys vanderhaegei (Testudines, Chelidae) in a silvicultural system in southeastern Brazil. Herpetology Notes 6: 179-182.

Métrailler, S. 2005. Observation platemyde de Vanderhaege, Bufocephala vanderhaegei (Bour, 1973) au Paraguay. Manouria 8(27): 33-39.

Vinke, T.; Vinke, S.; Köhler, G. 2013. Mesoclemmys vanderhaegei (Bour, 1973), die Südliche Krötenkopfschildkröte. Schildkröten im Fokus 10(4): 3-25.

Mesoclemmys zuliae

Pritchard, P.C.H. 1987. Phrynops zuliae. Catalogue of American Amphibians and Reptiles 404: 1-2.

Rojas-Runjaic, F.J.M. 2009. MesoClemmys zuliae. Die geheimnisvolle Krötenkopfschildkröte aus dem Süden des Maracaibo-Sees. Reptilia (D) 14(80): 34-39.

Vargas-Ramírez, M.; Michels, J.; Castaño-Mora, O.V. 2012. Weak genetic divergence between the two South American toad-headed turtles Mesoclemmys dahli and M. zuliae (Testudines: Pleurodira: Chelidae). Amphibia-Reptilia 33(3-4): 373-385.