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Miralles, A.; Macleod, A.; Rodríguez, A.; Ibáñez, A.; Jiménez-Uzcategui, G.; Quezada, G.; Vences, M.; Steinfartz, S. 2017 Shedding light on the imps of darkness: an integrative taxonomic revision of the Galápagos Marine Iguanas (genus Amblyrhynchus). Zoological Journal of the Linnean Society 181: 678–710.