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Taxonomy of the family Liolaemidae
Bibliography of the genus Phymaturus
Biodiversity of the family Liolaemidae








Genus
Phymaturus

South American Chuckwallas

Hochgebirgsleguane

Sydamerikanske Højlandsleguaner

1837 Phymaturus Gravenhorst (type species: Phymaturus palluma sensu Gravenhorst 1837 [not Lacerta palluma Molina 1782])
1843 Centrura Bell (type species: Centrura flagellifer Bell 1843; syn. Peters & Donoso-Barros 1970 ICZN (2005): Opinion 2118)
Contents: 51 species, of which 50 (98.0%) are endemic.
Remarks: The type species of the genus Phymaturus, Lacerta palluma Molina 1782, was shown by Cei & Lescure (1985) to be identical with Callopistes maculatus Gravenhorst 1838 (family Teiidae). As a consequence of the rules of zoological nomenclature, the well known species Callopistes maculatus thus should be referred to as Callopistes palluma, this name being an older specific name than maculatus. Consequently, the iguanid species known until then as Phymaturus palluma, should from then on be referred to as Centrura flagellifer, the next available name, which has also been the case in some publications (e.g., Cei 1986). Furthermore, all other iguanid species until then placed in the genus Phymaturus, should also be assigned to the genus Centrura. However, the International Commission on Zoological Nomenclature decided to conserve the use of the name Phymaturus (International Commission on Zoological Nomenclature 2005: Opinion 2118).
Distribution: Argentina, Chile.

Phymaturus aguanegra

Agua Negra Chuckwalla

Agua-Negra-Hochgebirgsleguan

Agua Negra-højlandsleguan

2013 Phymaturus aguanegra Lobo, Laspiur & Acosta

Distribution: Argentina.


Phymaturus aguedae

Provincia Mountain Chuckwalla

Provincia-Hochgebirgsleguan

Provincia-højlandsleguan

2014 Phymaturus aguedae Troncoso-Palacios & Esquerré

Distribution: Chile.


Phymaturus alicahuense

Valparaiso Chuckwalla

Valparaiso-Hochgebirgsleguan

Valparaiso-højlandsleguan

2010 Phymaturus alicahuense Nunez, Veloso, Espejo, Veloso, Cortes & Araya

Distribution: Chile.


Phymaturus antofagastensis

Antofagasta Chuckwalla

Antofagasta-Hochgebirgsleguan

Antofagasta-højlandsleguan

1985 Phymaturus antofagastensis Pereyra

Distribution: Argentina.


Phymaturus bibronii

Ovalle Chuckwalla

Ovalle-Hochgebirgsleguan

Ovalle-højlandsleguan

1848 Oplurus bibronii Guichenot
Phymaturus bibronii Troncoso-Palacios, Lobo, Etheridge, Acosta & Laspiur 2013
2010 Phymaturus paihuanense Nunez, Veloso, Espejo, Veloso, Cortes & Araya (Troncoso-Palacios, Lobo, Etheridge, Acosta & Laspiur 2013)

Remarks: Previously regarded as a synonym of palluma (e.g., Boulenger 1885; Peters & Donoso-Barros 1970). Revalidated as a separate species by Troncoso-Palacios, Lobo, Etheridge, Acosta & Laspiur (2013).
Distribution: Chile.


Phymaturus cacivioi

El Cuy Chuckwalla

El-Cuy-Hochgebirgsleguan

El Cuy-højlandsleguan

2015 Phymaturus cacivioi Lobo & Nenda

Distribution: Argentina.


Phymaturus calcogaster

Chubut Chuckwalla

Chubut-Hochgebirgsleguan

Chubut-højlandsleguan

2003 Phymaturus calcogaster Scolaro & Cei

Distribution: Argentina.


Phymaturus camilae

Sacanana River Chuckwalla, (Camila's Patagonian Rocky Lizard)

Sacanana-Hochgebirgsleguan

Sacanana-højlandsleguan

2013 Phymaturus camilae Scolaro, Jara & Pincheira-Donoso

Distribution: Argentina.


Phymaturus castillensis

Sierra del Castillo Chuckwalla

Castillo-Hochgebirgsleguan

Castillo-højlandsleguan

2010 Phymaturus castillensis Scolaro & Pincheira-Donoso

Distribution: Argentina.


Phymaturus ceii

Chasicó Chuckwalla

Chasicó-Hochgebirgsleguan

Chasicó-højlandsleguan

2007 Phymaturus ceii Scolaro & Ibargüengoytía

Distribution: Argentina.


Phymaturus curivilcun

Argentinian Black Chuckwalla, (Black Lizard)

Schwarze Hochgebirgsleguan

Sort Højlandsleguan

2016 Phymaturus curivilcun Scolaro, Corbalán, Tappari & Streitenberger

Distribution: Argentina.


Phymaturus damasense

Las Damas River Chuckwalla

Las-Damas-River-Hochgebirgsleguan

Las Damas River-højlandsleguan

2012 Phymaturus damasense Troncoso-Palacios & Lobo

Distribution: Chile.


Phymaturus darwini

Riecillo Valley Chuckwalla

Riecillo-Hochgebirgsleguan

Riecillo-højlandsleguan

2010 Phymaturus darwini Nunez, Veloso, Espejo, Veloso, Cortes & Araya

Distribution: Chile.


Phymaturus delheyi

Butacó Creek Chuckwalla

Butacó-Hochgebirgsleguan

Butacó-højlandsleguan

2011 Phymaturus delheyi Avila, Perez, Perez & Morando

Distribution: Argentina.


Phymaturus denotatus

Laguna Blanca Chuckwalla

Laguna-Blanca-Hochgebirgsleguan

Laguna Blanca-højlandsleguan

2012 Phymaturus denotatus Lobo, Nenda & Slodki

Distribution: Argentina.


Phymaturus desuetus

Rio Negro Chuckwalla

Rio-Negro-Hochgebirgsleguan

Rio Negro-højlandsleguan

2009 Phymaturus desuetus Scolaro & Tappari

Distribution: Argentina.


Phymaturus dorsimaculatus

Copahue Chuckwalla

Copahue-Hochgebirgsleguan

Copahue-højlandsleguan

2005 Phymaturus dorsimaculatus Lobo & Quinteros

Remarks: Regarded as a synonym of vociferator by Pincheira-Donoso, Scolaro & Sura (2008) and Pincheira-Donoso (2011), however, Lobo, Espinoza & Quinteros (2010) questioned the synonymization, and also Avila, Martinez & Morando (2013) listed dorsimaculatus as a valid species. Pending further studies, it is retained here as a valid species.
Distribution: Argentina.


Phymaturus etheridgei

White-spotted Chuckwalla

Weißflecken-Hochgebirgsleguan

Hvidplettet Højlandsleguan

2010 Phymaturus etheridgei Lobo, Abdala & Valdecantos

Distribution: Argentina.


Phymaturus excelsus

Pied Chuckwalla

Bunte Hochgebirgsleguan

Broget Højlandsleguan

2005 Phymaturus excelsus Lobo & Quinteros

Distribution: Argentina.


Phymaturus extrilidus

Dauntless Chuckwalla

Invernada-Hochgebirgsleguan

Invernada-højlandsleguan

2012 Phymaturus extrilidus Lobo, Espinoza, Sanabria & Quiroga

Distribution: Argentina.


Phymaturus felixi

Paso de Indios Chuckwalla

Paso-de-Indios-Hochgebirgsleguan

Paso de Indios-højlandsleguan

2010 Phymaturus felixi Lobo, Abdala & Valdecantos

Distribution: Argentina.


Phymaturus fiambala

Sierra de Fiambalá Chuckwalla

Sierra-de-Fiambalá-Hochgebirgsleguan

Sierra de Fiambalá-højlandsleguan

2019 Phymaturus fiambala Lobo, Hibbard, Quipildor & Valdecantos

Distribution: Argentina.





Phymaturus gynechlomus

Cruz de Piedra Chuckwalla

Cruz-de-Piedra-Hochgebirgsleguan

Cruz de Piedra-højlandsleguan

2009 Phymaturus gynechlomus Corbalán, Scolaro & Debandi

Remarks: Status uncertain. Considered a synonym of palluma by Lobo, Abdala & Valdecantos (2010) and Abdala & Quinteros (2014), but Scolaro (2010) rejected this. Also Avila, Martinez & Morando (2013) retained gynechlomus as a valid species, pending further studies.
Distribution: Argentina.


Phymaturus indistinctus

Las Pulgas Chuckwalla

Las-Pulgas-Hochgebirgsleguan

Las Pulgas-højlandsleguan

1975 Phymaturus patagonicus indistinctus Cei & Castro
Centrura patagonica indistincta Cei 1986
Phymaturus indistinctus Etheridge 1995

Distribution: Argentina.


Phymaturus laurenti

El Peñón Chuckwalla

El-Peñón-Hochgebirgsleguan

El Peñón-højlandsleguan

2010 Phymaturus laurenti Lobo, Abdala & Valdecantos

Distribution: Argentina.


Phymaturus loboi

Baños Campanario Chuckwalla, (Lobo's Rocky Lizard)

Baños-Campanario-Hochgebirgsleguan

Baños Campanario-højlandsleguan

2018 Phymaturus loboi Troncoso-Palacios, Ferri-Yáñez, Laspiur & Aguilar

Distribution: Argentina, Chile.





Phymaturus mallimaccii

Cueva de Peez Chuckwalla

Cueva-de-Peez-Hochgebirgsleguan

Cueva de Peez-højlandsleguan

1980 Phymaturus mallimacci Cei
Centrura mallimaccii Cei 1986

Distribution: Argentina.


Phymaturus manuelae

Comallo Chuckwalla

Comallo-Hochgebirgsleguan

Comallo-højlandsleguan

2008 Phymaturus manuelae Scolaro & Ibargüengoytía

Distribution: Argentina.


Phymaturus maulense

Maule Chuckwalla

Maule-Hochgebirgsleguan

Maule-højlandsleguan

2010 Phymaturus maulense Nunez, Veloso, Espejo, Veloso, Cortes & Araya

Distribution: Chile.


Phymaturus nevadoi

Nevado Mountains Chuckwalla

Nevado-Hochgebirgsleguan

Nevado-højlandsleguan

1975 Phymaturus patagonicus nevadoi Cei & Roig
Centrura patagonica nevadoi Cei 1986
Phymaturus nevadoi Etheridge 1995

Distribution: Argentina.


Phymaturus palluma

Cordón del Portillo Chuckwalla

Cordón-del-Portillo-Hochgebirgsleguan

Cordón del Portillo-højlandsleguan

1837 Phymaturus palluma Gravenhorst [not Lacerta palluma Molina 1782]
1843 Centrura flagellifer Bell (Peters & Donoso-Barros 1970)
Phymaturus flagellifer Lescure & Cei 1991
1848 Oplurus bibroni Guichenot (Peters & Donoso-Barros 1970)

Remarks: Now restricted to Argentina. The type locality was restricted to Cordón del Portillo, Mendoza Province, by Lobo & Etheridge (2013). Records from Chile refer to aguedae (Troncoso-Palacios & Esquerré 2014). See remarks under gynechlomus.
Distribution: Argentina.


Phymaturus patagonicus

Patagonian Chuckwalla, (Argentine Chuckwalla)

Patagonische Hochgebirgsleguan

Patagonisk Højlandsleguan

1896 Phymaturus patagonicus Koslowsky
Phymaturus palluma patagonicus Peters & Donoso-Barros 1970
Centrura patagonica Cei 1986

Remarks: Revalidated as a distinct species by Cei & Castro (1973).
Distribution: Argentina.


Phymaturus payuniae

Payun Plateau Chuckwalla

Payun-Hochgebirgsleguan

Payun-højlandsleguan

1975 Phymaturus patagonicus payuniae Cei & Castro
Centrura patagonica payuniae Cei 1986
Phymaturus payuniae Etheridge 1995

Distribution: Argentina.


Phymaturus punae

Puna Chuckwalla

Puna-Hochgebirgsleguan

Puna-højlandsleguan

1985 Phymaturus punae Cei, Etheridge & Videla
Centrura punae Cei 1986

Distribution: Argentina.


Phymaturus querque

Neuquén Chuckwalla

Neuquén-Hochgebirgsleguan

Neuquén-højlandsleguan

2010 Phymaturus querque Lobo, Abdala & Valdecantos

Distribution: Argentina.


Phymaturus rahuensis

Bajada del Rahue Chuckwalla

Rahue-Hochgebirgsleguan

Rahue-højlandsleguan

2016 Phymaturus rahuensis Marín, Pérez, Minoli, Morando & Avila

Distribution: Argentina.


Phymaturus roigorum

Mendoza Yellow-tailed Chuckwalla

Gelbschwanz-Hochgebirgsleguan

Gulhalet Højlandsleguan

2007 Phymaturus roigorum Lobo & Abdala

Distribution: Argentina.


Phymaturus sinervoi

Cari Laufquen Chuckwalla

Cari-Laufquen-Hochgebirgsleguan

Cari Laufquen-højlandsleguan

2012 Phymaturus sinervoi Scolaro, Cruz & Ibargüengoytía

Distribution: Argentina.


Phymaturus sitesi

Auca Mahuida Volcano Chuckwalla

Auca-Mahuida-Hochgebirgsleguan

Auca Mahuida-højlandsleguan

2011 Phymaturus sitesi Avila, Perez, Perez & Morando

Distribution: Argentina.


Phymaturus somuncurensis

Somuncura Plateau Chuckwalla

Somuncura-Hochgebirgsleguan

Somuncura-højlandsleguan

1975 Phymaturus patagonicus somuncurensis Cei & Castro
Centrura patagonica somuncurensis Cei 1986
Phymaturus somuncurensis Etheridge 1995

Distribution: Argentina.


Phymaturus spectabilis

Climbing Chuckwalla

Klätternde Hochgebirgsleguan

Klatrende Højlandsleguan

2005 Phymaturus spectabilis Lobo & Quinteros
2008 Phymaturus agilis Scolaro, Ibargüengoytía & Pincheira-Donoso (Lobo, Cruz & Abdala 2012)

Remarks: Scolaro, Corbalan, Tappari & Streitenberger (2016) treated agilis as a valid species.
Distribution: Argentina.


Phymaturus spurcus

Huanuluan Chuckwalla

Huanuluan-Hochgebirgsleguan

Huanuluan-højlandsleguan

1921 Phymaturus spurcus Barbour

Remarks: Previously regarded as a synonym of patagonicus (e.g., Burt & Burt 1931; Cei 1986). Revalidated by Lobo & Quinteros (2005).
Distribution: Argentina.


Phymaturus tenebrosus

Dark Chuckwalla

Dunkle Hochgebirgsleguan

Mørk Højlandsleguan

2005 Phymaturus tenebrosus Lobo & Quinteros

Distribution: Argentina.


Phymaturus timi

Pehuenches Chuckwalla

Pehuenches-Hochgebirgsleguan

Pehuenches-højlandsleguan

2019 Phymaturus timi Hibbard, Nenda & Lobo

Distribution: Argentina.





Phymaturus tromen

Tromen Volcano Chuckwalla

Tromen-Hochgebirgsleguan

Tromen-højlandsleguan

2015 Phymaturus tromen Lobo & Nenda

Distribution: Argentina.


Phymaturus verdugo

Volcán Peteroa Chuckwalla

Peteroa-Hochgebirgsleguan

Peteroa-højlandsleguan

2003 Phymaturus verdugo Cei & Videla

Distribution: Argentina.


Phymaturus videlai

Buen Pasto Chuckwalla

Buen-Pasto-Hochgebirgsleguan

Buen Pasto-højlandsleguan

2010 Phymaturus videlai Scolaro & Pincheira-Donoso

Distribution: Argentina.


Phymaturus vociferator

Antuco Chuckwalla

Antuco-Hochgebirgsleguan

Antuco-højlandsleguan

2004 Phymaturus vociferator Pincheira-Donoso

Remarks: Some authors regard dorsimaculatus as a synonym of vociferator (see remarks under dorsimaculatus). Records from Argentina refer to dorsimaculatus, if accepted as a valid species.
Distribution: Chile.


Phymaturus williamsi

Calingasta Chuckwalla

Calingasta-Hochgebirgsleguan

Calingasta-højlandsleguan

2013 Phymaturus williamsi Lobo, Laspiur & Acosta

Distribution: Argentina.


Phymaturus yachanana

Sierra Grande Chuckwalla

Sierra-Grande-Hochgebirgsleguan

Sierra Grande-højlandsleguan

2014 Phymaturus yachanana Avila, Pérez, Minoli & Morando

Distribution: Argentina.


Phymaturus zapalensis

Zapala Chuckwalla

Zapala-Hochgebirgsleguan

Zapala-højlandsleguan

1975 Phymaturus patagonicus zapalensis Cei & Castro
Centrura patagonica zapalensis Cei 1986
Phymaturus zapalensis Etheridge 1995

Distribution: Argentina.