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Eliades, S.J.; Brown, R.M.; Huang, W.S.; Siler, C.D. 2021 Taxonomic revision of Scaly-toed Geckos (Reptilia: Gekkonidae: Lepidodactylus) in the northern Philippines, with descriptions of four new species. Herpetological Monographs 35(1): 90-111.