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Taxonomy of the genus Acanthodactylus Biodiversity of the family Lacertidae

Bibliography of the genus
Acanthodactylus (Fringe-toed Lizards)

(Reptilia: Sauria: Lacertidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Acanthodactylus in general

Arnold, E.N. 1983. Osteology, genitalia and the relationships of Acanthodactylus (Reptilia: Lacertidae). Bulletin of the British Museum of Natural History (Zoology) 44(5): 291-339.

Bischoff, W. 1990. Übersicht der Arten und Unterarten der Familie Lacertidae. 1. Die Gattungen Acanthodactylus, Adolfus, Algyroides und Australolacerta. Die Eidechse 1(1): 18-23.

Blanc, C.P. 1979. Études sur les acanthodactyles de Tunisie: 2. Relations biométriques. Archives de l'Institut Pasteur de Tunis 56(1-2): 57-65.

Blanc, C.P. 1980. Etudes sur les acanthodactyles de Tunisie (Sauria: Lacertidae). 3. Variabilite morphologique et ses implications systematiques. Bulletin de la Societe Zoologique de France 104(4): 445-465.

Blanc, C.P. 1980. Studies on the Acanthodactylus of Tunisia. IV. Geographic distribution and habitats. Journal of Herpetology 14(4): 391-398.

Blanc, C.P.; Ineich, I. 1985. Etudes sur les acanthodactyles de Tunisie: 7 - les acanthodactyles de l'extreme-sud tunisien. Amphibia-Reptilia 6(1): 45-52.

Blanc, F.; Cariou, M.L. 1980. High genetic variability of lizards of the sand-dwelling lacertid genus Acanthodactylus. Genetica (Dordrecht) 54(2): 141-147.

Blanc, F.; Cariou, M.L. 1987. Geographical variability of allozyme variation and between Tunisian lizards of the sand-dwelling lacertid genus Acanthodactylus. Genetica (Dordrecht) 72(1): 13-25.

Bons, J.; Geniez, P. 1995. Contribution to the systematics of the lizard Acanthodactylus erythrurus (Sauria, Lacertidae) in Morocco. Herpetological Journal 5(3): 271-280.

Boulenger, G.A. 1918. Sur les lezards du genre Acanthodactylus Wieg. Bulletin de la Société Zoologique de France 43: 143-155.

Boulenger, G.A. 1919. Sur les lezards du genre Acanthodactylus Wiegm. Bulletin de la Société Zoologique de France 43: 143-155.

Brito, J.C.; Rebelo, H.; Crochet, P.A.; Geniez, P. 2008. Data on the distribution of amphibians and reptiles from north and west Africa, with emphasis on Acanthodactylus lizards and the Sahara Desert. Herpetological Bulletin 105: 19-27.

Carretero, M.A.; Fonseca, M.M.; García-Muñoz, E.; Brito, J.C.; Harris, D.J. 2011. Adding Acanthodactylus beershebensis to the mtDNA phylogeny of the Acanthodactylus pardalis group. North-Western Journal of Zoology 7(1): 138-142.

Corti, C.; Bischoff, W.; Böhme, W. 1999. Die Lacertiden der Mittelmeer-Inseln. Ein erste Übersicht. Die Eidechse 10(2): 41-58.

Crochet, P.A.; Geniez, P.; Ineich, I. 2003. A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): Systematic and biogeographical implications. Zoological Journal of the Linnean Society 137(1): 117-155.

Duvdevani, I. 1971. Nasal salt secretion and electrolyte balance in Acanthodactylus lizards. Int. J. Biomet. 15: 87.

Duvdevani, I.; Borut, A. 1974. Mean body temperature and heat absorption in four species of Acanthodactylus lizards (Lacertidae). Herpetologica 30(2): 176-181.

Duvdevani, I.; Borut, A. 1974. Oxygen consumption and evaporative water loss in four species of Acanthodactylus (Lacertidae). Copeia 1974(1): 155-164.

Ferreira, J.B. 1892. Sobre o "Acanthodactylus" de Portugal. Jornal de Sciencias Mathematicas, Physicas e Naturaes (Ser. 2) (Lisboa) 2: 188-194.

Fonseca, M.M.; Brito, J.C.; Paulo, O.S.; Carretero, M.A.; Harris, D.J. 2009. Systematic and phylogeographical assessment of the Acanthodactylus erythrurus group (Reptilia: Lacertidae) based on phylogenetic analyses of mitochondrial and nuclear DNA. Molecular Phylogenetics and Evolution 51(2): 131-142.

Fonseca, M.M.; Brito, J.C.; Rebelo, H.; Kalboussi, M.; Larbes, S.; Carretero, M.A.; Harris, D.J. 2008. Genetic variation among spiny-footed lizards in the Acanthodactylus pardalis group from North Africa. African Zoology 43(1): 8-15.

Geniez, P.; Lluch, P. 2003. New data on fringe-toed lizards of the Acanthodactylus scutellatus group in Mauritania. Herpetozoa 17(1-2): 88-89.

Harris, D.J.; Arnold, E.N. 2000. Elucidation of the relationships of spiny-footed lizards, Acanthodactylus spp. (Reptilia: Lacertidae) using mitochondrial DNA sequence, with comments on their biogeography and evolution. Journal of Zoology (London) 252(3): 351-362.

Heidari, N.; Rastegar-Pouyani, E.; Rastegar-Pouyani, N.; Faizi, H. 2014. Molecular phylogeny and biogeography of the genus Acanthodactylus Fitzinger, 1834 (Reptilia: Lacertidae) in Iran, inferred from mtDNA Sequences. Zootaxa 3860(4): 379-395.

Mayer, W. 1989. Comments on Szczerbak's (1975) catalogue of the African sand lizards (Reptilia: Sauria: Eremiainae). Herpetozoa 1(3-4): 133-137.

Mellado, J.; Olmedo, G. 1990. El genero Acanthodactylus en Marruecos: problemas de identification en los grupos de especies A. pardalis y A. scutellatus. Amphibia-Reptilia 11(2): 131-146.

Moravec, J.; Baha el Din, S.; Seligmann, H.; Sivan, N.; Werner, Y.L. 1999. Systematics and distribution of the Acanthodactylus pardalis group (Lacertidae) in Egypt and Israel. Zoology in the Middle East 17: 21-50.

Pollo, C.J.; Perez-Mellado, V. 1988. Trophic ecology of a taxocenosis of Mediterranean Lacertidae. Ecologia Mediterranea 14(3-4): 131-147.

Salvador, A. 1982. A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologische Monographien 16: 1-167.

Schlüter, U. 2002. Die Fransenfingereidechsen (Acanthodactylus) Nordafrikas. Draco 3(2) (10): 58-65.

Schlüter, U. 2007. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. 1. Allgemeines zu Ökologie und Verhalten. Reptilia (D) 12(66): 68-76.

Schlüter, U. 2007. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. 2. Grundlagen der Terrarienhaltung und Reproduktionsbiologie. Reptilia (D) 12(67): 56-65.

Schlüter, U. 2008. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. 3. Die Acanthodactylus-cantoris-Gruppe. Reptilia (D) 13(70): 70-75.

Schlüter, U. 2008. Genus Acanthodactylus fringe-fingered lizards in the wild and in the terrarium. Part 1: general ecology and behavior. Reptilia (GB) 57: 64-71.

Schlüter, U. 2008. Genus Acanthodactylus: fringe-fingered lizards in the wild and in the terrarium. Part 2: captive care and reproduction. Reptilia (GB) 58: 63-71.

Schlüter, U. 2009. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. 4. Die Acanthodactylus-pardalis-Gruppe. Reptilia (D) 14(76): 52-60.

Schlüter, U. 2010. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium Teil 5a: Die Acanthodactylus-scutellatus- Gruppe. Reptilia (D) 15(83): 40-45.

Schlüter, U. 2010. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. Teil 5b: Die Acanthodactylus-scutellatus-Gruppe. Reptilia (D) 15(84): 62-66.

Schlüter, U. 2013. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. Teil 6: Die Acanthodactylus-boskianus-Gruppe. Reptilia (D) 18(99): 58-64.

Sinha, P.P. 1985. Some observations on food habits of two species of lizards of western Rajasthan. Plant Protection Bulletin (Faridabad) 37(3-4): 35-36.

Szczerbak, N.N. 1975. [Catalog of African [and Middle Eastern] Desert Lizards]. (In Russian). Nauka Dymka, Kiev. 84 pp.

Szczerbak, N.N. 1989. Catalogue of the African sand lizards (Reptilia: Sauria: Eremiainae: Lampreremias, Pseuderemias, Taenieremias, Mesalina, Meroles). Herpetozoa 1(3-4): 119-132. (Translation of 1975 Russian original).

Werner, Y.L.; Gajst, O.; Talbi, R.; Bouskila, A. 2012. Acanthodactylus opheodurus Arnold, 1980 in the Levant revisited, and the striped patterns of Levantine Acanthodactylus (Reptilia: Lacertidae). Zoology in the Middle East 56: 31-38.

Zaady, E.; Bouskila, A. 2002. Lizard burrows association with successional stages of biological soil crusts in an arid sandy region. Journal of Arid Environments 50(2): 235-246.

Acanthodactylus aegyptius

Baha el Din, S.M. 2007. A new lizard of the Acanthodactylus scutellatus group (Squamata: Lacertidae) from Egypt. Zoology in the Middle East 40: 21-32.

Stark, G.; David, D.; Lwein, A.; Meiri, S. 2018. Natural history notes: Acanthodactylus aegyptius (Egyptian Fringed-fingered Lizard). Tail bifurcation. Herpetological Review 49(2): 324-325.

Acanthodactylus ahmaddisii

Werner, Y.L. 2004. A new species of the Acanthodactylus pardalis group (Reptilia: Lacertidae) from Jordan. Zoology in the Middle East 32: 39-45.

Acanthodactylus arabicus

Haas, G.; Battersby, J.C. 1959. Amphibians and reptiles from Arabia. Copeia 1959: 196-202.

Acanthodactylus aureus

Böhme, W. 1978. Zur Herpetofaunistik des Senegal. Bonner Zoologische Beiträge 29(4): 360-417.

Crochet, P.A.; Geniez, P.; Ineich, I. 2003. A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): Systematic and biogeographical implications. Zoological Journal of the Linnean Society 137(1): 117-155.

Schweiger, M. 2013. [Acanthodactylus aureus Guenther, 1903 - Fringe-toed Lizard]. (In German). Die Eidechse 24(2): 43-50.

Acanthodactylus beershebensis

Carretero, M.A.; Fonseca, M.M.; García-Muñoz, E.; Brito, J.C.; Harris, D.J. 2011. Adding Acanthodactylus beershebensis to the mtDNA phylogeny of the Acanthodactylus pardalis group. North-Western Journal of Zoology 7(1): 138-142.

Goldberg, S.R. 2014. Reproduction in the Beerr Sheva Fringe-fingered Lizard, Acanthodactylus beershebensis (Squamata: Lacertidae) from Israel. Herpetological Bulletin 128: 20-21.

Hawlena, D. 2009. Colorful tails fade when lizards adopt less risky behaviors. Behavioral Ecology and Sociobiology 64(2): 205-213.

Hawlena, D.; Boochnik, R.; Abramsky, Z.; Bouskila, A. 2006. Blue tail and striped body: why do lizards change their infant costume when growing up? Behavioral Ecology 17(6): 889-896.

Hawlena, D.; Pérez-Mellado, V. 2009. Change your diet or die: predator-induced shifts in insectivorous lizard feeding ecology. Oecologia (Berlin) 161(2): 411-419.

Hawlena, D.; Saltz, D.; Abramsky, Z.; Bouskila, A. 2010. Ecological trap for desert lizards caused by anthropogenic changes in habitat structure that favor predator activity. Conservation Biology 24(3): 803-809.

Moravec, J.; Baha el Din, S.; Seligmann, H.; Sivan, N.; Werner, Y.L. 1999. Systematics and distribution of the Acanthodactylus pardalis group (Lacertidae) in Egypt and Israel. Zoology in the Middle East 17: 21-50.

Acanthodactylus blanci

Blanc, C.P. 1980. Studies on the Acanthodactylus of Tunisia. IV. Geographic distribution and habitats. Journal of Herpetology 14(4): 391-398.

Schlüter, U. 2000. Zur Kenntnis von Blancs Fransenfinger (Acanthodactylus blanci). Elaphe 8(3): 70-72.

Acanthodactylus blanfordii

Gardner, A.S. 2005. Blanford's fringe-toed lizard (Lacertidae: Acanthodactylus blanfordii Boulenger, 1918): a new species record for the United Arab Emirates. Tribulus 15(2): 25-26.

Heidari, N.; Faizi, H.; Rastegar-Pouyani, N. 2012. Sexual dimorphism in Blanford's Fringe-toed Lizard, Acanthodactylus blanfordi Boulenger, 1918, from southern Iran (Sauria: Lacertidae). Zoology in the Middle East 55: 35-40.

Heidari, N.; Faizi, H.; Rastegar-Pouyani, N.; Rastegar-Pouyani, E. 2012. A new record of Acanthodactylus cantoris (Sauria: Lacertidae) and its comparison with closely related A. blanfordi in southeastern Iran. Asian Herpetological Research 3(1): 79-82.

Rastegar-Pouyani, N.; Heydari, N.; Faizi, H. 2011. Analysis of geographic variation in four populations of Acanthodactylus blanfordi Boulenger, 1918 (Sauria: Lacertidae) from southern and southeastern Iran. Russian Journal of Herpetology 18(3): 228-234.

Sharma, R.C.; Singh, C. 1976. Records of some little known reptiles from Afghanistan. Newsletter of the Zoological Survey of India 2(2): 70-74.

Acanthodactylus boskianus

Al-Johany, A.M. 1987. Ecology and reproductive biology of Acanthodactylus schmidti in central Arabia. Dissertation Abstracts International C European Abstracts 44(4): 678.

Al-Johany, A.M.; Spellerberg, I.F. 1989. Home range and vagility of the lizards Acanthodactylus schmidti and Acanthodactylus boskianus in Saudi Arabia. Journal of Arid Environments 16(1): 79-86.

Al-Shammari, A.M. 2012. Additional records of lizards in Ha'il Province, Saudi Arabia. Russian Journal of Herpetology 19(4): 287-291.

Amer, S.A.M.; Sallam, F.A.E. 2006. Mitochondrial DNA sequences of the spiny-footed lizards, Acanthodactylus boskianus (Squamata: Lacertidae): systematic implications. Journal of the Egyptian German Society of Zoology 51(C): 1-13.

Angel, F. 1950. Contribution a l'étude de l'Air (Mission L. Chopard et A. Villiers). Lezards. Mem. Inst. Franc. Afr. Noire 10: 331-336.

Baird, D.L.; Crews, D.; Wilczynski, W. 1999. Spatial and reversal learning in congeneric lizards with different foraging strategies. Animal Behaviour 57(2): 393-407.

Bedir, M.A.; El Naggar, M.H.; Kayed, A.N.; Orabi, G.M. 1999. Ecological studies on two lizard species inhabiting Wadi El-Arbein at Saint Katherine area, South Sinai with special reference to biotic elements. Journal of the Egyptian German Society of Zoology (A) 28: 485-503.

Blanc, C.P. 1980. Studies on the Acanthodactylus of Tunisia. IV. Geographic distribution and habitats. Journal of Herpetology 14(4): 391-398.

Blanc, F.; Cariou, M.L.; Blanc, C. 1981. Etudes sur les acantodactyles de Tunisie. 5.- Differenciation genetique et divergence. Amphibia-Reptilia 1(3-4): 329-335.

Böhme, W. 1973. Erstnachweis zweier Eidechsengattungen für die Türkei. Bonner Zoologische Beiträge 24(4): 394-398.

Boulenger, G.A. 1919. On a new variety of Acanthodactylus boskianus Daudin from the Euphrates. Annals and Magazine of Natural History (Ser. 9) 3: 549-550.

Bustard, R. 1959. Lizards from southern Europe. Aquarist and Pondkeeper 24: 70-71.

Calabresi, E. 1923. Missione zoologica del Dr. E. Festa in Cirenaica. Boll. Mus. Zool. Anat. Comp. Univ. Torino 38(7): 1-28.

Cooper, W.E. 1999. Supplementation of phylogenetically correct data by two-species comparison: support for correlated evolution of foraging mode and prey chemical discrimination in lizards extended by first intrageneric evidence. Oikos 87(1): 97-104.

Curtin, N.A.; Woledge, R.C.; Aerts, P. 2005. Muscle directly meets the vast power demands in agile lizards. Proceedings of the Royal Society Biological Sciences Series B 272(1563): 581-584.

Dehlawi, G.Y. 1987. Reproductive activity in the female lizard Acanthodactylus boskianus. Proceedings of the Zoological Society of the Arab Republic Egypt 13: 203-216

Fathinia, B.; Rastegar-Pouyani, N.; Sampour, M.; Bahrami, A.M.; Jaafari, G. 2009. The lizard fauna of Ilam province, Southwestern Iran. Iranian Journal of Animal Biosystematics 5(2): 65-79.

Gilad, A.; Klass, K.; Werner, Y.L. 2012. New methods and old pictures illuminate the status of Acanthodactylus boskianus asper (Audouin, 1827) (Reptilia: Lacertidae). North-Western Journal of Zoology 8(1): 154-163.

Goldberg, S.R. 2013. Reproduction in Bosk's Fringe-fingered Lizard, Acanthodactylus boskianus from Israel (Squamata: Lacertidae). Zoology in the Middle East 59(1): 16-19.

Haas, G.; Battersby, J.C. 1959. Amphibians and reptiles from Arabia. Copeia 1959: 196-202.

Hussein, M.F.; Boulos, R.; Al-Badry, K.S. 1977. Activity of some Egyptian reptiles with reference to light, humidity and temperature. Bulletin of the Faculty of Science Cairo University 46: 313-328.

Hussein, M.F.; Boulos, R.; Al-Badry, K.S. 1977. On the heat death of some Egyptian reptiles. Bulletin of the Faculty of Science Cairo University 46: 299-312.

Kayed, A.N.; Bedir, M.A.; El Naggar, M.H.; Orabi, G.M. 1998. Food consumption, food assimilation and assimilation efficiency of Acanthodactylus boskianus lizard. Journal of the Egyptian German Society of Zoology (A) 25: 273-291.

Khalil, M.W. 2008. Genetic and physiological variabilities among the Egyptian populations of Acanthodactylus boskianus (Lacertidae: Reptilia). Egyptian Journal of Zoology 51: 63-72.

Khannoon, E.; Breithaupt, T.; El-Gendy, A.; Hardege, J.D. 2010. Sexual differences in behavioural response to femoral gland pheromones of Acanthodactylus boskianus. Herpetological Journal 20(4): 225-229.

Mellado, J.; Olmedo, G. 1991. Use of space in Moroccan Sand Lizards, Acanthodactylus (Reptilia, Lacertidae). Journal of Arid Environments 20(3): 339-355.

Mohammed, M.B.H. 1989. Carpus and tarsus patterns in some Arabian lizards (Reptilia: Agamidae, Lacertidae, Scincidae). Folia Morphologica (Prague) 37(2): 201-207.

Nasri, I.; Hammouda, A.; Hamza, F.; Selmi, S. 2015. Sexual size dimorphism in a Tunisian population of Bosk's Fringe-toed Lizards Acanthodactylus boskianus asper. African Journal of Herpetology 64(2): 103-115.

Nazarov, R.A.; Rajabizadeh, M.; Heidari, N.; Faizi, H. 2011. Geographic distribution: Acanthodactylus boskianus (Bose's Fringe-toed Lizard). Herpetological Review 42(4): 567.

Nietzke, G. 1975. Temperaturmessungen im Biotop von Acanthodactylus boscianus. Aquarium Aqua Terra 9(68): 80-81.

Perez-Mellado, V. 1992. Ecology of lacertid lizards in a desert area of eastern Morocco. Journal of Zoology (London) 226(3): 369-386.

Perry, G. 1990. Size and mass in five Israeli lizards (Reptilia: Lacertidae). Israel Journal of Zoology 36(3-4): 129-132.

Perry, G.; Lampl, I.; Lerner, A.; Rothenstein, D.; Shani, E.; Sivan, N.; Werner, Y.L. 1990. Foraging mode in lacertid lizards: variation and correlates. Amphibia-Reptilia 11(4): 373-384.

Rastegar-Pouyani, N. 1999. First record of the lacertid Acanthodactylus boskianus (Sauria: Lacertidae) for Iran. Asiatic Herpetological Research 8: 85-89.

Robson, G.M.; Lambert, M.R.K. 1980. Observations on the insect food of some semi-desert lizards in southern Morocco. Journal of Arid Environments 3(2): 141-151.

Roobas, B.; Feulner, G.R. 2013. A population of Bosk's fringe-toed lizard Acanthodactylus boskianus (Daudin, 1802) in the Hajar Mountain foothills of the UAE. Tribulus 21: 24-37.

Roobas, B.; Feulner, G.R.; Thakur, Y. 2014. Bosk's Fringe-toed Lizard Acanthodactylus boskianus: follow-up study of a population in the Hajar Mountain foothills of the UAE. Tribulus 22: 35-40.

Sarhan, M.; Islam, S.; El-Shehaby, M.; Saber, S. 2015. Molecular phylogenetic relationship among three Acanthodactylus species (Reptilia: Lacertidae) in Egypt based on 18S rRNA sequences. Egyptian Academic Journal of Biological Sciences C Physiology & Molecular Biology 7(1): 39-49.

Seifan, M.; Gilad, A.; Klass, K.; Werner, Y.L. 2009. Ontogenetically stable dimorphism in a lacertid lizard (Acanthodactylus boskianus) with tests of methodology and comments on life-history. Biological Journal of the Linnean Society 97(2): 275-288.

Tamar, K.; Carranza, S.; Sindaco, R.; Moravec, J.; Meiri, S. 2014. Systematics and phylogeography of Acanthodactylus schreiberi and its relationships with Acanthodactylus boskianus (Reptilia: Squamata: Lacertidae). Zoological Journal of the Linnean Society 172(3): 720-739.

Tamar, K.; Maza, E.; Meiri, S. 2013. Natural history notes: Acanthodactylus boskianus (Bosk's Fringe-fingered Lizard). Bifurcation. Herpetological Review 44(1): 135-136.

Trape, J.F.; Trape, S.; Chirio, L. 2012. Lézards, crocodiles et tortues d'Afrique occidentale et du Sahara. IRD Éditions, Marseille. 503 pp.

Üzüm, N.; Ilgaz, C.; Kumlutas, Y.; Cicek, G.; Avci, A. 2014. The body size, age structure, and growth of Bosc's fringe-toed lizard, Acanthodactylus boskianus (Daudin, 1802). Turkish Journal of Zoology 38(4): 383-388.

Volynchik, S. 2014. Climate-related variation in body dimensions within four lacertid species. International Journal of Zoology (Article 795387): 1.

Werner, Y.L. 1986. Geographic sympatry of Acanthodactylus opheodurus with A. boskianus in the Levant. Zoology in the Middle East 1: 92-95.

Werner, Y.L.; Gajst, O.; Talbi, R.; Bouskila, A. 2012. Acanthodactylus opheodurus Arnold, 1980 in the Levant revisited, and the striped patterns of Levantine Acanthodactylus (Reptilia: Lacertidae). Zoology in the Middle East 56: 31-38.

Werner, Y.L.; Ibrahim, A.A. 2012. A specific pholidotic character in Acanthodactylus boskianus boskianus (Daudin, 1802) (Squamata: Sauria: Lacertidae). Herpetozoa 25(1-2): 13-19.

Wilms, T.; Hulbert, F. 2003. Acanthodactylus boskianus (Daudin, 1802). Sauria (Berlin) 25(3): 597-602 (Suppl).

Zari, T.A. 1992. Seasonal variation in time of activity and body temperature of the fringe-toed lizard, Acanthodactylus boskianus. Proceedings of the Zoological Society A.R. Egypt 23(2): 57-66.

Zimmermann, H.; Zimmermann, E. 1983. Durch Nachzucht erhalten: die Fransenfinger-Eidechse Acanthodactylus boskianus. Aquarien Magazin 1983(4): 386-390.

Acanthodactylus busacki

Donaire, D.; Mateo, J.A.; Hasi, M.; Geniez, P. 2000. Nuevos datos sobre la fauna reptiliana de la hamada de Tinduf (Argelia [Algeria]). Boletin de la Asociacion Herpetologica Espanola 11(1): 8-12.

Salvador, A. 1982. A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologische Monographien 16: 1-167.

Schlüter, U. 2002. Zur Kenntnis von Busacks Fransenfinger (Acanthodactylus busacki Salvador, 1982). Elaphe 10(4): 58-61.

Tamar, K.; Geniez, P.; Brito, J.C.; Crochet, P.A. 2017. Systematic revision of Acanthodactylus busacki (Squamata: Lacertidae) with a description of a new species from Morocco. Zootaxa 4276(3): 357–386.

Acanthodactylus cantoris

Agarwal, I.; Goyal, S.P.; Qureshi, Q. 2015. Lizards of the Thar Desert - resource partitioning and community composition. Journal of Arid Environments 118: 58-64.

Bhanotar, R.K.; Srivastava, Y.N. 1985. Impact of reptiles on food production and environment. AISWS Research Publication 2: 116-120, 265-266.

Bons, J.; Girot, B.; Pasteur, G. 1960. Un Acanthodactylus cantoris du Sinai (Sauriens, Lacertidés). Compte Rendus des Seances Mensuelles de la Societe des Sciences Naturelles et Physiques du Maroc 26(1): 18-19.

Heidari, N.; Faizi, H.; Rastegar-Pouyani, N.; Rastegar-Pouyani, E. 2012. A new record of Acanthodactylus cantoris (Sauria: Lacertidae) and its comparison with closely related A. blanfordi in southeastern Iran. Asian Herpetological Research 3(1): 79-82.

Joshi, M.; Das, S.K. 2015. Notes on unusual behavioural ecology of Acanthodactylus cantoris feeding on Bufoniceps laungwalensis. Herpetology Notes 8: 563-565.

Mozaffari, O. 2010. Geographic distribution: Acanthodactylus cantoris (Indian Fringe-fingered Lizard). Herpetological Review 41(4): 510-511.

Pardeshi, M.; Kumar, V.V.; Das, S.K. 2010. Observations on unusual foraging behaviour of Acanthodactylus cantoris Gunther, 1864, in Western Kachchh, Gujarat, India. Journal of the Bombay Natural History Society 106(2): 209-210.

Parshad, B. 1914. Range of Acanthodactylus cantoris, Gunther. Records of the Indian Museum 10: 271-272.

Rehman, H.; Ahmed, S.I.; Fakhri, S. 2002. Home range and growth rate of Fringe-toed Sand Lizard (Acanthodactylus cantoris cantoris) at Hawksbay area, Karachi. Records Zoological Survey of Pakistan 14: 49-54.

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Acanthodactylus micropholis

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Acanthodactylus nilsoni

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Acanthodactylus savignyi

Blanc, F.; Cariou, M.L.; Blanc, C. 1981. Etudes sur les acantodactyles de Tunisie. 5.- Differenciation genetique et divergence. Amphibia-Reptilia 1(3-4): 329-335.

Bons, J. 1968. Addition a la faune du Maroc: Acanthodactylus savignyi savignyi Aud. (Sauria, Lacertidae). Bull. Soc. Sci. Nat. Phys. Maroc 47: 387-394.

Acanthodactylus schmidti

Aldakhil, M.A.; Silva, P.H.D.H. de 1988. The habitat and systematics of Acanthodactylus schmidti Hass (1957) (Sauria, Lacertidae). Journal of the College of Science King Saud University 19(2): 89-95.

Al-Johany, A.M. 1987. Ecology and reproductive biology of Acanthodactylus schmidti in central Arabia. Dissertation Abstracts International C European Abstracts 44(4): 678.

Al-Johany, A.M.; Spellerberg, I.F. 1988. Reproductive biology of the lizard Acanthodactylus schmidti, Weigmann in central Arabia. Journal of Arid Environments 15(2): 197-207.

Al-Johany, A.M.; Spellerberg, I.F. 1989. Home range and vagility of the lizards Acanthodactylus schmidti and Acanthodactylus boskianus in Saudi Arabia. Journal of Arid Environments 16(1): 79-86.

Al-Sadoon, M.K.; Abdo, N.M. 1991. Temperature and body mass effects on the metabolic rate of Acanthodactylus schmidti Weigmann (Reptilia: Lacertidae). Journal of Arid Environments 21(3): 351-361.

Al-Sadoon, M.K.; Al-Otaibi, F.S. 2014. Ecology of the Sand Boa, Eryx jayakari in Riyadh Region of Saudi Arabia. Saudi Journal of Biological Sciences 21(5): 391-393.

Al-Shammari, A.M. 2012. Additional records of lizards in Ha'il Province, Saudi Arabia. Russian Journal of Herpetology 19(4): 287-291.

Haas, G. 1957. Some amphibians and reptiles from Arabia. Proceedings of the California Academy of Sciences (Ser. 4) 29(4): 47-86.

Meinig, H. 1994. Beobachtungen an zwei nur wenig bekannten Fransenfingerarten im Südosten des Oman: Acanthodactylus masirae Arnold, 1980 und Acanthodactylus schmidti Haas, 1957. Die Eidechse 5(11): 24-29.

Meinig, H.; Kessler, H. 1998. Herpetologische Beobachtungen im Rahmen einer Nationalparkplanung: Barr al Hikman und Masirah Island, Sultanat von Oman (Amphibia; Reptilia). Faunistische Abhandlungen (Dresden) 21(Supplement): 89-97.

Ross, W. 1989. Observations on three species of Acanthodactylus (Reptilia: Lacertidae) from eastern Saudi Arabia. Fauna of Saudi Arabia 10: 423-432.

Acanthodactylus schreiberi

Akiki, Y.S.; Saadé, J.H.; Hokayem, M.L.; Hraoui-Bloquet, S.; Nassar, F. 2015. Female reproductive cycle of the lacertid lizard Acanthodactylus schreiberi syriacus (Reptilia: Squamata) from Lebanon. Herpetology Notes 8: 439-443.

Disi, A.M.; Böhme, W. 1996. Zoogeography of the amphibians and reptiles of Syria, with additional new records. Herpetozoa 9(1-2): 63-70.

Franzen, M. 1998. Erstnachweis von Acanthodactylus schreiberi schreiberi Boulenger, 1879 für die Türkei. (Squamata: Sauria: Lacertidae). Herpetozoa 11(1-2): 27-36.

Hraoui-Bloquet, S.; Sadek, R.A.; Lahoud-Hokayem, M.; Hakim-Saade, J. 2007. Cycle de reproduction chez les males d'Acanthodactylus schreiberi syriacus Boettger, 1879 (Reptilia, Lacertidae) du Liban. Bulletin de la Societe Herpetologique de France 122: 5-18.

Lahoud-Hokayem, M.; Hraoui-Bloquet, S.; Sadek, R.A.; Hakim-Saade, J. 2006. Regime alimentaire d'une population d'Acanthodactylus schreiberi syriacus Boettger, 1879 (Reptilia Lacertidae) de la cote sud du Liban. Bulletin de la Societe Herpetologique de France 119: 17-25.

Perry, G. 1990. Size and mass in five Israeli lizards (Reptilia: Lacertidae). Israel Journal of Zoology 36(3-4): 129-132.

Perry, G.; Lampl, I.; Lerner, A.; Rothenstein, D.; Shani, E.; Sivan, N.; Werner, Y.L. 1990. Foraging mode in lacertid lizards: variation and correlates. Amphibia-Reptilia 11(4): 373-384.

Poulakakis, N.; Kapli, P.; Kardamaki, A.; Skourtanioti, E.; Göcmen, B.; Ilgaz, C.; Kumlutas, Y.; Avci, A.; Lymberakis, P. 2013. Comparative phylogeography of six herpetofauna species in Cyprus: late Miocene to Pleistocene colonization routes. Biological Journal of the Linnean Society 108(3): 619-635.

Tamar, K.; Carranza, S.; Sindaco, R.; Moravec, J.; Meiri, S. 2014. Systematics and phylogeography of Acanthodactylus schreiberi and its relationships with Acanthodactylus boskianus (Reptilia: Squamata: Lacertidae). Zoological Journal of the Linnean Society 172(3): 720-739.

Werner, Y.L.; Gajst, O.; Talbi, R.; Bouskila, A. 2012. Acanthodactylus opheodurus Arnold, 1980 in the Levant revisited, and the striped patterns of Levantine Acanthodactylus (Reptilia: Lacertidae). Zoology in the Middle East 56: 31-38.

Yalcinkaya, D.; Göcmen, B. 2012. A new subspecies from Anatolia, Acanthodactylus schreiberi Boulenger, 1879 ataturi n. ssp. (Squamata: Lacertidae). Biharean Biologist 6(1): 19-31.

Zotos, S.; Adamopoulou, C.; Chondropoulos, B.; Kadis, C.; Hadjichambis, A.C.; Legakis, A. 2012. Evidence of sperm storage in Schreiber's Fringe-fingered Lizard, Acanthodactylus schreiberi schreiberi, from Cyprus (Reptilia: Lacertidae). Zoology in the Middle East 56: 39-47.

Acanthodactylus scutellatus

Al-Badry, K.S. 1978. Activity and heat death of the lizard Acanthodactylus scutellatus with reference to the influence of the parietal eye. Proceedings of the Egyptian Academy of Sciences 28[1975]: 139-144.

Al-Hashem, M. 2009. Effects of oil pollution on the population size of the desert lizard Acanthodactylus scutellatus and their prey at Kuwait's Al-Burgan Oil Field. Journal of Biological Sciences 9(7): 682-686.

Al-Hashem, M. 2009. Monitoring population abundance of the Sand Lizard Acanthodactylus scutellatus and their ant prey in oil polluted soils at Kuwait's greater Al-Burgan oil field. Pakistan Journal of Biological Sciences 12(21): 1425-1429.

Al-Hashem, M.; Brain, P.F. 2009. Effects of oil pollution on body size and weight of the Sand Lizard Acanthodactylus scutellatus at the Greater Al-Burgan oil field in Kuwait. Research Journal of Environmental Toxicology 3(1): 56-59.

Al-Hashem, M.A. 2011. Evidence of hepatotoxicity in the sand lizard Acanthodactylus scutellatus from Kuwait's Greater Al-Burgan oil field. Ecotoxicology and Environmental Safety 74(5): 1391-1395.

Al-Hashem, M.A.; Brain, P.F. 2009. Changed substrate preferences shown by fringe-toed Lizards, Acanthodactylus scutellatus, from Kuwait's Al-Burgan oil field (Reptilia: Lacertidae). Zoology in the Middle East 46: 41-45.

Al-Hashem, M.A.; Brain, P.F.; Omar, S.A. 2008. Effects of oil pollution at Kuwait's Greater Al-Burgan oil field on the timing of morning emergence, basking and foraging behaviors by the Sand Lizard Acanthodactylus scutellatus. Pakistan Journal of Biological Sciences 11(4): 589-594.

Baird, D.L.; Crews, D.; Wilczynski, W. 1999. Spatial and reversal learning in congeneric lizards with different foraging strategies. Animal Behaviour 57(2): 393-407.

Bergmans, W. 1964. Acanthodactylus scutellatus Audouin, 1829. Lacerta 22: 36.

Blanc, C.P. 1980. Studies on the Acanthodactylus of Tunisia. IV. Geographic distribution and habitats. Journal of Herpetology 14(4): 391-398.

Blanc, C.P.; Nouira, S. 1987. Etudes sur les acanthodactyles de Tunisie: 8. Les acanthodactyles Meridionaux au sud du Chott El Djerid. Archives de l'Institut Pasteur de Tunis 64(1-2): 89-100.

Blanc, F.; Cariou, M.L.; Blanc, C. 1981. Etudes sur les acantodactyles de Tunisie. 5.- Differenciation genetique et divergence. Amphibia-Reptilia 1(3-4): 329-335.

Bons, J.; Girot, B. 1963. Revision de l'espèce Acanthodactylus scutellatus (Lacertidae, Saurien). Bull. Soc. Sci. Nat. Phys. Maroc 42: 311-334.

Busack, S.D. 1959. Methods of eating and general behavior in some Old World saurians in captivity. Herpetologica 15(4): 234.

Cooper, W.E. 1999. Supplementation of phylogenetically correct data by two-species comparison: support for correlated evolution of foraging mode and prey chemical discrimination in lizards extended by first intrageneric evidence. Oikos 87(1): 97-104.

Crochet, P.A.; Geniez, P.; Ineich, I. 2003. A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): Systematic and biogeographical implications. Zoological Journal of the Linnean Society 137(1): 117-155.

El Hennawy, H.K. 1990. Arachnida in the diet of Acanthodactylus scutellatus (Audouin, 1825) (Reptilia: Lacertidae). Serket 2(1): 1-8.

Geniez, P.; Lluch, P. 2003. New data on fringe-toed lizards of the Acanthodactylus scutellatus group in Mauritania. Herpetozoa 17(1-2): 88-89.

Gramentz, D. 2002. Zur Ökologie und zum Verhalten von Acanthodactylus scutellatus (Audouin, 1809) in West-Tunesien. Die Eidechse 13(1): 14-21.

Ibrahim, A.A.; Saleh, M.A.; Dixon, J.R.; Mahmoud, I.M. 2000. On the ecology of the fringe-toed lizard, Acanthodactylus scutellatus (Sauria: Lacertidae) in north Sinai, Egypt. Journal of the Egyptian German Society of Zoology 32(A): 335-355.

Kroniger, M. 1995. Zur Haltung von Acanthodactylus scutellatus (Audouin, 1809). Die Eidechse 6(14): 1-3.

Mellado, J.; Olmedo, G. 1990. El genero Acanthodactylus en Marruecos: problemas de identification en los grupos de especies A. pardalis y A. scutellatus. Amphibia-Reptilia 11(2): 131-146.

Mellado, J.; Olmedo, G. 1991. Use of space in Moroccan Sand Lizards, Acanthodactylus (Reptilia, Lacertidae). Journal of Arid Environments 20(3): 339-355.

Mohammed, M.B.H. 1989. Carpus and tarsus patterns in some Arabian lizards (Reptilia: Agamidae, Lacertidae, Scincidae). Folia Morphologica (Prague) 37(2): 201-207.

Perez-Mellado, V. 1992. Ecology of lacertid lizards in a desert area of eastern Morocco. Journal of Zoology (London) 226(3): 369-386.

Perry, G. 1990. Size and mass in five Israeli lizards (Reptilia: Lacertidae). Israel Journal of Zoology 36(3-4): 129-132.

Perry, G.; Dmi'el, R. 1994. Needles and haystacks: the location of lizard eggs in sand dunes. Amphibia-Reptilia 15(4): 395-401.

Perry, G.; Dmi'el, R. 1994. Reproductive and population biology of the fringe-toed lizard, Acanthodactylus scutellatus, in Israel. Journal of Arid Environments 27(3): 257-263.

Perry, G.; Lampl, I.; Lerner, A.; Rothenstein, D.; Shani, E.; Sivan, N.; Werner, Y.L. 1990. Foraging mode in lacertid lizards: variation and correlates. Amphibia-Reptilia 11(4): 373-384.

Sarhan, M.; Islam, S.; El-Shehaby, M.; Saber, S. 2015. Molecular phylogenetic relationship among three Acanthodactylus species (Reptilia: Lacertidae) in Egypt based on 18S rRNA sequences. Egyptian Academic Journal of Biological Sciences C Physiology & Molecular Biology 7(1): 39-49.

Schlüter, U. 2000. Ägyptische Dunenfransenfinger. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 53(2): 30-31.

Scortecci, G. 1940. Biologia del Tibesti. Ann. Mus. libico Stor. nat. 2: 257-275.

Scortecci, G. 1946. Tentativo di analisi biologica condotto sulla specie Acanthodaclylus scutellatus Audouin. Riv. Biol. Colon 7(1): 6-16.

Subach, A.; Scharf, I.; Ovadia, O. 2009. Foraging behavior and predation success of the sand viper (Cerastes vipera). Canadian Journal of Zoology 87(6): 520-528.

Acanthodactylus spinicaudus

Dellaoui, B.; Beddek, M.; Peyre, O.; Geniez, P.; Allegrini, B.; Koudache, F.; Crochet, P.A. 2015. Rediscovery of Acanthodactylus spinicauda Doumergue, 1901 in Algeria. Herpetology Notes 8: 511-515.

Acanthodactylus taghitensis

Crochet, P.A.; Geniez, P.; Ineich, I. 2003. A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): Systematic and biogeographical implications. Zoological Journal of the Linnean Society 137(1): 117-155.

Donaire, D.; Mateo, J.A.; Hasi, M.; Geniez, P. 2000. Nuevos datos sobre la fauna reptiliana de la hamada de Tinduf (Argelia [Algeria]). Boletin de la Asociacion Herpetologica Espanola 11(1): 8-12.

Geniez, P.; Foucart, A. 1995. Un nouvel acanthodactyle en Algerie: Acanthodactylus taghitensis n. sp. (Reptilia, Sauria, Lacertidae). Bulletin du Museum National d'Histoire Naturelle Section A Zoologie Biologie et Ecologie Animales 17(1-2): 3-9.

Geniez, P.; Lluch, P. 2003. New data on fringe-toed lizards of the Acanthodactylus scutellatus group in Mauritania. Herpetozoa 17(1-2): 88-89.

Hasi, M.; Lopez-Jurado, L.F.; Mateo, J.A.; Saint Andrieux, J.P.; Geniez, P. 1997. Nouvelles observations herpetologiques au Sahara Occidental, 3. Bulletin de la Societe Herpetologique de France 84: 33-38.

Acanthodactylus tilburyi

Arnold, E.N. 1986. A new spiny-footed lizard (Acanthodactylus: Lacertidae) from Saudi Arabia. Fauna of Saudi Arabia 8: 378-384.

Modry, D.; Al-Oran, R.M.; Amr, Z.S.; Necas, P. 1999. A new record of the Tilbury's spiny-footed lizard, Acanthodactylus tilburyi Arnold, 1986 (Reptilia: Lacertidae), from the Hashemite Kingdom of Jordan. Casopis Narodniho Muzea Rada Prirodovedna 168(1-4): 121-124.

Acanthodactylus yemenicus

Salvador, A. 1982. A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zoologische Monographien 16: 1-167.