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Taxonomy of the genus Chelonoidis Biodiversity of the family Testudinidae

Bibliography of the genus
Chelonoidis (South American Tortoises)

(Reptilia: Testudines: Testudinidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Chelonoidis in general

Arroyave Bermudez, F.J.; Romero Goyeneche, O.Y.; Bonilla Gómez, M.A.; Hurtado Heredia, R.G. 2014. [Illegal trade of tortoises (Testudinata) in Colombia: a network analysis approach]. (In Spanish). Acta Biologica Colombiana 19(3): 381-392.

Bastille-Rousseau, G.; Gibbs, J.P.; Yackulic, C.B.; Frair, J.L.; Cabrera, F.; Rousseau, L.P.; Wikelski, M.; Kuemmeth, F.; Blake, S. 2017. Animal movement in the absence of predation: environmental drivers of movement strategies in a partial migration system. Oikos 126(7): 1004-1019

Beheregaray, L.B.; Ciofi, C.; Geist, D.; Gibbs, J.P.; Caccone, A.; Powell, J.R. 2003. Genes record a prehistoric volcano eruption in the Galapagos. Science (Washington, D.C.) 302(5642): 75.

Bergen, T. 2013. Decline and rise of Galápagos tortoises. IRCF Reptiles & Amphibians 20(1): 23–29.

Blake, S.; Guezou, A.; Deem, S.L.; Yackulic, C.B.; Cabrera, F. 2015. The dominance of introduced plant species in the diets of migratory Galapagos Tortoises increases with elevation on a human-occupied island. Biotropica 47(2): 246-258.

Cayot, L.; Snell, H.L.; Llerena, W.; Snell, H.M. 1994. Conservation biology of Galapagos reptiles: twenty-five years of successful research and management. pp. 297-305. In: Murphy, J.B.; Adler, K. & Collins, J.T. (eds.). Captive management and conservation of amphibians and reptiles. SSAR (Society for the Study of Amphibians and Reptiles). 408 pp. (Contributions to Herpetology no. 11).

Chiari, Y.; Hyseni, C.; Fritts, T.H.; Glaberman, S.; Marquez, C.; Gibbs, J.P.; Claude, J.; Caccone, A. 2009. Morphometrics parallel genetics in a newly discovered and endangered taxon of Galapagos tortoise. PLoS ONE 4(7): e6272, 1-11.

Ciofi, C.; Wilson, G.A.; Beheregaray, L.B.; Marquez, C.; Gibbs, J.P.; Tapia, W.; Snell, H.L.; Caccone, A.; Powell, J.R. 2006. Phylogeographic history and gene flow among giant Galapagos tortoises on southern Isabela Island. Genetics 172(3): 1727-1744.

Denburgh, J. van 1998. The Gigantic Land Tortoises of the Galapagos Archipelago. Society for the Study of Amphibians and Reptiles, Ithaca. 290 pp.

Edwards, D.L.; Garrick, R.C.; Tapia, W.; Caccone, A. 2014. Cryptic structure and niche divergence within threatened Galapagos giant tortoises from southern Isabela Island. Conservation Genetics 15(6): 1357-1369.

Ellis-Soto, D.; Blake, S.; Soultan, A.; Guezou, A.; Cabrera, F.; Lötters, S. 2017. Plant species dispersed by Galapagos tortoises surf the wave of habitat suitability under anthropogenic climate change. PLoS ONE 12(7): e0181333

Farias, I.P.; Jerozolimski, A.; Melo, A.; Neves Viana, M. das; Martins, M.; Santos Monjelo, L.A. dos 2007. Population genetics of the Amazonian tortoises, Chelonoidis denticulata and C. carbonaria, (Cryptodira: Testudinidae) in an area of sympatry. Amphibia-Reptilia 28(3): 357-365.

Garrick, R.C.; Kajdacsi, B.; Russello, M.A.; Benavides, E.; Hyseni, C.; Gibbs, J.P.; Tapia, W.; Caccone, A. 2015. Naturally rare versus newly rare: demographic inferences on two timescales inform conservation of Galapagos giant tortoises. Ecology and Evolution 5(3): 676-694.

Gerlach, J. 1998. Famous Tortoises. Nature Protection Trust of the Seychelles, Cambridge. 51 pp.

Hunter, E.A.; Gibbs, J.P. 2014. Densities of ecological replacement herbivores required to restore plant communities: a case study of giant tortoises on Pinta Island, Galapagos. Restoration Ecology 22(2): 248-256.

Knaack, J. 2006. Beobachtungen zum Vorkommen von Geochelone (Chelonoidis) im südlichen Bolivien. Reptilia (D) 11(59): 77-82.

Marquez, C.; Wiedenfeld, D.A.; Landazuri, S.; Chavez, J. 2007. Human-caused and natural mortality of giant tortoises in the Galapagos Islands during 1995-2004. Oryx 41(3): 337-342.

McCord, W.P.; Joseph-Ouni, M. 2004. Chelonian illustrations. 14. North and South American tortoises. Reptilia (GB) 34: 47-50.

Olson, S.L.; David, N. 2014. The gender of the tortoise genus Chelonoidis Fitzinger, 1835 (Testudines: Testudinidae). Proceedings of the Biological Society of Washington 126(4): 393-394.

Papa, L.M. del 2015. Utilización de reptiles durante el periodo Agroalfarero de la región Chaco-Santiagueña, Argentina. Archaeofauna 24: 7-26.

Patton, J.L. 1984. Genetical processes in the Galapagos. Biological Journal of the Linnean Society 21(1-2): 97-111.

Poulakakis, N.; Glaberman, S.; Russello, M.; Beheregaray, L.B.; Ciofi, C.; Powell, J.R.; Caccone, A. 2008. Historical DNA analysis reveals living descendents of an extinct species of Galapagos tortoise. Proceedings of the National Academy of Sciences of the United States of America 105(40): 15464-15469.

Renaud, M.L.; Gitschlag, G.R.; Hale, J.K. 1993. Retention of imitation satellite transmitters fiberglassed to the carapace of Sea Turtles. Herpetological Review 24(3): 94-99.

Rothschild, Lord 1925. Giant tortoises of the Galapagos and Mascarene Islands. Proceedings of the Linnean Society of London 137: 40.

Sachs, T. 2014. Lonesome George, icon of extinction. The Tortoise 1(3): 174-179.

Sims, J. 1991. Captive care. South American tortoises. Rephiberary 168: 4-5.

Sims, J. 1991. South American tortoises. Captive conditions part 2. Rephiberary 169: 4-6.

Stanford, C.B. 2010. The Last Tortoise: A Tale of Extinction in Our Lifetime. Belknap Press, Cambridge, Massachusetts. 210 pp.

Sulloway, F.J. 1984. Darwin and the Galapagos. Biological Journal of the Linnean Society 21(1-2): 29-59.

Wüthrich, F. 2006. Die Geschichte der Galápagos-Riesenschildkröte Harriet. Testudo 15(4): 15-27.

Chelonoidis abingdonii

Anonymous. 2012. Pinta Island, Ecuador. 6. A Galapagos legend dies. Science (Washington D C) 336(6089): 1625.

Boulenger, E.G. 1914. Giant saddle-backed tortoise. Proceedings of the Zoological Society of London 1914: 220-222.

Cayot, L.J.; Gibbs, J.P.; Tapia, W.; Caccone, A. 2016. Chelonoidis abingdonii. The IUCN Red List of Threatened Species 2016: e.T9017A65487433. http://dx.doi.org/10.2305/IUCN.UK.2016-1.RLTS.T9017A65487433.en.

Dowling, H.G. 1961. Vanishing giants and enduring dwarfs - the tortoises. Animal Kingdom 64: 66-75.

Edwards, D.L.; Benavides, E.; Garrick, R.C.; Gibbs, J.P.; Russello, M.A.; Dion, K.B.; Hyseni, C.; Flanagan, J.P.; Tapia, W.; Caccone, A. 2013. The genetic legacy of Lonesome George survives: giant tortoises with Pinta Island ancestry identified in Galapagos. Biological Conservation 157: 225-228.

Garrick, R.C.; Benavides, E.; Russello, M.A.; Gibbs, J.P.; Poulakakis, N.; Dion, K.B.; Hyseni, C.; Kajdacsi, B.; Márquez, L.D.; Bahan, S.; Ciofi, C.; Tapia Aguilera, W.; Caccone, A. 2012. Genetic rediscovery of an 'extinct' Galápagos giant tortoise species. Current Biology 22(1): R10-11.

Jones, J. 2013. Lonesome George: the legacy. Testudo 7(5): 69-78.

Nicholls, H. 2006. Lonesome George and the origins and evolution of Galapagos giant tortoises. Testudo 6(3): 64-73.

Nicholls, H. 2006. Lonesome George: the life and loves of a conservation icon. Macmillan, London & New York. 231 pp.

Philippen, H.D. 2012. [The Pinta Island tortoise Chelonoidis (nigra) abingdoni (Günther, 1877)]. (In German). Marginata 35: 31-34.

Pritchard, P.C.H. 1977. Three, two, one tortoise. Natural History (New York) 86(8): 91-100.

Pritchard, P.C.H. 2005. The Pinta Tortoise: Globalization and the Extinction of Island Species. Global Connections Foundation, Orlando, Florida. 48 pp.

Rowley, A. 2012. Restoration of Pinta Island through the repatriation of giant tortoises. Testudo 7(4): 50-66.

Chelonoidis becki

Emerson, B.C.; Faria, C.M.A. 2014. Fission and fusion in island taxa - serendipity, or something to be expected. Molecular Ecology 23(21): 5132-5134.

Garrick, R.C.; Benavides, E.; Russello, M.A.; Gibbs, J.P.; Poulakakis, N.; Dion, K.B.; Hyseni, C.; Kajdacsi, B.; Márquez, L.D.; Bahan, S.; Ciofi, C.; Tapia Aguilera, W.; Caccone, A. 2012. Genetic rediscovery of an 'extinct' Galápagos giant tortoise species. Current Biology 22(1): R10-11.

Garrick, R.C.; Benavides, E.; Russello, M.A.; Hyseni, C.; Edwards, D.L.; Gibbs, J.P.; Tapia, W.H.; Ciofi, C.; Caccone, A. 2014. Lineage fusion in Galapagos Giant Tortoises. Molecular Ecology 23(21): 5276-5290.

IUCN Tortoise & Freshwater Turtle Specialist Group. 2016. Chelonoidis becki. The IUCN Red List of Threatened Species 2016: e.T9018A97225095. http://dx.doi.org/10.2305/IUCN.UK.2016-1.RLTS.T9018A97225095.en.

Marquez, C. 1986. Herpetology program of the CDRS. Estacion Cientifica Charles Darwin Informe Anual 1983: 28-29.

Chelonoidis carbonarius

Anonymous. 1994. Hatching red-foots. Tropical Fish Hobbyist 43(4): 128-129.

Aponte, C.; Barreto, G.R.; Terborgh, J. 2003. Consequences of habitat fragmentation on age structure and life history in a tortoise population. Biotropica 35(4): 550-555.

Auffenberg, W. 1965. Sex and species discrimination in two sympatric South American tortoises. Copeia 1965: 335-342.

Bakowsky, R. 2014. [Successful care of Red-footed Tortoises in Venezuela]. (In German). Marginata 42: 36-45.

Barros, M.S.; Resende, L.C.; Silva, A.G.; Ferreira-Júnior, P.D. 2012. Morphological variations and sexual dimorphism in Chelonoidis carbonaria (Spix, 1824) and Chelonoidis denticulata (Linnaeus, 1766) (Testudinidae). Brazilian Journal of Biology 72(1): 153-161.

Bernardo, P.H.; Guerra-Fuentes, R.A.; Matiazzi, W.; Zaher, H. 2012. Checklist of amphibians and reptiles of Reserva Biológica do Tapirapé, Pará, Brazil. Check List 8(5): 839-846.

Bertoni, A. de W. 1926. Nuevos quelonios Paraguayos. Rev. Soc. Cien. Paraguay Asuncion 2(1): 71.

Bjorndal, K.A. 1989. Flexibility of digestive responses in two generalist herbivores, the tortoises Geochelone carbonaria and Geochelone denticulata. Oecologia (Berlin) 78(3): 317-321.

Borini, J.F.; Petrucci, B.B.; Krohling, W.; Rossi Junior, J.L.; Santos, M.R.D.; Ferreira-Júnior, P.D. 2014. Site fidelity and movement of Chelonoidis carbonaria (Spix, 1824) (Testudinidae) in cocoa plantations in southeastern Brazil. Brazilian Journal of Biology (Suppl. S) 74(3): 135-141.

Buchert, P. 2011. Köhlerschildkröten - begehrte, aber anspruchsvolle Pfleglinge. Elaphe 19(3): 44-50.

Castano Mora, O.V. 1985. Notas adicionales sobre la reproduccion yel crecimiento de los morrocoyes (Geochelone carbonaria y G. denticulata, Testudines, Testudinidae). Lozania 52: 1-5.

Castano Mora, O.V.; Lugo Rugeles, M. 1981. Estudio comparativo del comportamiento de dos especies de morrocoy: Geochelone carbonaria y Geochelone denticulata y aspectos comparables de su morfologia externa. Cespedesia 10(37-38): 55-122.

Censky, E.J. 1988. Geochelone carbonaria (Reptilia: Testudines) in the West Indies. Florida Scientist 51(2): 108-114.

Colvee, S. 2002. Reproductive aspects of the redfoot tortoise in captivity. Reptilia (GB) 21: 58-66.

Conceicao, A.M.; Almeida, B.M.; Brasileiro, M.T.R.; Barros, L.E.D.; Wiltshire, S.A.; Sarmento, C.A.P. 2009. Hermafroditismo em jabuti piranga (Geochelone carbonaria). Arquivo Brasileiro de Medicina Veterinaria e Zootecnia 61(6): 1478-1481.

Cunha, O.R. da; Nascimento, F.P. do; Avila Pires, T.C.S. 1985. Os repteis da area de Carajas, Para, Brasil (Testudines e Squamata). 1. Museu Paraense Emilio Goeldi Publicacoes Avulsas 40: 9-92.

Czybulinski, R. 2015. Die Köhlerschildkröte Chelonoidis carbonarius – Faszination, Pflege und Zucht. Kleintierverlag Thorsten Geier, Biebertal. 161 pp.

Dathe, F. 2011. Haltung und Vermehrung von Köhlerschildkröten, Chelonoidis carbonaria (Spix, 1824), im Tierpark Berlin. Milu 13(3): 413-425.

Davis, S. 1979. Husbandry and breeding of the red-footed tortoise Geochelone carbonaria at the National Zoological Park, Washington. International Zoo Yearbook 19: 50-53.

Dowling, H.G. 1961. Vanishing giants and enduring dwarfs - the tortoises. Animal Kingdom 64: 66-75.

Dubois, B.; Vuylsteker, J. 2007. Elevage et reproduction en captivite de la Tortue charbonniere, Chelonoidis carbonaria (Spix, 1824). Cheloniens 5: 38-44.

Ernst, C.H.; Leuteritz, T.E.J. 1999. Geochelone carbonaria. Catalogue of American Amphibians and Reptiles 690: 1-7.

Fehringer, P. 2010. Köhlerschildkröten (Chelonoidis carbonaria ssp.) auf den Kleinen Antillen in der Natur beobachtet. Marginata 27: 16-19.

Fehringer, P. 2010. Köhlerschildkröten auf der Insel Barbuda - eine ungewöhnliche Erfahrung bei einem Kurzaufenthalt. Marginata 27: 28-29.

Flanigan, W.F. 1974. Sleep and wakefulness in chelonian reptiles. 2. The red-footed tortoise, Geochelone carbonaria. Archives ital. Biol. 112(3): 253-277.

Fritz, C. 2001. Erfolgreiche Haltung und Vermehrung der Köhlerschildkröte (Geochelone carbonaria) aus dem nördlichen Südamerika. Draco 2(4) (8): 72-77.

Gorzula, S. 1989. Recent population changes: Chelonoidis carbonaria (red footed tortoise). Herpetological Review 20(2): 56.

Guix, J.C.; Fedullo, D.L.; Molina, F.B. 2001. Masculinization of captive females of Chelonoidis carbonaria (Testudinidae). Revista Española de Herpetología 15: 67-75.

Guix, J.C.C.; Salvatti, M.; Peroni, M.A.; Lima-Verde, J.S. 1989. Observations additionnelles sur le comportement et la reproduction de Chelonoidis carbonaria (Spix, 1824) en captivite (Testudines-Testudinidae). Grupo de Estudos Ecologicos Serie Documentos 1: 20-31.

Gutsche, A. 2012. Klassiker: Die Köhlerschildkröte. Terraria-Elaphe 2012(38): 68-69.

Hedman, H.D.; Muniz, D.N.; Powell, R. 2010. Natural history notes: Chelonoidis carbonaria (Red-footed Tortoise). Size and thermal biology. Herpetological Review 41(4): 484-485.

Hernandez, O.; Boede, E.O. 2002. Efectos de la densidad y la proporcion de sexos en la reproduccion en cautiverio del Morrocoy Geochelone (Chelonoidis) carbonaria (Spix 1824). Acta Biologica Venezuelica 21(2): 29-37.

Hernández-Montoya, V.; Páez, V.P.; Ceballos, C.P. 2017. Effects of temperature on sex determination and embryonic development in the Red-footed Tortoise, Chelonoidis carbonarius. Chelonian Conservation and Biology 16(2): 164-171.

Hernandez-P., O.E. 1997. Reproduccion y crecimiento del morrocoy, Geochelone (Chelonoidis) carbonaria (Spix, 1824) (Reptilia, Testudinidae). BioLlania 13: 165-183.

Hernandez-P., O.E.; Boede, E.O. 2000. Efecto de la alimentacion sobre el crecimiento y produccion de huevos de Geochelone (Chelonoidis) carbonaria (Spix, 1824) bajo condiciones de cautiverio. Acta Biologica Venezuelica 20(2): 37-43.

Herz, M. 2015. [Encounters with Testudines in the Venezuelan state of Amazonas. Part 2: Tortoises]. (In German). Reptilia (D) 20(111): 64-68.

Hoogmoed, M.S.; Gruber, U. 1983. Spix and Wagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (the Netherlands). Spixiana Supplement 9: 319-415.

Houbrix, P. 2007. Elevage et reproduction en captivite de la tortue charbonniere, Chelonoidis carbonaria (Spix, 1824). Manouria 10(36): 12-18.

Houtman, H.; Fosse, A. de la 1988. Care and breeding of Geochelone carbonaria. Lacerta 47(2) [1988-1989]: 40-46.

Knaack, J. 2004. Köhler- und Waldschildkröte. Beobachtungen zum Vorkommen von Geochelone (Chelonoides) carbonaria und G. (C.) denticulata in Bolivien. Aquarium (Bornheim) 416: 70-74.

Kradolfer, K.D. 2004. Beobachtungen an Köhlerschildkröten während des Freilaufs im Garten. Testudo 13(1): 37-40.

Kundert, S. 2004. Bemerkungen zur Bestimmung der Herkunft von Köhlerschildkröten aufgrund der Plastrongefärbung. Testudo 13(1): 41-42.

Kundert, S. 2004. Haltung un Vermehrung der Köhlerschildkröte Geochelone carbonaria (Spix 1824). Testudo 13(1): 21-36.

Legler, J.M. 1963. Tortoises (Geochelone carbonaria) in Panama: distribution and variation. American Midland Naturalist 70: 490-503.

Lowe, P. 1997. Redfoot tortoises: South American treasure. Reptile & Amphibian Magazine 46: 36-43.

Marcellini, D.L.; Davis, S.W. 1982. Effects of handling on reptile egg hatching. Herpetological Review 13(2): 43-44.

May, C. 2004. Pflege und Vermehrung der rotköpfigen Köhlerschildkröte "Cherryhead", Geochelone carbonaria. Schildkröten im Fokus 1(4): 3-12.

May, C. 2012. An incident of triplets in the Red-footed Tortoise, Chelonoidis carbonaria (Spix, 1824). Schildkröten im Fokus 9(1): 3-12.

May, C.D.; Bidmon, H.J.; Buskirk, J.R.; Vinke, S. 2010. Über das Auftreten eines Nackenschildes bei einer Südamerikanischen Landschildkröte, Chelonoidis carbonaria. Schildkröten im Fokus 7(2): 11-20.

McBride, R.; Giordano, A.; Ballard, W.B. 2010. Note on the winter diet of jaguars Panthera onca in the Paraguayan transitional chaco. Bellbird 4 [2009-2010]: 4 pp.

McKeown, S. 1997. How to repair and hatch damaged turtle eggs. Vivarium (Lakeside) 8(5): 6-7.

Medem, F.; Castano, O.V.; Lugo, R.M. 1979. Contribucion al conocimiento sobre la reproducion y el creamiento de los 'morrocoyes' (Geochelone carbonaria y G. denticulata; Testudines, Testudinidae). Caldasia (Bogota) 12(59): 497-511.

Merchan, M.; Coll, M.; Gil, J.R. 2002. Confirman la presencia de tortugas carboneras silvestres en Nicaragua. Quercus 193: 44-45.

Merchan, M.; Fidalgo, A.M.; Perez, C. 1998. Biology, distribution and conservation of the redfoot tortoise Geochelone carbonaria. Reptilia (GB) 2: 30-38.

Merchan, M.; Fidalgo, A.M.; Perez, C. 1999. Biologie, Verbreitung und Schutz der Köhlerschildkröte Geochelone carbonaria. Reptilia (D) 4(20): 56-63.

Metrailler, S. 1997. Geochelone carbonaria. Reptilia (D) 2(3): 53-55.

Mislin, V. 2010. Nahezu 40 Jahre Schildkrötenhaltung und -zucht am Biespiel der Köhlerschildkröte (Chelonoidis carbonaria). Marginata 27: 20-27.

Moll, D.; Tucker, J.K. 1976. Growth and sexual maturity of the red-footed tortoise, Geochelone carbonaria. Bulletin of the Maryland Herpetological Society 12(3): 96-98.

Montes-Correa, A.C.; Saboya-Acosta, L.P.; Páez, V.P.; Vega, K.; Renjifo, J.M. 2014. [Distribution of tortoises and freshwater turtles of the Colombian Caribbean]. (In Spanish). Acta Biologica Colombiana 19(3): 341-350.

Moreira, G. 1991. Observacoes sobre Geochelone denticulata (Linnaeus, 1766) e Geochelone carbonaria (Spix, 1824) na bacia do Rio Uatuma, Amazonia central. Boletim do Museu Paraense Emilio Goeldi (Zoologia) 7(2): 183-188.

Moreira, G.R.S. 1989. Sympatry of the turtles Geochelone carbonaria and G. denticulata in the Rio Uatuma Basin, central Amazonia. Journal of Herpetology 23(2): 183-185.

Moskovits, D. 1998. Population and ecology of the tortoises Geochelone carbonaria and G. denticulata on the Ilha de Maraca. pp. 263-284. In: Milliken, W. & Ratter, J.A. (eds.). Maraca: the biodiversity and environment of an Amazonian rainforest. John Wiley & Sons, Chichester, New York, etc. 508 pp.

Moskovits, D.K. 1988. Sexual dimorphism and population estimates of the two Amazonian tortoises (Geochelone carbonaria and G. denticulata) in northwestern Brazil. Herpetologica 44(2): 209-217.

Moskovits, D.K.; Bjorndal, K.A. 1990. Diet and food preferences of the tortoises Geochelone carbonaria and G. denticulata in northwestern Brazil. Herpetologica 46(2): 207-218.

Moskovits, D.K.; Kiester, A.R. 1987. Activity levels and ranging behaviour of the two Amazonian tortoises, Geochelone carbonaria and Geochelone denticulata, in north-western Brazil. Functional Ecology 1(3): 203-214.

Mourthé, Í. 2016. Natural history notes: Chelonoidis carbonarius (Red-footed Tortoise). Antipredator behavior. Herpetological Review 47(4): 651-652.

Mourthé, Í.; Castro, A. 2017. Natural history notes: Chelonoidis carbonaria (Red-footed Tortoise). Scavenging behavior. Herpetological Review 48(2): 422-423.

Mueller-Paul, J.; Wilkinson, A.; Aust, U.; Steurer, M.; Hall, G.; Huber, L. 2014. Touchscreen performance and knowledge transfer in the red-footed tortoise (Chelonoidis carbonaria). Behavioural Processes 106: 187-192.

Mueller-Paul, J.; Wilkinson, A.; Hall, G.; Huber, L. 2012. Radial-arm-maze behavior of the Red-footed Tortoise (Geochelone carbonaria). Journal of Comparative Psychology 126(3): 305-317.

Müller, P. 1971. Beobachtungen an brasilianischen Geochelone carbonaria. Aqua Terra 8: 69-75.

Netten, H.; Zuurmond, F. 1985. Geochelone carbonaria in captivity. Lacerta 43(5): 84-89.

Nietzke, G. 1974. Testudo carbonaria - Köhlerschildkröten. Aquarium Aqua Terra 8(55): 33-34.

Nijs, J.; Zwartepoorte, H.; Boonstra, J. 1996. Divergent incubation times in clutches of Chelonia incubated under the same temperature conditions. Bulletin of the Chicago Herpetological Society 31(10): 187.

Noss, A.J.; Montaño, R.R.; Soria, F.; Deem, S.L.; Fiorello, C.V.; Fitzgerald, L.A. 2013. Chelonoidis carbonaria (Testudines: Testudinidae) activity patterns and burrow use in the Bolivian Chaco. South American Journal of Herpetology 8(1): 19-28.

Orchard, K. 2010. Chelonoidis carbonaria (Red-footed Tortoise). Distribution. Caribbean Herpetology 11: 1.

Philippen, H.D. 1990. Köhlerschildkröten vor der regionalen Ausrottung. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 43: 584.

Philippen, H.D. 2010. Albino-Köhlerschildkröten (Chelonoidis carbonaria) - eine häufiger auftretende Mutation. Marginata 27: 30.

Philippen, H.D. 2010. Die Köhlerschildkröte (Chelonoidis carbonaria Spix, 1824). Marginata 27: 10-15.

Pingleton, M. 2006. Red-footed Tortoises: natural history, behavior and their influences on captive care. Proceedings of the Annual Conference of the Association of Reptilian and Amphibian Veterinarians 13: 26-31.

Pritchard, P.C.H. 1980. Record size turtles from Florida and South America. Chelonologica 1(3): 113-123.

Richard, E.; Nakhle, J. 1989. Geographic distribution: Chelonoidis carbonaria (South American Red-footed Tortoise). Herpetological Review 20(1): 14.

Rogner, M. 2012. [The red-footed tortoise. Care and breeding of a tropical beauty]. (In German). Reptilia (D) 17(97): 58-62.

Salazar-Saavedra, M.; Loza, J.C.; Fernández, M.; Martínez-Fonseca, J.G.; Dwyer, Q.; Sunyer, J. 2015. Miscellaneous notes: Chelonoidis carbonarius (Spix, 1824): a member of the Nicaraguan herpetofauna. Mesoamerican Herpetology 2(4): 571-573.

Schall, O. 1978. Das Schildkrötenportrat. Die Köhlerschildkröte. Aquarien Magazin 12(5): 260.

Schlegel, H. 1968. Testudo carbonaria, omnivore oder Aasfreser. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 21: 52-55.

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Chelonoidis denticulatus

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Chelonoidis phantasticus

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Chelonoidis porteri

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Chelonoidis vicina

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