Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus Recent updates Related bibliographies: Reptiles Turtles Testudinidae Central America North America South America West Indies Pacific Ocean Taxonomy of the genus Chelonoidis Biodiversity of the family Testudinidae Bibliography of the genus Chelonoidis (South American Tortoises) (Reptilia: Testudines: Testudinidae) Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted. Chelonoidis in general Arroyave Bermudez, F.J.; Romero Goyeneche, O.Y.; Bonilla Gómez, M.A.; Hurtado Heredia, R.G. 2014. [Illegal trade of tortoises (Testudinata) in Colombia: a network analysis approach]. (In Spanish). Acta Biologica Colombiana 19(3): 381-392. Bastille-Rousseau, G.; Gibbs, J.P.; Yackulic, C.B.; Frair, J.L.; Cabrera, F.; Rousseau, L.P.; Wikelski, M.; Kuemmeth, F.; Blake, S. 2017. Animal movement in the absence of predation: environmental drivers of movement strategies in a partial migration system. Oikos 126(7): 1004-1019 Beheregaray, L.B.; Ciofi, C.; Geist, D.; Gibbs, J.P.; Caccone, A.; Powell, J.R. 2003. Genes record a prehistoric volcano eruption in the Galapagos. Science (Washington, D.C.) 302(5642): 75. Bergen, T. 2013. Decline and rise of Galápagos tortoises. IRCF Reptiles & Amphibians 20(1): 23–29. Blake, S.; Guezou, A.; Deem, S.L.; Yackulic, C.B.; Cabrera, F. 2015. The dominance of introduced plant species in the diets of migratory Galapagos Tortoises increases with elevation on a human-occupied island. Biotropica 47(2): 246-258. Cayot, L.; Snell, H.L.; Llerena, W.; Snell, H.M. 1994. Conservation biology of Galapagos reptiles: twenty-five years of successful research and management. pp. 297-305. In: Murphy, J.B.; Adler, K. & Collins, J.T. (eds.). Captive management and conservation of amphibians and reptiles. SSAR (Society for the Study of Amphibians and Reptiles). 408 pp. (Contributions to Herpetology no. 11). Chiari, Y.; Hyseni, C.; Fritts, T.H.; Glaberman, S.; Marquez, C.; Gibbs, J.P.; Claude, J.; Caccone, A. 2009. Morphometrics parallel genetics in a newly discovered and endangered taxon of Galapagos tortoise. PLoS ONE 4(7): e6272, 1-11. Ciofi, C.; Wilson, G.A.; Beheregaray, L.B.; Marquez, C.; Gibbs, J.P.; Tapia, W.; Snell, H.L.; Caccone, A.; Powell, J.R. 2006. Phylogeographic history and gene flow among giant Galapagos tortoises on southern Isabela Island. Genetics 172(3): 1727-1744. Denburgh, J. van 1998. The Gigantic Land Tortoises of the Galapagos Archipelago. Society for the Study of Amphibians and Reptiles, Ithaca. 290 pp. Edwards, D.L.; Garrick, R.C.; Tapia, W.; Caccone, A. 2014. Cryptic structure and niche divergence within threatened Galapagos giant tortoises from southern Isabela Island. Conservation Genetics 15(6): 1357-1369. Ellis-Soto, D.; Blake, S.; Soultan, A.; Guezou, A.; Cabrera, F.; Lötters, S. 2017. Plant species dispersed by Galapagos tortoises surf the wave of habitat suitability under anthropogenic climate change. PLoS ONE 12(7): e0181333 Farias, I.P.; Jerozolimski, A.; Melo, A.; Neves Viana, M. das; Martins, M.; Santos Monjelo, L.A. dos 2007. Population genetics of the Amazonian tortoises, Chelonoidis denticulata and C. carbonaria, (Cryptodira: Testudinidae) in an area of sympatry. Amphibia-Reptilia 28(3): 357-365. Garrick, R.C.; Kajdacsi, B.; Russello, M.A.; Benavides, E.; Hyseni, C.; Gibbs, J.P.; Tapia, W.; Caccone, A. 2015. Naturally rare versus newly rare: demographic inferences on two timescales inform conservation of Galapagos giant tortoises. Ecology and Evolution 5(3): 676-694. Gerlach, J. 1998. Famous Tortoises. Nature Protection Trust of the Seychelles, Cambridge. 51 pp. Hunter, E.A.; Gibbs, J.P. 2014. Densities of ecological replacement herbivores required to restore plant communities: a case study of giant tortoises on Pinta Island, Galapagos. Restoration Ecology 22(2): 248-256. Knaack, J. 2006. Beobachtungen zum Vorkommen von Geochelone (Chelonoidis) im südlichen Bolivien. Reptilia (D) 11(59): 77-82. Marquez, C.; Wiedenfeld, D.A.; Landazuri, S.; Chavez, J. 2007. Human-caused and natural mortality of giant tortoises in the Galapagos Islands during 1995-2004. Oryx 41(3): 337-342. McCord, W.P.; Joseph-Ouni, M. 2004. Chelonian illustrations. 14. North and South American tortoises. Reptilia (GB) 34: 47-50. Olson, S.L.; David, N. 2014. The gender of the tortoise genus Chelonoidis Fitzinger, 1835 (Testudines: Testudinidae). Proceedings of the Biological Society of Washington 126(4): 393-394. Papa, L.M. del 2015. Utilización de reptiles durante el periodo Agroalfarero de la región Chaco-Santiagueña, Argentina. Archaeofauna 24: 7-26. Patton, J.L. 1984. Genetical processes in the Galapagos. Biological Journal of the Linnean Society 21(1-2): 97-111. Poulakakis, N.; Glaberman, S.; Russello, M.; Beheregaray, L.B.; Ciofi, C.; Powell, J.R.; Caccone, A. 2008. Historical DNA analysis reveals living descendents of an extinct species of Galapagos tortoise. Proceedings of the National Academy of Sciences of the United States of America 105(40): 15464-15469. Renaud, M.L.; Gitschlag, G.R.; Hale, J.K. 1993. Retention of imitation satellite transmitters fiberglassed to the carapace of Sea Turtles. Herpetological Review 24(3): 94-99. Rothschild, Lord 1925. Giant tortoises of the Galapagos and Mascarene Islands. Proceedings of the Linnean Society of London 137: 40. Sachs, T. 2014. Lonesome George, icon of extinction. The Tortoise 1(3): 174-179. Sims, J. 1991. Captive care. South American tortoises. Rephiberary 168: 4-5. Sims, J. 1991. South American tortoises. Captive conditions part 2. Rephiberary 169: 4-6. Stanford, C.B. 2010. The Last Tortoise: A Tale of Extinction in Our Lifetime. Belknap Press, Cambridge, Massachusetts. 210 pp. Sulloway, F.J. 1984. Darwin and the Galapagos. Biological Journal of the Linnean Society 21(1-2): 29-59. Wüthrich, F. 2006. Die Geschichte der Galápagos-Riesenschildkröte Harriet. Testudo 15(4): 15-27. Chelonoidis abingdonii Anonymous. 2012. Pinta Island, Ecuador. 6. A Galapagos legend dies. Science (Washington D C) 336(6089): 1625. Boulenger, E.G. 1914. Giant saddle-backed tortoise. Proceedings of the Zoological Society of London 1914: 220-222. Cayot, L.J.; Gibbs, J.P.; Tapia, W.; Caccone, A. 2016. Chelonoidis abingdonii. The IUCN Red List of Threatened Species 2016: e.T9017A65487433. http://dx.doi.org/10.2305/IUCN.UK.2016-1.RLTS.T9017A65487433.en. Dowling, H.G. 1961. Vanishing giants and enduring dwarfs - the tortoises. Animal Kingdom 64: 66-75. Edwards, D.L.; Benavides, E.; Garrick, R.C.; Gibbs, J.P.; Russello, M.A.; Dion, K.B.; Hyseni, C.; Flanagan, J.P.; Tapia, W.; Caccone, A. 2013. The genetic legacy of Lonesome George survives: giant tortoises with Pinta Island ancestry identified in Galapagos. Biological Conservation 157: 225-228. Garrick, R.C.; Benavides, E.; Russello, M.A.; Gibbs, J.P.; Poulakakis, N.; Dion, K.B.; Hyseni, C.; Kajdacsi, B.; Márquez, L.D.; Bahan, S.; Ciofi, C.; Tapia Aguilera, W.; Caccone, A. 2012. Genetic rediscovery of an 'extinct' Galápagos giant tortoise species. Current Biology 22(1): R10-11. Jones, J. 2013. Lonesome George: the legacy. Testudo 7(5): 69-78. Nicholls, H. 2006. Lonesome George and the origins and evolution of Galapagos giant tortoises. Testudo 6(3): 64-73. Nicholls, H. 2006. Lonesome George: the life and loves of a conservation icon. Macmillan, London & New York. 231 pp. Philippen, H.D. 2012. [The Pinta Island tortoise Chelonoidis (nigra) abingdoni (Günther, 1877)]. (In German). Marginata 35: 31-34. Pritchard, P.C.H. 1977. Three, two, one tortoise. Natural History (New York) 86(8): 91-100. Pritchard, P.C.H. 2005. The Pinta Tortoise: Globalization and the Extinction of Island Species. Global Connections Foundation, Orlando, Florida. 48 pp. Rowley, A. 2012. Restoration of Pinta Island through the repatriation of giant tortoises. Testudo 7(4): 50-66. Chelonoidis becki Emerson, B.C.; Faria, C.M.A. 2014. Fission and fusion in island taxa - serendipity, or something to be expected. Molecular Ecology 23(21): 5132-5134. Garrick, R.C.; Benavides, E.; Russello, M.A.; Gibbs, J.P.; Poulakakis, N.; Dion, K.B.; Hyseni, C.; Kajdacsi, B.; Márquez, L.D.; Bahan, S.; Ciofi, C.; Tapia Aguilera, W.; Caccone, A. 2012. Genetic rediscovery of an 'extinct' Galápagos giant tortoise species. Current Biology 22(1): R10-11. Garrick, R.C.; Benavides, E.; Russello, M.A.; Hyseni, C.; Edwards, D.L.; Gibbs, J.P.; Tapia, W.H.; Ciofi, C.; Caccone, A. 2014. Lineage fusion in Galapagos Giant Tortoises. Molecular Ecology 23(21): 5276-5290. IUCN Tortoise & Freshwater Turtle Specialist Group. 2016. Chelonoidis becki. The IUCN Red List of Threatened Species 2016: e.T9018A97225095. http://dx.doi.org/10.2305/IUCN.UK.2016-1.RLTS.T9018A97225095.en. Marquez, C. 1986. Herpetology program of the CDRS. Estacion Cientifica Charles Darwin Informe Anual 1983: 28-29. Chelonoidis carbonarius Anonymous. 1994. Hatching red-foots. Tropical Fish Hobbyist 43(4): 128-129. Aponte, C.; Barreto, G.R.; Terborgh, J. 2003. Consequences of habitat fragmentation on age structure and life history in a tortoise population. Biotropica 35(4): 550-555. Auffenberg, W. 1965. Sex and species discrimination in two sympatric South American tortoises. Copeia 1965: 335-342. Bakowsky, R. 2014. [Successful care of Red-footed Tortoises in Venezuela]. (In German). Marginata 42: 36-45. Barros, M.S.; Resende, L.C.; Silva, A.G.; Ferreira-Júnior, P.D. 2012. Morphological variations and sexual dimorphism in Chelonoidis carbonaria (Spix, 1824) and Chelonoidis denticulata (Linnaeus, 1766) (Testudinidae). Brazilian Journal of Biology 72(1): 153-161. Bernardo, P.H.; Guerra-Fuentes, R.A.; Matiazzi, W.; Zaher, H. 2012. Checklist of amphibians and reptiles of Reserva Biológica do Tapirapé, Pará, Brazil. Check List 8(5): 839-846. Bertoni, A. de W. 1926. Nuevos quelonios Paraguayos. Rev. Soc. Cien. Paraguay Asuncion 2(1): 71. Bjorndal, K.A. 1989. Flexibility of digestive responses in two generalist herbivores, the tortoises Geochelone carbonaria and Geochelone denticulata. Oecologia (Berlin) 78(3): 317-321. Borini, J.F.; Petrucci, B.B.; Krohling, W.; Rossi Junior, J.L.; Santos, M.R.D.; Ferreira-Júnior, P.D. 2014. Site fidelity and movement of Chelonoidis carbonaria (Spix, 1824) (Testudinidae) in cocoa plantations in southeastern Brazil. Brazilian Journal of Biology (Suppl. S) 74(3): 135-141. Buchert, P. 2011. Köhlerschildkröten - begehrte, aber anspruchsvolle Pfleglinge. Elaphe 19(3): 44-50. Castano Mora, O.V. 1985. Notas adicionales sobre la reproduccion yel crecimiento de los morrocoyes (Geochelone carbonaria y G. denticulata, Testudines, Testudinidae). Lozania 52: 1-5. Castano Mora, O.V.; Lugo Rugeles, M. 1981. Estudio comparativo del comportamiento de dos especies de morrocoy: Geochelone carbonaria y Geochelone denticulata y aspectos comparables de su morfologia externa. Cespedesia 10(37-38): 55-122. Censky, E.J. 1988. Geochelone carbonaria (Reptilia: Testudines) in the West Indies. Florida Scientist 51(2): 108-114. Colvee, S. 2002. Reproductive aspects of the redfoot tortoise in captivity. Reptilia (GB) 21: 58-66. Conceicao, A.M.; Almeida, B.M.; Brasileiro, M.T.R.; Barros, L.E.D.; Wiltshire, S.A.; Sarmento, C.A.P. 2009. Hermafroditismo em jabuti piranga (Geochelone carbonaria). Arquivo Brasileiro de Medicina Veterinaria e Zootecnia 61(6): 1478-1481. Cunha, O.R. da; Nascimento, F.P. do; Avila Pires, T.C.S. 1985. Os repteis da area de Carajas, Para, Brasil (Testudines e Squamata). 1. Museu Paraense Emilio Goeldi Publicacoes Avulsas 40: 9-92. Czybulinski, R. 2015. Die Köhlerschildkröte Chelonoidis carbonarius – Faszination, Pflege und Zucht. Kleintierverlag Thorsten Geier, Biebertal. 161 pp. Dathe, F. 2011. Haltung und Vermehrung von Köhlerschildkröten, Chelonoidis carbonaria (Spix, 1824), im Tierpark Berlin. Milu 13(3): 413-425. Davis, S. 1979. Husbandry and breeding of the red-footed tortoise Geochelone carbonaria at the National Zoological Park, Washington. International Zoo Yearbook 19: 50-53. Dowling, H.G. 1961. Vanishing giants and enduring dwarfs - the tortoises. Animal Kingdom 64: 66-75. Dubois, B.; Vuylsteker, J. 2007. Elevage et reproduction en captivite de la Tortue charbonniere, Chelonoidis carbonaria (Spix, 1824). Cheloniens 5: 38-44. Ernst, C.H.; Leuteritz, T.E.J. 1999. Geochelone carbonaria. Catalogue of American Amphibians and Reptiles 690: 1-7. Fehringer, P. 2010. Köhlerschildkröten (Chelonoidis carbonaria ssp.) auf den Kleinen Antillen in der Natur beobachtet. Marginata 27: 16-19. Fehringer, P. 2010. Köhlerschildkröten auf der Insel Barbuda - eine ungewöhnliche Erfahrung bei einem Kurzaufenthalt. Marginata 27: 28-29. Flanigan, W.F. 1974. Sleep and wakefulness in chelonian reptiles. 2. The red-footed tortoise, Geochelone carbonaria. Archives ital. Biol. 112(3): 253-277. Fritz, C. 2001. Erfolgreiche Haltung und Vermehrung der Köhlerschildkröte (Geochelone carbonaria) aus dem nördlichen Südamerika. Draco 2(4) (8): 72-77. Gorzula, S. 1989. Recent population changes: Chelonoidis carbonaria (red footed tortoise). Herpetological Review 20(2): 56. Guix, J.C.; Fedullo, D.L.; Molina, F.B. 2001. Masculinization of captive females of Chelonoidis carbonaria (Testudinidae). Revista Española de Herpetología 15: 67-75. Guix, J.C.C.; Salvatti, M.; Peroni, M.A.; Lima-Verde, J.S. 1989. Observations additionnelles sur le comportement et la reproduction de Chelonoidis carbonaria (Spix, 1824) en captivite (Testudines-Testudinidae). Grupo de Estudos Ecologicos Serie Documentos 1: 20-31. Gutsche, A. 2012. Klassiker: Die Köhlerschildkröte. Terraria-Elaphe 2012(38): 68-69. Hedman, H.D.; Muniz, D.N.; Powell, R. 2010. Natural history notes: Chelonoidis carbonaria (Red-footed Tortoise). Size and thermal biology. Herpetological Review 41(4): 484-485. Hernandez, O.; Boede, E.O. 2002. Efectos de la densidad y la proporcion de sexos en la reproduccion en cautiverio del Morrocoy Geochelone (Chelonoidis) carbonaria (Spix 1824). Acta Biologica Venezuelica 21(2): 29-37. Hernández-Montoya, V.; Páez, V.P.; Ceballos, C.P. 2017. Effects of temperature on sex determination and embryonic development in the Red-footed Tortoise, Chelonoidis carbonarius. Chelonian Conservation and Biology 16(2): 164-171. Hernandez-P., O.E. 1997. Reproduccion y crecimiento del morrocoy, Geochelone (Chelonoidis) carbonaria (Spix, 1824) (Reptilia, Testudinidae). BioLlania 13: 165-183. Hernandez-P., O.E.; Boede, E.O. 2000. Efecto de la alimentacion sobre el crecimiento y produccion de huevos de Geochelone (Chelonoidis) carbonaria (Spix, 1824) bajo condiciones de cautiverio. Acta Biologica Venezuelica 20(2): 37-43. Herz, M. 2015. [Encounters with Testudines in the Venezuelan state of Amazonas. Part 2: Tortoises]. (In German). Reptilia (D) 20(111): 64-68. Hoogmoed, M.S.; Gruber, U. 1983. Spix and Wagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (the Netherlands). Spixiana Supplement 9: 319-415. Houbrix, P. 2007. Elevage et reproduction en captivite de la tortue charbonniere, Chelonoidis carbonaria (Spix, 1824). Manouria 10(36): 12-18. Houtman, H.; Fosse, A. de la 1988. Care and breeding of Geochelone carbonaria. Lacerta 47(2) [1988-1989]: 40-46. Knaack, J. 2004. Köhler- und Waldschildkröte. Beobachtungen zum Vorkommen von Geochelone (Chelonoides) carbonaria und G. (C.) denticulata in Bolivien. Aquarium (Bornheim) 416: 70-74. Kradolfer, K.D. 2004. Beobachtungen an Köhlerschildkröten während des Freilaufs im Garten. Testudo 13(1): 37-40. Kundert, S. 2004. Bemerkungen zur Bestimmung der Herkunft von Köhlerschildkröten aufgrund der Plastrongefärbung. Testudo 13(1): 41-42. Kundert, S. 2004. Haltung un Vermehrung der Köhlerschildkröte Geochelone carbonaria (Spix 1824). Testudo 13(1): 21-36. Legler, J.M. 1963. Tortoises (Geochelone carbonaria) in Panama: distribution and variation. American Midland Naturalist 70: 490-503. Lowe, P. 1997. Redfoot tortoises: South American treasure. Reptile & Amphibian Magazine 46: 36-43. Marcellini, D.L.; Davis, S.W. 1982. Effects of handling on reptile egg hatching. Herpetological Review 13(2): 43-44. May, C. 2004. Pflege und Vermehrung der rotköpfigen Köhlerschildkröte "Cherryhead", Geochelone carbonaria. 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