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Wüster, W.; Crookes, S.; Ineich, I.; Mané, Y.; Pook, C.E.; Trape, J.F.; Broadley, D.G. 2007 The phylogeny of cobras inferred from mitochondrial DNA sequences: Evolution of venom spitting and the phylogeography of the African spitting cobras (Serpentes: Elapidae: Naja nigricollis complex). Molecular Phylogenetics & Evolution 45(2): 437-453.