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Jaramillo, C.; Wilson, L.D.; Ibañez-D., R.; Jaramillo, F.
2010
The herpetofauna of Panama: distribution and conservation status. pp. 604-671.
In: Wilson, L.D.; Townsend, J.H.; Johnson, J.D. (eds.). Conservation of Mesoamerican amphibians and reptiles. Eagle Mountain Publishing, LC, Eagle Mountain, Utah. 812 pp.