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Kindler, C.; Branch, W.R.; Hofmeyr, M.D.; Maran, J.; Siroky, P.; Vences, M.; Harvey, J.; Hauswaldt, S.; Schleicher, A.; Stuckas, H.; Fritz, U. 2012 Molecular phylogeny of African Hinge-back Tortoises (Kinixys): implications for phylogeography and taxonomy (Testudines: Testudinidae). Journal of Zoological Systematics and Evolutionary Research 50(3): 192-201.