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Welton, L.J.; Siler, C.D.; Grismer, L.L.; Diesmos, A.C.; Sites, J.W.; Brown, R.M. 2017 Archipelago-wide survey of Philippine forest dragons (Agamidae: Gonocephalus): multilocus phylogeny uncovers unprecedented levels of genetic diversity in a biodiversity hotspot. Biological Journal of the Linnean Society 120: 410-426.