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Solovyeva, E.N.; Dunayev, E.N.; Nazarov, R.A.; Radjabizadeh, M.; Poyarkov, N.A. 2018 Molecular and morphological differentiation of Secret Toad-headed agama, Phrynocephalus mystaceus, with the description of a new subspecies from Iran (Reptilia, Agamidae). ZooKeys 748: 97-129.