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Taxonomy of the genus Amblyrhynchus Biodiversity of the family Iguanidae

Bibliography of the genus
Amblyrhynchus (Marine Iguana)

(Reptilia: Sauria: Iguanidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Amblyrhynchus cristatus

Anonymous. 1981. Marine iguanas and feral dogs. Oryx 16(2): 156.

Anonymous. 1986. Recovery of the marine iguana population after the El Nino catastrophe. Noticias de Galapagos 44: 4.

Amadon, D. 1967. Galapagos finches grooming marine iguanas. Condor 69: 311.

Ananjeva, N.B. 1978. [The Galapagos iguana - a unique marine reptile]. (In Russian). Priroda (Moscow) 1978(11): 139.

Baird, T.M. 1981. Dragons of the shore. Marine iguanas. Sea Frontiers 27(5): 306-311.

Bartholomew, G.A. 1966. A field study of temperature relations in the Galápagos marine iguana. Copeia 1966: 241-250.

Bartholomew, G.A.; Bennett, A.F.; Dawson, W.R. 1976. Swimming, diving and lactate production of the marine iguana, Amblyrhynchus cristatus. Copeia 1976(4): 709-720.

Bartholomew, G.A.; Lasiewski, R.C. 1965. Heating and cooling rates, heart rate and simulated diving in the Galapagos marine iguana. Comparative Biochemistry and Physiology 16: 573-582.

Bartholomew, G.A.; Vleck, D. 1979. The relation of oxygen consumption to body size and to heating and cooling in the Galapagos marine iguana, Amblyrhynchus cristatus. Journal of Comparative Physiology 132(4): 285-288.

Bennett, A.F.; Dawson, W.R.; Bartholomew, G.A. 1975. Effects of activity and temperature on aerobic and anaerobic metabolism in the Galapagos marine iguana. Journal of Comparative Physiology 100(4): 317-329.

Berger, S.; Wikelski, M.; Romero, L.M.; Kalko, E.K.V.; Roedl, T. 2007. Behavioral and physiological adjustments to new predators in an endemic island species, the Galapagos marine iguana. Hormones and Behavior 52(5): 653-663.

Boersma, P.D. 1982. The benefits of sleeping aggregations in marine iguanas, Amblyrhynchus cristatus. pp. 292-302. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Boersma, P.D. 1983. An ecological study of the Galapagos marine iguana. pp. 157-176. In: Bowman, R.I.; Berson, M. & Leviton, A.E. (eds.). Patterns of evolution in Galapagos organisms. Pacific Division AAAS, San Francisco. 568 pp.

Butler, P.J.; Frappell, P.B.; Wang, T.; Wikelski, M. 2002. The relationship between heart rate and rate of oxygen consumption in Galapagos marine iguanas (Amblyrhynchus cristatus) at two different temperatures. Journal of Experimental Biology 205(13): 1917-1924.

Buttemer, W.A.; Dawson, W.R. 1993. Temporal pattern of foraging and microhabitat use by Galapagos marine iguanas, Amblyrhynchus cristatus. Oecologia (Heidelberg) 96(1): 56-64.

Carpenter, C.C. 1966. The marine iguana of the Galapagos Islands, its behavior and ecology. Proceedings of the California Academy of Sciences (Ser. 4) 34: 329-376.

Castillo-Briceno, P.; Marquez, C.; Wiedenfeld, D.; Snell, H.; Jaramillo, A. 2004. El Nino oscilacion del Sur" y Estado de Salud de Las iguanas marinas (Amblyrhynchus cristatus) en Galapagos. Ecologia Aplicada 3(1-2): 92-97.

Cayot, L.; Rassmann, K.; Trillmich, F. 1994. Are marine iguanas endangered on islands with introduced predators? Noticias de Galapagos 53: 13-15.

Cayot, L.; Rossmann, K.; Trillmich, F. 1993. Status of the marine iguana: are they threatened by introduced predators? Galapagos Bulletin 1993(Fall): 2-3.

Chiari, Y.; Glaberman, S.; Tarroso, P.; Caccone, A.; Julien, C. 2016. Ecological and evolutionary influences on body size and shape in the Galapagos Marine Iguana (Amblyrhynchus cristatus). Oecologia (Berlin) 181(3): 885-894.

Cooper, J.E.; Laurie, W.A. 1987. Investigation of deaths in marine iguanas (Amblyrhynchus cristatus) on Galapagos. Journal of Comparative Pathology 97(2): 129-136.

Dawson, E.Y. 1962. The giants of Galápagos. Natural History (New York) 71(9): 52-57.

Dawson, W.R.; Bartholomew, G.A.; Bennett, A.F. 1978. A reappraisal of the aquatic specializations of the Galapagos marine iguana (Amblyrhynchus cristatus). Evolution 31(4): 891-897.

Dellinger, T. 1996. Population dynamics and energetics of reproduction of marine iguanas (Amblyrhynchus cristatus) in Galapagos. Charles Darwin Research Station Annual Report 1988-1989: 83-85.

Dellinger, T.; Hegel, G. von 1990. Sex identification through cloacal probing in juvenile marine iguanas (Amblyrhynchus cristatus). Journal of Herpetology 24(4): 424-426.

Denburgh, J. van; Slevin, J.R. 1913. The Galapagoan lizards of the genus Tropidurus; with notes on the iguanas of the genera Conolophus and Amblyrhynchus. (Expedition of the California academy of Sciences to the Galapagos islands, 1905-1906, 9.). Proceedings of the California Academy of Sciences 2: 132-202.

Dowling, H.G. 1962. Sea dragons of the Galapagos: the marine iguanas. Animal Kingdom 65: 169-174.

Drent, J.; Marken Lichtenbelt, W.D. van; Wikelski, M. 1999. Effects of foraging mode and season on the energetics of the marine iguana, Amblyrhynchus cristatus. Functional Ecology 13(4): 493-499.

Dunson, W.A. 1969. Electrolyte excretion by the salt gland of the Galapagos marine iguana. American Journal of Physiology 216: 995-1002.

Eibl-Eibesfeldt, I. 1955. Der Kommentkampf der Meerechse (Amblyrhynchus cristatus Bell), nebst einigen Notizen zur Biologie dieser Art. Zeitschrift Tierpsychol. 12: 49-62.

Eibl-Eibesfeldt, I. 1956. Eine neue Rasse der Meerechse, Amblyrhynchus cristatus venustissimus, nebst einigen Bemerkungen über Amblyrhynchus cristatus cristatus. Senckenbergiana Biologica 37: 87-100.

Eibl-Eibesfeldt, I. 1962. Neue Unterarten der Meerechse, Amblyrhynchus cristatus, nebst weiteren Angaben zur Biologie der Art. Senckenbergiana Biologica 43: 177-199.

Eibl-Eibesfeldt, I. 1963. Meerechsen in Gefangenschaft. Natur und Museum (Frankfurt Am Main) 93: 410-414.

Eibl-Eibesfeldt, I. 1964. The art of keeping marine iguanas. Animal Kingdom 67: 75-77.

Eibl-Eibesfeldt, I. 1966. Das Verteidigen der Eiablageplätze bei der Hood-Meerechse (Amblyrhynchus cristatus venustissimus). Zeitschrift für Tierpsychologie 23: 627-631.

Eibl-Eibesfeldt, I. 1984. The large iguanas of the Galapagos Islands. pp. 157-173. In: Perry, R. (ed.). Key environments. Galapagos. Pergamon Press, Oxford, New York etc. 321 pp.

Ellis, H.I.; Ross, J.P. 1978. Field observations of cooling rates of Galapagos land iguanas (Conolophus subcristatus). Comparative Biochemistry and Physiology A Comparative Physiology 59(2): 205-209.

Estes, G.; Jackson, M.; Meatyard, B. 1983. Feeding ecology of marine iguanas (Amblyrhynchus cristatus) at Pta. Nunez, Sta. Cruz. Estacion Cientifica Charles Darwin Informe Anual 1982: 318-320.

Etheridge, R.E. 1982. Checklist of the iguanine and Malagasy iguanid lizards. pp. 7-37. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

French, S.S.; DeNardo, D.F.; Greives, T.J.; Strand, C.R.; Demas, G.E. 2010. Human disturbance alters endocrine and immune responses in the Galapagos Marine Iguana (Amblyrhynchus cristatus). Hormones and Behavior 58(5): 792-799.

Fritts, T.H.; Snell, H.L.; Cayot, L.; MacFarland, C.; Earsom, S.; Marquez, C.; Llerena, W.; Llerena, F. 2000. Progress and priorities in research for the conservation of reptiles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie 70(Supplement): 39-45.

Frost, I.; Beck Frost, M. 1983. Limitations to algal growth in the Galapagos Islands: its consequences on the breeding strategy of the marine iguana Amblyrhynchus cristatus. Noticias de Galapagos 37: 23-24.

Gleeson, T.T. 1979. Foraging and transport costs in the Galapagos marine iguana, Amblyrhynchus cristatus. Physiological Zoology 52(4): 549-557.

Hayes, W.K.; Carter, R.L.; Wikelski, M.; Sonnentag, J.A. 2004. Determinants of lek mating success in male Galapagos marine iguanas: behavior, body size, condition, ornamentation, ectoparasite load and female choice. pp. 127-147. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Higgins, P.J. 1978. The Galapagos iguanas: models of reptilian differentiation. Bioscience 28(8): 512-515.

Higgins, P.J.; Rand, C.S.; Haynes, J.L. 1974. Galapagos iguanas: Amblyrhynchus and Conolophus serum protein relationships. Journal of Experimental Zoology 189(2): 255-260.

Hobson, E.S. 1965. Observations on diving in the Galapagos marine iguana, Amblyrhynchus cristatus (Bell). Copeia 1965: 249-250.

Hobson, E.S. 1969. Remarks on aquatic habits of the Galapagos marine Iguana, including submergence times, cleaning symbiosis, and the shark threat. Copeia 1969: 401-402.

Hobson, E.S. 1974. The marine iguana. Oceans 7(2): 48-49.

Holmes, M. 1991. Sea trek. BBC Books, London. 240 pp.

Hugi, J.; Sánchez-Villagra, M.R. 2012. Life history. and skeletal adaptations in the Galapagos Marine Iguana (Amblyrhynchus cristatus) as reconstructed with bone histological data - a comparative study of Iguanines. Journal of Herpetology 46(3): 312-324.

Ineich, I.; Miralles, A. 2014. Natural history notes: Amblyrhynchus cristatus (Marine Iguana). Polydactyly. Herpetological Review 45(2): 322-323.

Knudtson, P.M. 1980. Imps of darkness. Oceans 13(5): 46-50.

Krefft, G. 1953. Herpetologische Eindrücke einer Walfangreise. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 6: 100-103.

Kruuk, H.; Snell, H. 1981. Prey selection by feral dogs from a population of marine iguanas (Amblyrhynchus cristatus). Journal of Applied Ecology 18(1): 197-204.

Lanterbecq, D.; Glaberman, S.; Vitousek, M.N.; Steinfartz, S.; Benavides, E.; Wikelski, M.; Caccone, A. 2010. Genetic differentiation between Marine Iguanas from different breeding sites on the Island of Santa Fe (Galapagos Archipelago). Journal of Heredity 101(6): 663-675.

Laurie, A. 1982. Marine iguanas - where have all their babies gone? Noticias de Galapagos 35: 17-19.

Laurie, A. 1982. Population dynamics and social organization of marine iguanas on Galapagos. Estacion Cientifica Charles Darwin Informe Anual 1981: 111-134.

Laurie, A. 1982. Santa Fe newsletter. Noticias de Galapagos 36: 19-20.

Laurie, A. 1983. An ill wind for iguanas. New Scientist 100(1379): 108.

Laurie, A. 1983. Marine iguanas in Galapagos. Oryx 17(1): 18-25.

Laurie, A. 1983. Marine iguanas suffer as El Nino breaks all records. Noticias de Galapagos 38: 11.

Laurie, A. 1983. Population dynamics and social organization of marine iguanas on Galapagos. (Progress report No. 2: August 1982). Estacion Cientifica Charles Darwin Informe Anual 1982: 321-335.

Laurie, A. 1984. Interim report on the marine iguana situation in the aftermath of the 1982-3 El Nino. Noticias de Galapagos 40: 9-11.

Laurie, A. 1985. Santa Fe newsletter. Noticias de Galapagos 41: 20-21.

Laurie, A. 1985. The effects of the 1982-83 El Nino on marine iguanas. pp. 199-209. In: Robinson, G. & del Pino, E.M. (eds.). El Nino en las Islas Galapagos: el evento de 1982-1983. Charles Darwin Foundation for the Galapagos Islands, Quito. 534 pp.

Laurie, A. 1986. Population dynamics and social organisation of marine iguanas on Galapagos. Progress report No. 3 September 1983. Estacion Cientifica Charles Darwin Informe Anual 1983: 56-58.

Laurie, A. 1987. Marine iguana project to continue. Noticias de Galapagos 45: 19-22.

Laurie, A. 1987. Marine iguanas: living on the ocean margin. Oceanus 30(2): 54-60.

Laurie, W.A. 1990. Effects of the 1982-83 El Nino-Southern Oscillation event on marine iguana (Amblyrhynchus cristatus Bell, 1925) populations on Galapagos. Elsevier Oceanography Series 52: 361-380.

Laurie, W.A. 1990. Population biology of marine iguanas (Amblyrhynchus cristatus). 1. Changes in fecundity related to a population crash. Journal of Animal Ecology 59(2): 515-528.

Laurie, W.A.; Brown, D. 1990. Population biology of marine iguanas (Amblyrhynchus cristatus). 2. Changes in annual survival rates and the effects of size, sex, age and fecundity in a population crash. Journal of Animal Ecology 59(2): 529-544.

Laurie, W.A.; Brown, D. 1990. Population biology of marine iguanas (Amblyrhynchus cristatus). 3. Factors affecting survival. Journal of Animal Ecology 59(2): 545-568.

Lewbart, G.A.; Hirschfeld, M.; Ricaurte, O.; Lohmann, K.J. 2017. Natural history notes: Amblyrhynchus cristatus (Marine Iguana). Diet. Herpetological Review 48(2): 432.

Llerena, A.; Marquez, C.; Snell, H.L.; Jaramillo, A. 2004. Abundancia de Amblyrhynchus cristatus en El Nino (97-98) y La Nina (01-02) en Santa Cruz, Galapagos. Ecologia Aplicada 3(1-2): 87-91.

Lucker, H. 1994. Die Meeresechsen (Amblyrhynchus cristatus) von Galapagos. Sauria (Berlin) 16(2): 3-8.

Luecker, H.; Feiler, A. 2002. Beobachtung einer Paarung zwischen Meerechsen-Männchen (Amblyrhynchus cristatus) und Landleguan-Weibchen (Conolophus subcristatus) auf Plaza-Sur/Galapagos-Inseln sowie Beobachtungen an einem adulten Gattungshybriden (Amblyrhynchus X Conolophus). Elaphe 10(4): 49-54.

Mackie, R.I.; Rycyk, M.; Ruemmler, R.L.; Aminov, R.I.; Wikelski, M. 2004. Biochemical and microbiological evidence for fermentative digestion in free-living land iguanas (Conolophus pallidus) and marine iguanas (Amblyrhynchus cristatus) on the Galapagos Archipelago. Physiological and Biochemical Zoology 77(1): 127-138.

MacLeod, A.; Rodriguez, A.; Vences, M.; Orozco-Terwengel, P.; García, C.; Trillmich, F.; Gentile, G.; Caccone, A.; Quezada, G.; Steinfartz, S. 2015. Hybridization masks speciation in the evolutionary history of the Galapagos Marine Iguana. Proceedings of the Royal Society Biological Sciences Series B 282(1809): 20150425.

MacLeod, A.; Steinfartz, S. 2016. The conservation status of the Galápagos Marine Iguanas, Amblyrhynchus cristatus: a molecular perspective. Amphibia-Reptilia 37(1): 91-109.

MacLeod, A.; Unsworth, L.; Trillmich, F.; Steinfartz, S. 2016. Mark-resight estimates confirm a critically small population size in threatened marine iguanas (Amblyrhynchus cristatus) on San Cristobal Island, Galapagos. Salamandra 52(1): 58-62.

Meermann, H. 1978. Das Energie-Kalkül der Meerechsen. Kosmos (Stuttgart) 74(1): 12-14.

Merlen, G. 1985. The 1982-83 El Nino: some of its consequences for Galapagos wildlife. Noticias de Galapagos 41: 8-15.

Merlen, G. 1995. Fundamentals. Noticias de Galapagos 55: 4-6.

Mertens, R. 1927. Meeresechsen. Bericht der Senckenbergischen Naturforschenden Gesellschaft (Frankfurt) 57(2): 78-91.

Miles, D.B.; Fitzgerald, L.A.; Snell, H.L. 1995. Morphological correlates of locomotor performance in hatchling Amblyrhynchus cristatus. Oecologia (Heidelberg) 103(2): 261-264.

Milton, Y.; Cayot, L.J.; Jaramillo, A. 2012. Dispersal of neonatal Galapagos Marine Iguanas Amblyrhynchus cristatus from their nesting zone: natural history and conservation implications. Herpetological Conservation and Biology 7(3): 470-480.

Miralles, A.; Macleod, A.; Rodríguez, A.; Ibáńez, A.; Jiménez-Uzcategui, G.; Quezada, G.; Vences, M.; Steinfartz, S. 2017. Shedding light on the imps of darkness: an integrative taxonomic revision of the Galápagos Marine Iguanas (genus Amblyrhynchus). Zoological Journal of the Linnean Society 181(3): 678-710.

Nagy, K.A.; Shoemaker, V.H. 1984. Field energetics and food consumption of the Galapagos marine iguana, Amblyrhynchus cristatus. Physiological Zoology 57(3): 281-290.

Nelson, K.; Snell, H.; Wikelski, M. 2004. Amblyrhynchus cristatus. The IUCN Red List of Threatened Species 2004: e.T1086A3222951. http://dx.doi.org/10.2305/IUCN.UK.2004.RLTS.T1086A3222951.en.

O'Connor, M.P. 1999. Physiological and ecological implications of a simple model of heating and cooling in reptiles. Journal of Thermal Biology 24(2): 113-136.

Ogamba, E.N.; Abowei, J.F.N. 2012. Some aquatic reptiles in culture fisheries management. International Journal of Fisheries and Aquatic Sciences 1(1): 5-15.

Oliver, J.A. 1956. Lizards of the sea. Animal Kingdom 59: 148-151.

Olson, S.L. 2014. The early scientific history of Galapagos iguanas. Archives of Natural History 41(1): 141-153.

Pardi, L. 1982. Some observations on orientation in Talorchestia fritzi Stebbing, Grapsus grapsus L. and Amblyrhynchus cristatus Bell from the Galapagos Islands. pp. 129-147. Museo Zoologico dell'Universita di Firenze, Galapagos. Studi e ricerche. Spedizione 'L. Mares - G.R.S.T.S.' Gruppo Ricerche Scientifische e Techniche Subacquee. 426 pp.

Partecke, J.; Haeseler, A. von; Wikelski, M. 2002. Territory establishment in lekking marine iguanas, Amblyrhynchus cristatus: Support for the hotshot mechanism. Behavioral Ecology and Sociobiology 51(6): 579-587.

Pawley, R. 1965. Housekeeping for marine iguanas. Animal Kingdom 68: 146-150.

Pawley, R. 1966. Observations on the care and nutrition of a captive group of marine iguanas Amblyrhynchus cristatus. International Zoo Yearbook 6: 107-115.

Pawley, R. 1969. Further notes on a captive colony of marine iguanas Amblyrhynchus cristatus at Brookfield Zoo, Chicago. International Zoo Yearbook 9: 41-44.

Rasmussen, A.R.; Murphy, J.C.; Ompi, M.; Whitfield-Gibbons, J.; Uetz, P. 2011. Marine reptiles. PLoS ONE 6(11): e27373, 1-12.

Rassmann, K. 1996. Molecular genetics and conservation in the Galapagos. Noticias de Galapagos 56: 22-26.

Rassmann, K. 1997. Evolutionary age of the Galapagos iguanas predates the age of the present Galapagos islands. Molecular Phylogenetics and Evolution 7(2): 158-172.

Rassmann, K.; Markmann, M.; Trillmich, F.; Tautz, D. 2004. Tracing the evolution of the Galapagos iguanas: a molecular approach. pp. 71-83. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Rassmann, K.; Tautz, D.; Trillmich, F.; Gliddon, C. 1997. The microevolution of the Galapagos marine iguana Amblyrhynchus cristatus assessed by mitochondrial genetic analysis. Molecular Ecology 6(5): 437-452.

Rassmann, K.; Trillmich, F.; Tautz, D. 1997. Hybridization between the Galapagos land and marine iguana (Conolophus subcristatus and Amblyrhynchus cristatus) on Plaza Sur. Journal of Zoology (London) 242(4): 729-739.

Rauch, N. 1982. Reproductive behaviour of male and female marine iguanas. Estacion Cientifica Charles Darwin Informe Anual 1981: 135-138.

Rauch, N. 1983. Reproductive strategies of male and female marine iguana (Amblyrhynchus cristatus). Estacion Cientifica Charles Darwin Informe Anual 1982: 336-346.

Rauch, N. 1985. Female habitat choice as a determinant of the reproductive success of the territorial male marine iguana (Amblyrhynchus cristatus). Behavioral Ecology and Sociobiology 16(2): 125-134.

Rauch, N. 1988. Competition of marine iguana females (Amblyrhynchus cristatus) for egg-laying sites. Behaviour 107(1-2): 91-106.

Robinson, G.R. 1987. Negative effects of the 1982-83 El Nino on Galapagos marine life. Oceanus 30(2): 42-48.

Roedl, T.; Berger, S.; Romero, L.M.; Wikelski, M. 2007. Tameness and stress physiology in a predator-naive island species confronted with novel predation threat. Proceedings of the Royal Society Biological Sciences Series B 274(1609): 577-582.

Rohrbach, C. 1981. Ecological factors in the development of marine iguanas, Amblyrynchus cristatus. Charles Darwin Research Station Annual Report 1980: 164-165.

Romero, L.M.; Wikelski, M. 2001. Corticosterone levels predict survival probabilities of Galapagos marine iguanas during El Nino events. Proceedings of the National Academy of Sciences of the United States of America 98(13): 7366-7370.

Romero, L.M.; Wikelski, M. 2002. Exposure to tourism reduces stress-induced corticosterone levels in Galapagos marine iguanas. Biological Conservation 108(3): 371-374.

Romero, L.M.; Wikelski, M. 2010. Stress physiology as a predictor of survival in Galapagos Marine Iguanas. Proceedings of the Royal Society Biological Sciences Series B 277(1697): 3157-3162.

Romero-L., M. 2012. Using the reactive scope model to understand why stress physiology predicts survival during starvation in Galapagos Marine Iguanas. General and Comparative Endocrinology 176(3): 296-299.

Rosenfeld, A.E. 1978. The Galapagos marine iguana, a review. Bulletin of the Philadelphia Herpetological Society 26: 13-27.

Roy Moore, T. de 1980. Dragon of the sea. International Wildlife 10(2): 48-56.

Rubenstein, D.R.; Wikelski, M. 2003. Seasonal changes in food quality: a proximate cue for reproductive timing in marine iguanas. Ecology (Washington, D.C.) 84(11): 3013-3023.

Rubenstein, D.R.; Wikelski, M. 2005. Steroid hormones and aggression in female Galapagos marine iguanas. Hormones and Behavior 48(3): 329-341.

Sayre, R. 1984. Creatures. Audubon 86(1): 98-99.

Schellenger, K. 1974. The diving lizards of the Galapagos. Pacific Discovery 27(6): 24-28.

Shaw, C.E. 1967. The eggs and young of the marine iguana (Amblyrhynchus cristatus). Copeia 1966: 585.

Shepherd, S.A.; Hawkes, M.W. 2005. Algal food preferences and seasonal foraging strategy of the marine iguana, Amblyrhynchus cristatus, on Santa Cruz, Galapagos. Bulletin of Marine Science 77(1): 51-72.

Shoemaker, V.H.; Nagy, K.A. 1984. Osmoregulation in the Galapagos marine iguana, Amblyrhynchus cristatus. Physiological Zoology 57(3): 291-300.

Smith, M. 1956. Marine iguana. Zoo Life 11: 51-53.

Steinfartz, S.; Glaberman, S.; Lanterbecq, D.; Marquez, C.; Rassmann, K.; Caccone, A. 2007. Genetic impact of a severe El Nino event on Galapagos marine iguanas (Amblyrhynchus cristatus). PLoS ONE 2(12): e1285, 1-9.

Steinfartz, S.; Glaberman, S.; Lanterbecq, D.; Russello, M.A.; Rosa, S.; Hanley, T.C.; Marquez, C.; Snell, H.L.; Snell, H.M.; Gentile, G.; Dell'Omo, G.; Powell, A.M.; Caccone, A. 2009. Progressive colonization and restricted gene flow shape island-dependent population structure in Galapagos marine iguanas (Amblyrhynchus cristatus). BMC Evolutionary Biology 9(297): 1-18.

Steinfartz, S.; Glaberman, S.; Lanterbecq, D.; Russello, M.A.; Rosa, S.; Hanley, T.C.; Márquez, C.; Snell, H.L.; Snell, H.M.; Gentile, G.; Dell'Omo, G.; Powell, A.M.; Caccone, A. 2010. Fortschreitende Besiedlung und eingeschänkter Genfluß formen inselabhängige Populationsstrukturen der Galapagos-Meerechsen (Amblyrhynchus cristatus). Braunschweiger Naturkundliche Schriften 9(2): 157-183.

Swainston, J. (ed.) 1988. Report of the March meeting. Marine iguanas, by Dr. Andrew Laurie. Rephiberary 125: 1-4.

Switak, K.H. 1999. Die Meerechsen der Galapagos-Inseln. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 52(7): 50-54.

Tilby, J.G.W. 1935. Local variation of habits of the lizard, Amblyrhynchus cristatus. Nature (London) 135(3404): 151.

Tracy, C.R.; Christian, K.A. 1985. Are marine iguana tails flattened? British Journal of Herpetology 6(12): 434-435.

Trillmich, F. 1979. Feeding behaviour and social behaviour of the marine iguana. Noticias de Galapagos 29: 17-20.

Trillmich, F.; Trillmich, K.G.K. 1984. The mating systems of pinnipeds and marine iguanas: convergent evolution of polygyny. Biological Journal of the Linnean Society 21(1-2): 209-216.

Trillmich, K.G.K. 1983. The mating system of the marine iguana (Amblyrhynchus cristatus). Zeitschrift für Tierpsychologie 63(3-4): 141-172.

Trillmich, K.G.K.; Trillmich, F. 1986. Foraging strategies of the marine iguana, Amblyrhynchus cristatus. Behavioral Ecology and Sociobiology 18(4): 259-266.

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