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Taxonomy of the family Agamidae
Bibliography of the genus Ctenophorus
Biodiversity of the family Agamidae








Genus
Ctenophorus

Comb Dragons, Comb-bearing Dragons

Soldatenagamen

Australske Kamagamer

1843 Ctenophorus Fitzinger (type species: Grammatophora decresii Dumeril & Bibron 1837)
1843 Homalonotus Fitzinger [not Homalonotus Koenig 1825 (Trilobita); not Homalonotus Schoenherr 1836 (Insecta: Coleoptera); not Homalonotus Agassiz 1846 (Arachnida)] (type species: Grammatophora gaimardii Dumeril & Bibron 1837; syn. Cogger, Cameron & Cogger 1983)
1884 Macrops De Vis (type species: Macrops nuchalis De Vis 1884; syn. Cogger 1996)
1984 Licentia Wells & Wellington (type species: Grammatophora cristata Gray 1841; syn. Cogger 1996)
1984 Phthanodon Wells & Wellington (type species: Uromastyx maculatus Gray 1831; syn. Cogger 1996)
1985 Tachyon Wells & Wellington (type species: Grammatophora caudicincta Günther 1875; syn. Cogger 1996)
Contents: 33 species, all of which are endemic.
Remarks: Previously regarded as a synonym of Amphibolurus (e.g., Wermuth 1967). Revalidated by Storr, Smith & Johnstone (1983). Some species were previously placed in Rankinia or in Tympanocryptis (adelaidensis, butlerorum, chapmani, parviceps).
Distribution: Australia (New South Wales, Northern Territory, Queensland, South Australia [incl. Kangaroo Island], Victoria, Western Australia).

Ctenophorus adelaidensis

Western Heath Dragon, Prickly Dragon, Queen Adelaide Dragon

Westliche Heideagame

Vestlig Hedeagam

1841 Grammatophora muricata var. adelaidensis Gray
Grammatophora adelaidensis Günther 1875
Amphibolurus adelaidensis Boulenger 1885
Tympanocryptis adelaidensis Storr, Smith & Johnstone 1983
Rankinia adelaidensis Wells & Wellington 1984
Ctenophorus adelaidensis Melville, Schulte & Larson 2001
1845 Grammatophora angulifera Gray [part] (syn.Cogger, Cameron & Cogger 1983)
1885 Amphibolurus adelaidensis tasmaniensis Boulenger (syn.Zietz 1921; Cogger, Cameron & Cogger 1983)
1885 Amphibolurus pallidus Boulenger (syn.Loveridge 1934; Cogger, Cameron & Cogger 1983)
1885 Amphibolurus pulcherrimus Boulenger (syn.Loveridge 1934; Cogger, Cameron & Cogger 1983)
Amphibolurus adelaidensis pulcherrimus Glauert 1961

Remarks: Previously included butlerorum and chapmani. Records from South Australia refer to chapmani (Hutchinson 2010).
Distribution: Australia (Western Australia).


Ctenophorus butlerorum

Shark Bay Heath Dragon, (Butler's Dragon)

Shark-Bay-Heideagame

Shark Bay-hedeagam

1977 Tympanocryptis parviceps butleri Storr
Tympanocryptis butleri Storr, Smith & Johnstone 1983
Amphibolurus parviceps butleri Cogger 1983
Rankinia butleri Melville, Shoo & Doughty 2008
Ctenophorus butleri Wilson & Swan 2010
Ctenophorus butlerorum [justified emendation] Melville, Wilson, Ford & Macdonald 2017

Remarks: Previously regarded as a subspecies of parviceps (e.g., Aplin & Smith 2001). Often treated as a separate species (e.g., Storr, Smith & Johnstone 1983; Wilson & Knowles 1988, Wilson & Swan 2010). Revalidated by Melville, Shoo & Doughty (2008).
Distribution: Australia (Western Australia).


Ctenophorus caudicinctus

Ring-tailed Dragon, Western Ring-tailed Dragon

Ringelschwanz-Agame

Ringhalet Kamagam

1875 Grammatophora caudicincta Günther
Amphibolurus caudicinctus Boulenger 1885
Ctenophorus caudicinctus Storr, Smith & Johnstone 1983
Phthanodon caudicinctus Wells & Wellington 1984
Tachyon caudicinctus Wells & Wellington 1985
1876 Amphibolurus imbricatus Peters (syn.Cogger, Cameron & Cogger 1983)
1967 Amphibolurus caudicinctus mensarum Storr (syn. Melville, Hainesm, Hale, Chapple & Ritchie 2016)
Phthanodon mensarum Wells & Wellington 1984
Ctenophorus caudicinctus mensarum Cogger 1996

Other common names:
mensarum: Murchison Rock Dragon
Remarks: Previously included graafi, infans, and slateri.
Distribution: Australia (Western Australia).


Ctenophorus chapmani

Southern Heath Dragon, Eastern Heath Dragon, Prickly Dragon, (Chapman's Dragon)

Südliche Heideagame

Sydlig Hedeagam

1977 Amphibolurus adelaidensis chapmani Storr
Rankinia chapmani Wells & Wellington 1984
Amphibolurus chapmani Greer 1989
Ctenophorus chapmani Melville, Schulte & Larson 2001

Remarks: Raised to species status by Melville, Shoo & Doughty (2008), although already Wells & Wellington (1984) had treated it as such.
Distribution: Australia (South Australia, Western Australia).


Ctenophorus clayi

Black-shouldered Ground Dragon, Black-collared Dragon, (Clay's Dragon)

Schwarzschulter-Agame

Sortskuldret Kamagam

1966 Amphibolurus clayi Storr
Ctenophorus clayi Storr, Smith & Johnstone 1983
1985 Ctenophorus raffertyi Wells & Wellington (syn.Shea & Sadlier 1999)

Distribution: Australia (Northern Territory, South Australia, Western Australia).


Ctenophorus cristatus

Crested Dragon, Crested Bicycle Dragon

Kamm-Soldatenagame

Almindelig Kamagam

1841 Grammatophora cristata Gray
Amphibolurus cristatus Boulenger 1885
Ctenophorus cristatus Storr, Smith & Johnstone 1983
Licentia cristata Wells & Wellington 1984

Distribution: Australia (South Australia, Western Australia).


Ctenophorus decresii

Tawny Crevice Dragon, Tawny Rock Dragon, Tawny Dragon

Gelbbraune Felsenagame

Gulbrun Kamagam

1837 Grammatophora decresii Dumeril & Bibron
Ctenophorus decresii Fitzinger 1843
Amphibolurus decresii Peters 1864
1843 Agama decresiensis Fitzinger [substitute name for Grammatophora decresii Duméril & Bibron 1837]
1926 Amphibolurus modestus Ahl (syn.Cogger, Cameron & Cogger 1983)

Remarks: Now restricted to South Australia. Previously included the populations now known as mirrityana. Records from New South Wales refer to that species (McLean, Moussalli, Sass & Stuart-Fox 2013).
Distribution: Australia (South Australia [incl. Kangaroo Island]).


Ctenophorus femoralis

Long-tailed Sand Dragon

Langschwanz-Sandagame

Langhalet Kamagam

1965 Amphibolurus femoralis Storr
Ctenophorus femoralis Storr, Smith & Johnstone 1983
Phthanodon femoralis Wells & Wellington 1984

Distribution: Australia (Western Australia).


Ctenophorus fionni

Peninsula Dragon

Eyre-Halbinsel-Agame

Sydaustralsk Kamagam

1923 Amphibolurus fionni Procter
Ctenophorus fionni Swanson 1987

Distribution: Australia (South Australia).


Ctenophorus fordi

Mallee Dragon, Mallee Sand Dragon, Mallee Military Dragon

Mallee-Soldatenagame

Mallee-soldateragam

1965 Amphibolurus fordi Storr
Ctenophorus fordi Storr, Smith & Johnstone 1983
Phthanodon fordi Wells & Wellington 1984
1985 Phthanodon hawkeswoodi Wells & Wellington (syn.Shea & Sadlier 1999)

Distribution: Australia (New South Wales, South Australia, Victoria, Western Australia).


Ctenophorus gibba

Gibber Dragon, Bulldust Ground Dragon

Wüsten-Bodenagame

Ørken-kamagam

1974 Amphibolurus gibba Houston
Ctenophorus gibba Wilson & Knowles 1988

Distribution: Australia (South Australia).


Ctenophorus graafi

Mount Eveline Dragon, (Graaf's Ring-tailed Dragon)

Mount-Eveline-Agame

Mount Eveline-kamagam

1967 Amphibolurus caudicinctus graafi Storr
Phthanodon graafi Wells & Wellington 1984
Tachyon graafi Wells & Wellington 1985
Ctenophorus caudicinctus graafi Cogger 1996
Ctenophorus graafi Melville, Hainesm, Hale, Chapple & Ritchie 2016

Remarks: Previously regarded as a subspecies of caudicinctus. Raised to species status by Melville, Hainesm, Hale, Chapple & Ritchie (2016).
Distribution: Australia (Western Australia).





Ctenophorus infans

Laverton Dragon, Laverton Ring-tailed Dragon, Southern Murchison Rock Dragon

Laverton-Agame

Laverton-kamagam

1967 Amphibolurus caudicinctus infans Storr
Phthanodon infans Wells & Wellington 1984
Ctenophorus caudicinctus infans Cogger 1996
Ctenophorus infans Melville, Hainesm, Hale, Chapple & Ritchie 2016

Remarks: Previously regarded as a subspecies of caudicinctus. Raised to species status by Melville, Hainesm, Hale, Chapple & Ritchie (2016).
Distribution: Australia (Western Australia).





Ctenophorus isolepis

Central Military Dragon, Military Sand Dragon

Zentralaustralische Soldatenagame

Centralaustralsk Soldateragam

1881 Grammatophora isolepis Fischer
Amphibolurus isolepis Storr 1965
Ctenophorus isolepis Storr, Smith & Johnstone 1983
Phthanodon isolepis Wells & Wellington 1984
1924 Amphibolurus maculatus gularis Sternfeld (syn.Cogger 1996)
Amphibolurus isolepis gularis Storr 1965
Ctenophorus isolepis gularis Storr, Smith & Johnstone 1983
Phthanodon gularis Wells & Wellington 1984
1965 Amphibolurus isolepis citrinus Storr
Ctenophorus isolepis citrinus Storr, Smith & Johnstone 1983
Phthanodon citrinus Wells & Wellington 1984

Other common names:
citrinus: Goldfields Military Sand Dragon
Remarks: Previously included rubens.
Distribution: Australia (Northern Territory, Queensland, South Australia, Western Australia).


Ctenophorus maculatus

Spotted Military Dragon, Spotted Dragon

Gefleckte Soldatenagame

Plettet Soldateragam

1831 Uromastyx maculatus Gray
Grammatophora maculata Gray 1867
Amphibolurus maculatus Boulenger 1885
Ctenophorus maculatus Storr, Smith & Johnstone 1983
Phthanodon maculatus Wells & Wellington 1984
1837 Grammatophora gaimardii Duméril & Bibron (syn.Cogger, Cameron & Cogger 1983)
1965 Amphibolurus maculatus badius Storr
Phthanodon badius Wells & Wellington 1984
1965 Amphibolurus maculatus dualis Storr
Phthanodon dualis Wells & Wellington 1984
1965 Amphibolurus maculatus griseus Storr
Phthanodon griseus Wells & Wellington 1984

Other common names:
dualis: Nullarbor Spotted Sand Dragon
griseus: Wheatbelt Spotted Sand Dragon
maculatus: Coastal Spotted Sand Dragon
Distribution: Australia (South Australia, Western Australia).


Ctenophorus maculosus

Lake Eyre Dragon

Lake Eyre-Agame

Lake Eyre-agam

1948 Tympanocryptis maculosa Mitchell
Amphibolurus maculosus Cogger 1967
Ctenophorus maculosus Wilson & Knowles 1988

Distribution: Australia (South Australia).


Ctenophorus mckenziei

Dwarf Bicycle Dragon, (McKenzie's Chenopod Dragon, McKenzie's Dragon)

Zwerg-Soldatenagame

Dværg-kamagam

1981 Amphibolurus mckenziei Storr
Ctenophorus mckenziei Storr, Smith & Johnstone 1983
Licentia mckenziei Wells & Wellington 1984

Distribution: Australia (South Australia, Western Australia).


Ctenophorus mirrityana

Barrier Range Dragon

Barrier-Range-Felsengame

Barrier Range-kamagam

2013 Ctenophorus mirrityana McLean, Moussalli, Sass & Stuart-Fox

Distribution: Australia (New South Wales).


Ctenophorus nguyarna

Lake Disappointment Dragon

Lake-Disappointment-Agame

Nguyarna-kamagam

2007 Ctenophorus nguyarna Doughty, Maryan, Melville & Austin

Distribution: Australia (Western Australia).


Ctenophorus nuchalis

Central Netted Dragon, Central Netted Ground Dragon

Zentralaustralische Netzagame

Centralaustralsk Netagam

1884 Macrops nuchalis De Vis
Amphibolurus nuchalis Rankin 1977
Ctenophorus nuchalis Swanson 1987
1888 Grammatophora inermis De Vis (syn.Cogger, Cameron & Cogger 1983)
Amphibolurus reticulatus inermis Loveridge 1934
Amphibolurus inermis Storr 1966
Ctenophorus inermis Storr, Smith & Johnstone 1983
1919 Amphibolurus reticulatus major Sternfeld (syn.Cogger, Cameron & Cogger 1983)
Ctenophorus major Wells & Wellington 1984

Other common names:
inermis: Defenseless Dragon
Remarks: Some authors have maintained to refer to this species as inermis (e.g., Storr, Hanlon & Dunlop 1983; Storr, Smith & Johnstone 1983; Storr & Harold 1984; Chapman & Dell 1985), although inermis is clearly a junior synonym of nuchalis.
Distribution: Australia (New South Wales, Northern Territory, Queensland, South Australia, Western Australia).


Ctenophorus ornatus

Ornate Crevice Dragon, Ornate Dragon

Pracht-Felsenagame

Pragt-kamagam

1875 Grammatophora ornata Gray
Amphibolurus ornatus Boulenger 1885
Ctenophorus ornatus Storr, Smith & Johnstone 1983
1926 Amphibolurus tibialis Ahl (syn.Cogger, Cameron & Cogger 1983)

Distribution: Australia (Western Australia).


Ctenophorus parviceps

Northwestern Heath Dragon, Gnaraloo Earless Dragon, Gnaraloo Heath Dragon

Nordwestliche Heideagame

Nordvestlig Hedeagam

1964 Tympanocryptis parviceps Storr
Amphibolurus parviceps Storr 1977
Rankinia parviceps Wells & Wellington 1985
Ctenophorus parviceps Wilson & Swan 2010

Remarks: Previously included butlerorum.
Distribution: Australia (Western Australia).


Ctenophorus pictus

Painted Dragon, Painted Ground Dragon

Gemalte Bartagame

Malet Kamagam

1866 Amphibolurus pictus Peters
Grammatophora picta Günther 1875
Ctenophorus pictus Storr, Smith & Johnstone 1983
1985 Ctenophorus dudleyi Wells & Wellington (syn.Shea & Sadlier 1999)

Distribution: Australia (New South Wales, Northern Territory, Queensland, South Australia, Victoria, Western Australia).


Ctenophorus reticulatus

Western Netted Dragon, Western Netted Ground Dragon

Westaustralische Netzagame

Vestlig Netagam

1845 Grammatophora reticulata Gray
Amphibolurus reticulatus Boulenger 1885
Ctenophorus reticulatus Storr, Smith & Johnstone 1983
Licentia reticulata Wells & Wellington 1984
1867 Grammatophora laevis Günther (syn.Cogger, Cameron & Cogger 1983)
1932 Amphibolurus darlingtoni Loveridge (syn.Storr 1966; Cogger, Cameron & Cogger 1983)

Distribution: Australia (Northern Territory, South Australia, Western Australia).


Ctenophorus rubens

Greater Military Dragon, Russet Dragon

Große Soldatenagame

Stor Soldateragam

1965 Amphibolurus isolepis rubens Storr
Amphibolurus rubens Storr, Smith & Johnstone 1983
Phthanodon rubens Wells & Wellington 1984
Ctenophorus rubens Aplin & Smith 2001

Distribution: Australia (Western Australia).


Ctenophorus rufescens

Rusty Crevice Dragon, Rusty Dragon, Bicycle Lizard

Rost-Felsenagame

Rustrød Kamagam

1893 Amphibolurus rufescens Stirling & Zietz
Ctenophorus rufescens Storr, Smith & Johnstone 1983
Licentia rufescens Wells & Wellington 1984

Distribution: Australia (Northern Territory, South Australia, Western Australia).


Ctenophorus salinarum

Saltpan Ground Dragon, Claypan Dragon, Salt Lake Dragon

Salzsee-Bodenagame

Saltsø-agam

1966 Amphibolurus pictus salinorum Storr
Amphibolurus salinarum Cogger, Cameron & Cogger 1983
Ctenophorus salinarum Storr, Smith & Johnstone 1983

Distribution: Australia (South Australia, Western Australia).


Ctenophorus scutulatus

Lozenge-marked Dragon, Lozenge-marked Bicycle Dragon

Rauten-Agame

Rudemønstret Kamagam

1893 Amphibolurus scutulatus Stirling & Zietz
Ctenophorus scutulatus Storr, Smith & Johnstone 1983
Licentia scutulata Wells & Wellington 1984
1904 Amphibolurus websteri Boulenger (syn.Cogger, Cameron & Cogger 1983)
Licentia websteri Wells & Wellington 1984
1905 Amphibolurus holsti Rosén (syn.Loveridge 1934; Cogger, Cameron & Cogger 1983)

Remarks: Presence in South Australia needs confirmation. Known from only a single specimen (Loveridge 1938) which has been lost (Houston & Hutchinson 1998).
Distribution: Australia (Western Australia).


Ctenophorus slateri

Hermannsburg Dragon, (Slater's Ring-tailed Dragon)

Hermannsburg-Agame

Hermannsburg Kamagam

1967 Amphibolurus caudicinctus slateri Storr
Phthanodon slateri Wells & Wellington 1984
Ctenophorus caudicinctus slateri Cogger 1996
Ctenophorus slateri Melville, Hainesm, Hale, Chapple & Ritchie 2016
1967 Amphibolurus caudicinctus macropus Storr (syn. Melville, Hainesm, Hale, Chapple & Ritchie 2016)
Phthanodon macropus Wells & Wellington 1984
Ctenophorus caudicinctus macropus Cogger 1996

Remarks: Previously regarded as a subspecies of caudicinctus. Raised to species status by Melville, Hainesm, Hale, Chapple & Ritchie (2016).
Distribution: Australia (Northern Territory, Queensland, South Australia, Western Australia).





Ctenophorus spinodomus

Eastern Mallee Dragon

Östliche Mallee-Soldatenagame

Østlig Mallee-soldateragam

2019 Ctenophorus spinodomus Sadlier, Colgan, Beatson & Cogger

Distribution: Australia (New South Wales, South Australia).





Ctenophorus tjantjalka

Ochre Dragon

Ocker-Agame

Okkergul Kamagam

1992 Ctenophorus tjantjalka Johnston

Distribution: Australia (South Australia).


Ctenophorus vadnappa

Red-barred Dragon

Rotband-Agame

Rødbåndet Kamagam

1974 Amphibolurus vadnappa Houston
Ctenophorus vadnappa Wilson & Knowles 1988

Distribution: Australia (South Australia).


Ctenophorus yinnietharra

Yinnietharra Crevice Dragon, Yinnietharra Rock Dragon

Yinnietharra-Felsenagame

Yinnietharra-kamagam

1981 Amphibolurus yinnietharra Storr
Ctenophorus yinnetharra Storr, Smith & Johnstone 1983
Licentia yinnietharra Wells & Wellington 1984

Distribution: Australia (Western Australia).