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© Rune Midtgaard


Taxonomy of the family Agamidae
Bibliography of the genus Diploderma
Biodiversity of the family Agamidae








Genus
Diploderma

Eastern Mountain Agamas, (Mountain Dragons)

Östliche Bergwaldagamen

Østlige Bjergagamer

1861 Diploderma Hallowell (type species: Diploderma polygonatum Hallowell 1861)
Contents: 27 species, of which 24 (88.9%) are endemic.
Remarks: Previously regarded as a synonym of Japalura by (e.g., Wermuth 1967). Revalidated as a separate genus by Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler (2018).
Distribution: China (Chongqing, Gansu, Guangxi, Guizhou, Henan, Hubei, Hunan, Qinghai, Shaanxi, Shanxi, Sichuan, Tibet, Yunnan), Japan (Ryukyu Islands [Okinawa Islands, Sakishima Islands (Miyako Islands, Yaeyama Islands), Satsunan Islands]), Myanmar, Taiwan (incl. Lanyu Island), Thailand, Vietnam.

Diploderma batangense

Batang Mountain Agama

Batang-Bergwaldagame

Batang-bjergagam

2001 Japalura batangensis Li, Deng, Wu & Wang
Diploderma batangense Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Sichuan, Tibet).


Diploderma brevicaudum

Short-tailed Mountain Agama

Kurzschwanz-Bergwaldagame

Korthalet Bjergagam

2012 Japalura brevicauda Manthey, Denzer, Hou & Wang
Diploderma brevicaudum Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Yunnan).


Diploderma brevipes

Short-legged Mountain Agama

Kurzbeinige Bergwaldagame

Kortbenet Bjergagam

1936 Japalura brevipes Gressitt
Diploderma brevipes Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Validity as a separate species confirmed by Ota (1989).
Distribution: Taiwan.


Diploderma chapaense

Chapa Mountain Agama, Sapa Japalure

Chapa-Bergwaldagame

Chapa-bjergagam

1937 Japalura swinhonis chapaensis Bourret
Japalura chapaensis Ota 1989
Diploderma chapaense Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Previously regarded as a synonym of swinhonis. Revalidated by Ota (1989). Validity confirmed by Ota & Weidenhöfer (1992), however, some authors have continued to list it as a subspecies of swinhonis (e.g., Nguyen & Ho 1996; Nguyen, Ho & Nguyen 2005).
Distribution: Vietnam.


Diploderma drukdaypo

Chamdo Mountain Agama

Chamdo-Bergwaldagame

Chamdo-bjergagam

2019 Diploderma drukdaypo Wang, Jiang, Ren, Zou, Wu, Che & Siler

Distribution: China (Tibet).





Diploderma dymondi

Northern Yunnan Mountain Agama, (Dymond's Mountain Agama)

Nördliche Yunnan-Bergwaldagame

Nordlig Yunnan-bjergagam

1906 Acanthosaura dymondi Boulenger
Japalura dymondi Pope 1935
Diploderma dymondi Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Sichuan, Yunnan).


Diploderma fasciatum

Banded Mountain Agama

Gebänderte Bergwaldagame

Båndet Bjergagam

1926 Japalura fasciata Mertens
Diploderma fasciatum Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018
1966 Japalura szechwanensis Hu & Zhao (Ota 2000)

Distribution: China (Guangxi, Guizhou, Hunan, Sichuan), Vietnam.


Diploderma flaviceps

Yellow-headed Mountain Agama

Gelbkopf-Bergwaldagame

Gulhovedet Bjergagam

1919 Japalura flaviceps Barbour & Dunn
Japalura yunnanensis flaviceps Mell 1929
Diploderma flaviceps Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Wang, Jiang, Zou, Yan, Siler & Che (2016) claimed that the true flaviceps is restricted to Sichuan, and that the remaining populations represent undescribed species. However, since detailed studies of these populations have still not been published, they are provisionally retained herein as flaviceps.
Distribution: China (Chongqing, Gansu, Hubei, Qinghai, Shaanxi, Shanxi, Sichuan, Tibet, Yunnan).


Diploderma grahami

Sichuan Mountain Agama, (Graham's Mountain Agama)

Sichuan-Bergwaldagame

Sichuan-bjergagam

1924 Phoxophrys grahami Stejneger
Japalura grahami Inger 1960
Diploderma grahami Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Sichuan).


Diploderma hamptoni

Mogok Mountain Agama, (Hampton's Mountain Agama)

Mogok-Bergwaldagame

Mogok-bjergagam

1935 Japalura hamptoni Smith
Diploderma hamptoni Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: Myanmar.


Diploderma iadinum

Emerald Mountain Agama, Emerald Mountain Dragon

Smaragd-Bergwaldagame

Smaragd-bjergagam

2016 Japalura iadina Wang, Jiang, Siler & Che in Wang, Jiang, Zou, Yan, Siler & Che
Diploderma iadinum Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Yunnan).


Diploderma laeviventre

Smooth-bellied Mountain Agama, Smooth-venter Mountain Dragon

Glattbauch-Bergwaldagame

Glatbuget Bjergagam

2016 Japalura laeviventris Wang, Jiang, Siler & Che in Wang, Jiang, Zou, Yan, Siler & Che
Diploderma laeviventre Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Tibet).


Diploderma luei

Taipingshan Mountain Agama

Taipingshan-Bergwaldagame

Taipingshan-bjergagam

1998 Japalura luei Ota, Chen & Shang
Diploderma luei Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: Taiwan.


Diploderma makii

Chitou Mountain Agama, (Makis' Mountain Agama)

Chitou-Bergwaldagame

Chiyou-bjergagam

1989 Japalura makii Ota
Diploderma makii Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: Taiwan.


Diploderma menghaiense

Menghai Mountain Agama

Menghai-Bergwaldagame

Menghai-bjergagam

2020 Diploderma menghaiense Liu, Hou, Wang, Ananjeva & Rao

Distribution: China (Yunnan).





Diploderma micangshanense

Shaanxi Mountain Agama

Shaanxi-Bergwaldagame

Shaanxi-bjergagam

1987 Japalura micangshanensis Song
Diploderma micangshanense Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Gansu, Henan, Shaanxi, Sichuan).


Diploderma ngoclinense

Vietnamese Mountain Agama

Vietnamesische Bergwaldagame

Vietnamesisk Bjergagam

2017 Japalura ngoclinensis Ananjeva, Orlov & Nguyen 2017
Diploderma ngoclinense Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler (2018) questioned the validity of ngoclinense, which is closely related to (and possibly synonymous with) splendidum.
Distribution: Vietnam.


Diploderma polygonatum

Japanese Mountain Agama, Okinawan Tree Lizard

Japanische Bergwaldagame

Japansk Bjergagam

1861 Diploderma polygonatum Hallowell
Japalura polygonata Günther 1864
1912 Japalura polygonata ishigakiensis Van Denburgh
1912 Japalura polygonata miyakensis Van Denburgh
1991 Japalura polygonata xanthostoma Ota
2003 Japalura polygonata donan Ota

Other common names:
donan: Yonagunijima Mountain Agama
ishigakiense: Ishigakijima Mountain Agama, Sakishima Tree Lizard
polygonatum: Okinawajima Mountain Agama
Remarks: The Taiwanese population (subspecies xanthostomum) was previously confused with swinhonis (Ota 1991).
Introduced to: Japan (Kyushu, Ryukyu Islands [Satsunan Islands: Osumi Islands]).
Distribution: Japan (Ryukyu Islands [Okinawa Islands, Sakishima Islands (Miyako Islands, Yaeyama Islands), Satsunan Islands]), Taiwan.


Diploderma slowinskii

Nujiang Mountain Agama

Nujiang-Bergwaldagame

Nujiang-bjergagam

2017 Japalura slowinskii Rao, Vindum, Ma, Fu & Wilkinson
Diploderma slowinskii Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Previously regarded as a population of splendidum (Rao, Vindum, Ma, Fu & Wilkinson 2017).
Distribution: China (Yunnan).


Diploderma splendidum

Splendid Mountain Agama

Pracht-Bergwaldagame

Pragt-bjergagam

1919 Japalura splendida Barbour & Dunn
Japalura yunnanensis splendida Mell 1929
Diploderma splendidum Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Previously included the populations now assigned to slowinskii (Rao, Vindum, Ma, Fu & Wilkinson 2017).
Distribution: China (Chongqing, Gansu, Guizhou, Henan, Hubei, Hunan, Sichuan, Tibet, Yunnan).


Diploderma swild

Yanbian Mountain Dragon, (Swild Mountain Dragon)

Yanbian-Bergwaldagame

Yanbian-bjergagam

2019 Diploderma swild Wang, Wu, Jiang, Chen, Miao, Siler & Che

Distribution: China (Sichuan).





Diploderma swinhonis

Tamsui Mountain Agama, (Swinhoe's Mountain Agama)

Tamsui-Bergwaldagame

Tamsui-bjergagam

1864 Japalura swinhonis Günther
Diploderma swinhonis Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018
1885 Japalura swinhoei Boettger [substitute name for Japalura swinhonis Günther 1864] (Wermuth 1967)
Japalura polygonata swinhonis Nakamura & Ueno 1963
1898 Japalura mitsukurii Stejneger (Ota 1991)
Japalura swinhonis mitsukurii Denburgh 1912
Japalura polygonata mitsukurii Nakamura & Uéno 1963
1938 Japalura yunnanensis popei Wettstein (Ota 2000)
1976 Japalura swinhonis formosensis Liang & Wang (Ota 1988)
Japalura mitsukurii formosensis Chen 1984

Remarks: The Taiwanese population was previously confused with polygonatum (Ota 1991). J. y. popei Wettstein 1938 is known from only three specimens believed to have been collected in Cambodia. Ota (2000) suggested they actually came from Taiwan, a zoogeographically more likely place of origin.
Distribution: Taiwan (incl. Lanyu Island).


Diploderma varcoae

Chinese Mountain Agama, (Varcoe's Mountain Agama)

Chinesische Bergwaldagame

Kinesisk Bjergagam

1918 Acanthosaura varcoæ Boulenger
Japalura varcoae Smith 1935
Diploderma varcoae Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Guizhou, Yunnan).


Diploderma vela

Quzika Mountain Agama, (Sail Mountain Agama)

Quzika-Bergwaldagame

Quzika-bjergagam

2015 Japalura vela Wang, Jiang, Pan, Hou, Siler & Che
Diploderma vela Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Tibet).


Diploderma yulongense

Yulong Mountain Agama

Yulong-Bergwaldagame

Yulong-bjergagam

2012 Japalura yulongensis Manthey, Denzer, Hou & Wang
Diploderma yulongense Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Distribution: China (Yunnan).


Diploderma yunnanense

Southern Yunnan Mountain Agama, (Yunnan Mountain Agama)

Südliche Yunnan-Bergwaldagame

Sydlig Yunnan-bjergagam

1878 Japalura yunnanensis Anderson
Diploderma yunnanense Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Previously included J. y. popei Wettstein 1938, now regarded as a synonym of J. swinhonis. Records from Cambodia (e.g., Bourret 1939; Wermuth 1967; Welch, Cooke & Wright 1990; Zhao & Adler 1993) and Vietnam (e.g., Bobrov 1995; Ota & Weidenhöfer 1990) are considered erroneous (Ota 2000).
Distribution: China (Yunnan), Myanmar, Thailand.


Diploderma zhaoermii

Mianchi Mountain Agama, (Zhao's Mountain Agama)

Mianchi-Bergwaldagame

Mianchi-bjergagam

2002 Japalura zhaoermii Gao & Hou
Diploderma zhaoermii Wang, Che, Lin, Deepak, Datta-Roy, Jiang, Jin, Chen & Siler 2018

Remarks: Not mentioned by Ananjeva, Guo & Wang (2011).
Distribution: China (Sichuan).