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© Rune Midtgaard


Taxonomy of the family Leiosauridae
Bibliography of the genus Enyalius
Biodiversity of the family Leiosauridae








Genus
Enyalius

Fat-headed Anoles

Dickkopfanolis

Tykhovedanoler*

1830 Enyalius Wagler (type species: Agama catenata Wied 1821)
1843 Dryophilus Fitzinger (type species: Enyalus bilineatus Dumeril & Bibron 1837; syn. Peters & Donoso-Barros 1970)
1933 Garbesaura Amaral (type species: Garbesaura garbei Amaral 1933; syn. Vanzolini 1973)
Contents: 11 species, all of which are endemic.
Distribution: E. South America.
Reported from: Brazil (Alagoas, Bahia, Ceara, Distrito Federal, Espirito Santo, Goias, Maranhao, Mato Grosso, Minas Gerais, Para, Paraiba, Parana, Pernambuco, Piaui, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Rondonia, Santa Catarina, Sao Paulo, Sergipe).

Enyalius bibronii

Bahia Fat-headed Anole*

Bahia-Dickkopfanolis*

Bahia-tykhovedanole*

1885 Enyalius bibronii Boulenger
Enyalius catenatus bibronii Jackson 1978

Remarks: Previously regarded as a subspecies of catenatus (e.g., Jackson 1978), although some continued to regard bibronii as a valid species (e.g., Freire 1997). Revalidated by Rodrigues, Freitas, Silva & Bertolotto (2006).
Distribution: Brazil (Alagoas, Bahia, Ceara, Minas Gerais, Paraiba, Pernambuco, Piaui, Rio Grande do Norte, Sergipe).


Enyalius bilineatus

Two-lined Fat-headed Anole*

Zweistreifiger Dickkopfanolis*

Tostribet Tykhovedanole*

1837 Enyalus bilineatus Dumeril & Bibron
Hypsibatus bilineatus Fitzinger 1843
1885 Enyalius fitzingeri Boulenger (Jackson 1978)

Remarks: A record from Espirito Santo (Etheridge 1969) refers to pictus (Jackson 1978; Gogliath, Ribeiro & Freire 2010).
Distribution: Brazil (Bahia, Distrito Federal, Espirito Santo, Goias, Minas Gerais, Rio de Janeiro, Sao Paulo).


Enyalius boulengeri

Espirito Santo Fat-headed Anole*

Espirito-Santo-Dickkopfanolis*

Espirito Santo-tykhovedanole*

1968 Enyalius boulengeri Etheridge
Enyalius brasiliensis boulengeri Jackson 1978

Remarks: Revalidated as a separate species by Rodrigues, Bertolotto, Amaro, Yonenaga-Yassuda, Freire & Pellegrino (2014).
Distribution: Brazil (Espirito Santo, Minas Gerais).


Enyalius brasiliensis

Brazilian Fat-headed Anole*

Brasilianischer Dickkopfanolis*

Brasiliansk Tykhovedanole*

1828 Lophyrus brasiliensis Lesson
Enyalius brasiliensis Etheridge 1969
1837 Enyalius rhombifer Duméril & Bibron [not Lophyrus rhombifer Spix 1825] (Jackson 1978)

Remarks: Previously included boulengeri. Presence in Goias (Jackson 1978) and Santa Catarina (Etheridge 1969; Jackson 1978) is considered doubtful (Costa & Bérnils 2018).
Distribution: Brazil (Minas Gerais, Rio de Janeiro).


Enyalius capetinga

Cerrado Fat-headed Anole*

Cerrado-Dickkopfanolis*

Cerrado-tykhovedanole*

2018 Enyalius capetinga Breitman, Domingos, Bagley, Wiederhecker, Ferrari, Cavalcante, Pereira, Abreu, De-Lima, Morais, Prette, Silva, Mello, Carvalho, Lima, Silva, Matias, Carvalho, Pantoja, Gomes, Paschoaletto, Rodrigues, Talarico, Barreto-Lima & Colli

Distribution: Brazil (Distrito Federal, Minas Gerais).





Enyalius catenatus

Chain-backed Fat-headed Anole*, Brazilian Chameleon

Kettenband-Dickkopfanolis*

Kædebåndet Tykhovedanole*

1821 Agama catenata Wied
Ophryessa catenata Fitzinger 1826
Uraniscodon catenata Kaup 1827
Enyalius catenata Wagler 1830
Hypsibatus catenata Fitzinger 1843
Enyalius catenata Boulenger 1885
1825 Lophyrus albomaxillaris Spix (Jackson 1978)
1825 Lophyrus margaritaceus Spix (Jackson 1978)
1825 Lophyrus rhombifer Spix (Jackson 1978)
1898 Enyalius catenatus paulista Ihering (Peters & Donoso-Barros 1970)
1926 Enyalius zonatus Wettstein (Jackson 1978)

Remarks: Previously included bibronii and pictus. Presence in Goias (Jackson 1978, as catenatus X pictus) is considered doubtful (Costa & Bérnils 2018).
Distribution: Brazil (Alagoas, Bahia, Espirito Santo, Minas Gerais, Paraiba, Pernambuco, Rio Grande do Norte).


Enyalius erythroceneus

Red-flanked Fat-headed Anole*

Rotseitiger Dickkopfanolis*

Rødsidet Tykhovedanole*

2006 Enyalius erythroceneus Rodrigues, de Freitas, Santos Silva & Bertolotto

Distribution: Brazil (Bahia).


Enyalius iheringii

Southern Fat-headed Anole*

Südlicher Dickkopfanolis*

Sydlig Tykhovedanole*

1885 Enyalius iheringii Boulenger

Distribution: Brazil (Parana, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, Sao Paulo).


Enyalius leechii

Hexagon Fat-headed Anole*

Hexagon-Dickkopfanolis*

Hexagon-tykhovedanole*

1885 Enyalioides leechii Boulenger
Enyalius leechii Etheridge 1969
1933 Garbesaura garbei Amaral (Vanzolini 1973; Avila-Pires 1995)

Distribution: Brazil (Maranhao, Mato Grosso, Para, Rondonia).


Enyalius perditus

Boraceia Fat-headed Anole*

Boraceia-Dickkopfanolis*

Boraceia-tykhovedanole*

1978 Enyalius perditus Jackson

Distribution: Brazil (Espirito Santo, Minas Gerais, Parana, Rio de Janeiro, Sao Paulo).


Enyalius pictus

Painted Fat-headed Anole*

Gemalter Dickkopfanolis*

Malet Tykhovedanole*

1822 Agama picta Schinz
Enyalius pictus Peters & Donoso-Barros 1970
Enyalius catenatus pictus Jackson 1978

Remarks: Revalidated by Rodrigues, de Freitas, Silva & Bertolotto (2006). Authorship previously contributed to Wied 1923 (see Myers, Rodrigues & Vanzolini 2011).
Distribution: Brazil (Bahia, Espirito Santo, Minas Gerais).