Reptiles of Goias [Brazil] Related checklists:  Brazil Bibliography of the reptiles of Goias Taxonomic history of the reptiles of Goias Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Amphisbaena (Neotropical Worm Lizards)   Amphisbaena alba Common Red Worm Lizard Araujo-Esteves, Brandao & Viegas 2008 Barcelos-Ribeiro, Santos & Vaz-Silva 2007 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Gans 1962 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Mott & Vieites 2009 Mott 2006 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Amphisbaena anaemariae Serra da Mesa Worm Lizard Barcelos-Ribeiro, Santos & Vaz-Silva 2007 [as neglecta] Costa, Guedes & Bernils 2021 Gans 1962 [as neglecta] Gans 2005 Gomes & Maciel 2012 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Teixeira, Vechio, Neto & Rodrigues 2014 Vanzolini 1997   Amphisbaena crisae Cafeara Worm Lizard Costa, Guedes & Bernils 2021   Amphisbaena fuliginosa Spotted Worm Lizard Araujo-Esteves, Brandao & Viegas 2008 Barcelos-Ribeiro, Santos & Vaz-Silva 2007 Costa, Guedes & Bernils 2021 Moreira, Fenolio, Silva & Silva 2009 Roberto, Muniz Silva, Pinto & Avila 2021 Silveira 2007 Vanzolini 2002   Amphisbaena leeseri Urucum Worm Lizard Costa, Guedes & Bernils 2021   * Amphisbaena mebengokre Arenopolis Worm Lizard Costa, Guedes & Bernils 2021 Ribeiro, Sa, Santos-Jr., Graboski, Zaher, Guedes, Andrade & Vaz-Silva 2019   Amphisbaena mertensii Sao Paulo Worm Lizard Barcelos-Ribeiro, Santos & Vaz-Silva 2007 Costa, Guedes & Bernils 2021 Lima Silveira, Campos Brites & Valinhas e Valinhas 2012 Moreira, Fenolio, Silva & Silva 2009 Pena Correa & Meneses 2020 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Amphisbaena silvestrii Southern Mato Grosso Worm Lizard Costa, Guedes & Bernils 2021 Teixeira, Vechio, Neto & Rodrigues 2014 Valdujo & Nogueira 2001   Amphisbaena talisiae Serra da Pitomba Worm Lizard Barcelos-Ribeiro, Santos & Vaz-Silva 2007 [as A. mensae] Castro-Mello 2000 [as A. mensae] Costa, Grabosky & Zaher 2019 Costa, Guedes & Bernils 2021 Gans 2005 [as A. mensae] Moreira, Fenolio, Silva & Silva 2009 [as A. mensae]   Amphisbaena vermicularis Vermiculate Worm Lizard Araujo-Esteves, Brandao & Viegas 2008 Barcelos-Ribeiro, Santos & Vaz-Silva 2007 Costa, Guedes & Bernils 2021 Gans & Amdur 1966 Gans 1967 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Peters & Donoso-Barros 1970 Genus Cercolophia (Keel-tailed Worm Lizards)   Cercolophia roberti Ipiranga Worm Lizard Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Gans 1967 Peters & Donoso-Barros 1970 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Leposternon (South American Worm Lizards)   * Leposternon cerradense Cerrado Worm Lizard Costa, Guedes & Bernils 2021 Ribeiro, Nogueira, Cintra, Silva & Zaher 2011 Ribeiro, Vaz-Silva & Santos 2008   Leposternon infraorbitale Central Brazilian Worm Lizard Araujo-Esteves, Brandao & Viegas 2008 Costa, Guedes & Bernils 2021 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Leposternon maximum Buritinopolis Worm Lizard Costa, Guedes & Bernils 2021 Ribeiro, Nogueira, Cintra, Silva & Zaher 2011   Leposternon microcephalum Small-headed Worm Lizard Costa, Guedes & Bernils 2021 Moreira, Fenolio, Silva & Silva 2009 Ribeiro, Nogueira, Cintra, Silva & Zaher 2011   Leposternon polystegum East Brazilian Worm Lizard Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Alligatoridae (Alligators and Caimans) Genus Caiman (Typical Caimans)   Caiman crocodilus Common Caiman Costa, Guedes & Bernils 2021   Caiman latirostris Broad-snouted Caiman Costa, Guedes & Bernils 2021 Genus Melanosuchus (Black Caiman)   Melanosuchus niger Black Caiman Costa, Guedes & Bernils 2021 Ministry of Environment [Brasilia] 2010 Genus Paleosuchus (Dwarf Caimans)   Paleosuchus palpebrosus Cayenne Dwarf Caiman Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Ophiodes (South American Glass Lizards)   Ophiodes striatus Dark-striped Glass Lizard Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 [as "aff. striatus"]  Family Anolidae (Anoles) Genus Norops (Brilliant Anoles)   Norops brasiliensis Brazilian Golden-scaled Anole Avila-Pires 1995 Costa, Guedes & Bernils 2021 D'Angiolella, Gamble, Avila-Pires, Colli, Noonan & Vitt 2011 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro-Junior 2015   Norops meridionalis Paraguayan Anole Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 D'Angiolella, Gamble, Avila-Pires, Colli, Noonan & Vitt 2011 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro-Junior 2015 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007  Family Gymnophthalmidae (Scaly-tongued Spectacled Tejus) Genus Bachia (Earless Tejus)   Bachia bresslaui Sao Paulo Earless Teju Costa, Guedes & Bernils 2021   Bachia cacerensis Caceres Earless Teju Costa, Guedes & Bernils 2021 Valdujo & Nogueira 2001 Genus Cercosaura (Eastern Shade Tejus)   Cercosaura olivacea East Brazilian Shade Teju Costa, Guedes & Bernils 2021 Dantas Sales, Silva Jorge, Meira-Ribeiro & Xavier Freire 2014 [as ocellata] Sturaro, Rodrigues, Colli, Knowles & Avila-Pires 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 [as ocellata]   Cercosaura quadrilineata Four-lined Shade Teju Costa, Guedes & Bernils 2021 Marques-Souza, Prates, Fouquet, Camacho, Kok, Nunes, Vechio, Recoder, Mejia, Teixeira-Junior, Barrio-Amoros, Cassimiro, Lima, Sena & Rodrigues 2018 Pellegrino, Rodrigues, Yonenaga-Yassuda & Sites 2001 Torres-Carvajal, Lobos, Venegas, Chavez, Aguirre-Penafiel, Zurita & Echevarria 2016   Cercosaura schreibersii South Brazilian Shade Teju Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Ribeiro-Junior & Amaral 2017 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Colobosaura (Short-thumbed Spectacled Tejus)   Colobosaura modesta Morro da Garcza Spectacled Teju Avila-Pires 1995 Campos & Lage 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Freire, Jorge & Ribeiro 2012 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Muniz, Vega, Chaves, Moura, Santos & Moura 2012 Ribeiro-Junior & Amaral 2017 Rodrigues, Pellegrino, Dixo, Verdade, Pavan, Argolo & Sites 2007 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Micrablepharus (Snake-eyed Glossy Tejus)   Micrablepharus atticolus Cerrado Snake-eyed Glossy Teju Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro-Junior & Amaral 2017 Rodrigues 1996 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Micrablepharus maximiliani Blue-tailed Snake-eyed Glossy Teju Campos & Lage 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Moura, Dayrell & Avelar Sao Pedro 2010 Ribeiro-Junior & Amaral 2017 Rodrigues 1996 Genus Vanzosaura (Red-tailed Spectacled Tejus)   Vanzosaura rubricauda Southern Red-tailed Spectacled Teju Costa, Guedes & Bernils 2021 Recoder, Werneck, Teixeira, Colli, Sites & Rodrigues 2014  Family Hoplocercidae (Club-tailed Iguanas) Genus Hoplocercus (Eastern Club-tailed Iguana)   Hoplocercus spinosus Eastern Club-tailed Iguana Avila-Pires 1995 Costa, Guedes & Bernils 2021 Ribeiro-Junior 2015  Family Iguanidae (Greater Iguanas) Genus Iguana (Green Iguanas)   Iguana iguana South American Green Iguana Avila-Pires 1995 Costa, Guedes & Bernils 2021 Cunha, Moreira & Sousa-Silva 2010 Ribeiro-Junior 2015  Family Leiosauridae (Smooth Iguanas) Genus Enyalius (Fat-headed Anoles)   Enyalius bilineatus Two-lined Fat-headed Anole Costa, Guedes & Bernils 2021 Rodrigues, Freitas, Silva & Bertolotto 2006 Sales, Ribeiro & Freire 2015   Enyalius capetinga Cerrado Fat-headed Anole Costa, Guedes & Bernils 2021  Family Phyllodactylidae (Leaf-toed Geckos) Genus Gymnodactylus (Naked-toed Geckos)   Gymnodactylus amarali Maranhao Naked-toed Gecko Colli, Mesquita, Rodrigues & Kitayama 2003 Costa, Guedes & Bernils 2021 Gamble, Colli, Rodrigues, Werneck & Simons 2012 Moreira, Fenolio, Silva & Silva 2009 [as carvalhoi] Ribeiro-Junior 2015 Vanzolini 1972 Vanzolini 2004 Vanzolini 2005 [as carvalhoi] Genus Phyllopezus (South American Robust Geckos)   Phyllopezus pollicaris Common Robust Gecko Condez, Tonini, Pereira-Ribeiro & Dubeux 2021 Costa, Guedes & Bernils 2021 Gamble, Colli, Rodrigues, Werneck & Simons 2012 Ribeiro-Junior 2015  Family Polychrotidae (Monkey Lizards) Genus Polychrus (Monkey Lizards)   Polychrus acutirostris Sharp-nosed Monkey Lizard Avila-Pires 1995 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro-Junior 2015 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007  Family Scincidae (Skinks) Genus Aspronema (South American White-striped Skinks)   Aspronema dorsivittatum South American White-striped Skink Costa, Guedes & Bernils 2021 Harvey, Aguayo & Miralles 2008 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Brasiliscincus (Brazilian Small-headed Skinks)   Brasiliscincus heathi Caatinga Small-headed Skink Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Genus Copeoglossum (Neotropical Spotted Skinks)   Copeoglossum nigropunctatum Black-spotted Skink Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Hedges & Conn 2012 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Miralles & Carranza 2010 Miralles, Barrio-Amoros, Rivas & Chaparro-Auza 2006 Moreira, Fenolio, Silva & Silva 2009 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro-Junior & Amaral 2016 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Manciola (South American Small-handed Skink)   Manciola guaporicola South American Small-handed Skink Costa, Guedes & Bernils 2021 Harvey, Aguayo & Miralles 2008 Ribeiro-Junior & Amaral 2016 Genus Notomabuya (Southern Neotropical Skink)   Notomabuya frenata Southern Neotropical Skink Caldeira Costa, Dias Fernandes, Vrcibradic & Neves Feio 2008 Costa, Guedes & Bernils 2021 Harvey, Aguayo & Miralles 2008 Hedges & Conn 2012 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Miralles & Carranza 2010 Moreira, Fenolio, Silva & Silva 2009 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro-Junior & Amaral 2016 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007  Family Sphaerodactylidae (Least Geckos, etc.) Genus Coleodactylus (Sheath-toed Pygmy Geckos)   Coleodactylus brachystoma Cerrado Pygmy Gecko Costa, Guedes & Bernils 2021 Freitas, Franca & Oliveira-Lima 2011 Geurgas, Rodrigues & Moritz 2008 Goncalves, Torquato, Skuk & Sena 2012 Moreira, Fenolio, Silva & Silva 2009 Peters & Donoso-Barros 1970 Wermuth 1965   Coleodactylus meridionalis Atlantic Forest Pygmy Gecko Costa, Guedes & Bernils 2021 Ribeiro-Junior 2015  Family Teiidae (Greater Tejus) Genus Ameiva (South American Ameivas)   Ameiva ameiva South American Giant Ameiva Avila-Pires 1995 Campos & Lage 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Ribeiro-Junior & Amaral 2016 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Ameiva jacuba Jacuba River Ameiva Costa, Guedes & Bernils 2021 Giugliano, Nogueira, Valdujo, Collevatti & Colli 2013 Genus Ameivula (Eastern South American Whiptails)   Ameivula mumbuca Mateiros Whiptail Arias, Carvalho, Zaher & Rodrigues 2014 Arias, Recoder, Alvarez, Etchepare, Quipildor, Lobo & Rodrigues 2018 [as "aff. mumbuca"] Campos & Lage 2013 Costa, Guedes & Bernils 2021   Ameivula ocellifera Brazilian Ocellated Whiptail Aparicio, Avila-Pires, Moravec & Nogueira 2019 Costa, Guedes & Bernils 2021 Mesquita & Colli 2003 Moreira, Fenolio, Silva & Silva 2009 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Kentropyx (Spurred Whiptails)   Kentropyx calcarata Striped Forest Whiptail Costa, Guedes & Bernils 2021   Kentropyx paulensis Sao Paulo Whiptail Costa, Guedes & Bernils 2021 Gallagher & Dixon 1992 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Kentropyx vanzoi Mato Grosso Whiptail Costa, Guedes & Bernils 2021 Ribeiro-Junior & Amaral 2016 Genus Salvator (Southern Tejus)   Salvator duseni Parana Teju Colli, Peres & Cunha 1998 [as rufescens] Costa, Guedes & Bernils 2021 Presch 1973 [as Tupinambis rufescens] Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Salvator merianae Southern Black-and-White Teju Campos & Lage 2013 Cintra, Silva & Silva 2009 Colli, Peres & Cunha 1998 Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ribeiro-Junior & Amaral 2016 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Tupinambis (Northern Tejus)   Tupinambis matipu Xingu Teju Costa, Guedes & Bernils 2021 Silva, Ribeiro-Junior & Avila-Pires 2018   Tupinambis quadrilineatus Four-striped Teju Abrahao-Morato, Meer, Bornschein, Vieira Capela, Almeida Ulandowski & Zampier 2015 Colli, Peres & Cunha 1998 [as cerradensis] Costa, Guedes & Bernils 2021 Freitas, Oliveira-Lima & Franca 2011 Koehler & Langerwerf 2000 Manzani & Abe 1997 Moreira, Fenolio, Silva & Silva 2009 Ribeiro-Junior & Amaral 2016 Silva, Ribeiro-Junior & Avila-Pires 2018 Silveira 2009   Tupinambis teguixin Amazonian Black-and-White Teju Costa, Guedes & Bernils 2021 Murphy, Jowers, Lehtinen, Charles, Colli, Peres, Hendry & Pyron 2016 Passos, Lima-Araujo, Melo & Borges-Nojosa 2013 Ribeiro-Junior & Amaral 2016 Silva, Ribeiro-Junior & Avila-Pires 2018  Family Tropiduridae (Rough-tailed Iguanas) Genus Stenocercus (Whorl-tailed Iguanas)   Stenocercus sinesaccus Mato Grosso Whorl-tailed Iguana Costa, Guedes & Bernils 2021 Morais, Signorelli, Vieira & Bastos 2010 Ribeiro-Junior 2015 Torres-Carvajal 2007 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Tropidurus (Eastern Keel-tailed Iguanas)   Tropidurus etheridgei Cordoba Keel-tailed Iguana Carvalho 2013 Costa, Guedes & Bernils 2021   Tropidurus itambere Sorocaba Keel-tailed Iguana Costa, Guedes & Bernils 2021 Rodrigues 1987 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Tropidurus montanus Minas Gerais Keel-tailed Iguana Costa, Guedes & Bernils 2021   Tropidurus oreadicus Cerrado Keel-tailed Iguana Avila-Pires 1995 Campos & Lage 2013 Carvalho 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ribeiro-Junior 2015 Rodrigues 1987 Santos, Sousa Abreu & Frota 2002   Tropidurus torquatus Collared Keel-tailed Iguana Campos & Lage 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Kunz & Borges-Martins 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Rodrigues 1987

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Aniliidae (South American Pipe Snake) Genus Anilius (South American Pipe Snake)   Anilius scytale South American Pipe Snake Costa, Guedes & Bernils 2021 Maschio, Prudente, Rodrigues & Hoogmoed 2010 Silva, Souza & Puorto 2001 Wallach, Williams & Boundy 2014  Family Anomalepididae (Neotropical Blind Snakes) Genus Liotyphlops (Smooth Blind Snakes)   Liotyphlops ternetzii Paraguayan Blind Snake Cintra, Silva & Silva 2009 [as beui] Costa, Guedes & Bernils 2021 [as beui] Cruz-Centeno, Swaya & Germano 2010 [as beui] Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009 [as beui]  Family Boidae (Typical Boas) Genus Boa (Red-tailed Boas)   Boa constrictor South American Red-tailed Boa Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Cunha, Moreira & Sousa-Silva 2010 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Corallus (Tree Boas)   Corallus hortulana Amazon Tree Boa Costa, Guedes & Bernils 2021 Genus Epicrates (Rainbow Boas)   Epicrates cenchria Common Rainbow Boa Andrade-Junior, Soares Franca, Sudre & Passos 2020 Campos & Lage 2013 Cintra, Silva & Silva 2009 Moreira, Fenolio, Silva & Silva 2009 Passos & Fernandes 2008   Epicrates crassus Paraguayan Rainbow Boa Andrade-Junior, Soares Franca, Sudre & Passos 2020 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Passos & Fernandes 2008 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Peters & Orejas-Miranda 1970 [as cenchria polylepis] Genus Eunectes (Anacondas)   Eunectes murinus Green Anaconda Campos & Lage 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Cunha, Moreira & Sousa-Silva 2010 Dirksen 2002 Franca & Silva Braz 2013 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007  Family Colubridae (Typical Snakes or Colubrids) Genus Adelphostigma (Twin-spotted Leaf-litter Snakes)   Adelphostigma occipitalis Bahia Leaf-litter Snake Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Apostolepis (Black-tailed Blackheads)   Apostolepis albicollaris White-collared Blackhead Costa, Guedes & Bernils 2021 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 [as A. cerradoensis] Entiauspe-Neto, Koch, Guedes, Paredero, Tiutenko & Loebmann 2022 Lema & Renner 2012 [as A. cerradoensis] Lema 2002 Lema 2003 [as A. cerradoensis] Nogueira, Barbo & Ferrarezzi 2012 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Wallach, Williams & Boundy 2014 [as A. cerradoensis]   Apostolepis assimilis Common Red Blackhead Costa, Guedes & Bernils 2021 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 Ferrarezzi, Barbo & Albuquerque 2005 Harvey 1999 Lema & Renner 2011 Lema & Renner 2012 Lema 2002 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Nogueira, Barbo & Ferrarezzi 2012 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Wallach, Williams & Boundy 2014   Apostolepis flavotorquata Yellow-collared Blackhead Costa, Guedes & Bernils 2021 Ferrarezzi, Barbo & Albuquerque 2005 Lema & Renner 2005 Lema & Renner 2012 Lema 2001 Lema 2002 Lima Silveira 2014 Nogueira, Barbo & Ferrarezzi 2012 Wallach, Williams & Boundy 2014   Apostolepis goiasensis Goias Blackhead Costa, Guedes & Bernils 2021 Lema & Renner 2012 Lema 2001 Loebmann & Lema 2012 Maffei & Campos Nogueira 2021 Nogueira, Barbo & Ferrarezzi 2012 Peters & Orejas-Miranda 1970 Wallach, Williams & Boundy 2014   Apostolepis intermedia Mato Grosso Blackhead Costa, Guedes & Bernils 2021   Apostolepis nelsonjorgei Minacu Blackhead Costa, Guedes & Bernils 2021 Curcio, Nunes, Harvey & Rodrigues 2011 Entiauspe-Neto, Koch, Guedes & Tiutenko 2020 Lema & Renner 2004 Lema & Renner 2012 Wallach, Williams & Boundy 2014   Apostolepis sanctaeritae Tocantins Blackhead Cintra, Silva & Silva 2009 [as ammodites] Costa, Guedes & Bernils 2021 Ferrarezzi, Barbo & Albuquerque 2005 [as ammodites] Franca & Silva Braz 2013 [as ammodites] Lema & Renner 2007 [as ammodites] Lema & Renner 2012 [as ammodites] Moreira, Fenolio, Silva & Silva 2009 [as ammodites] Nogueira, Barbo & Ferrarezzi 2012 [as ammodites] Wallach, Williams & Boundy 2014 [as ammodites] Genus Atractus (Spindle Snakes)   Atractus albuquerquei Guama River Spindle Snake Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009 Passos & Fernandes 2008 Passos, Azevedo, Nogueira, Fernandes & Sawaya 2019 Passos, Chiesse, Torres-Carvajal & Savage 2009 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Silva, Silva, Ribeiro, Souza & Souza 2005 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Wallach, Williams & Boundy 2014 Zaher, Souza, Gower, Hingst-Zaher & Silva 2005   * Atractus edioi Minacu Spindle Snake Costa, Guedes & Bernils 2021 Silva, Silva, Ribeiro, Souza & Souza 2005   Atractus pantostictus Sao Paulo Spindle Snake Costa, Guedes & Bernils 2021 Moreira, Fenolio, Silva & Silva 2009 Passos, Fernandes, Bernils & Moura-Leite 2010 Genus Boiruna (Southern Snake Eaters)   Boiruna maculata South American Spotted Snake Eater Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Giraudo & Scrocchi 2002 Zaher 1996 Genus Chironius (Sipos)   Chironius bicarinatus Eastern Sipo Costa, Guedes & Bernils 2021 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Chironius brazili Southeast Brazilian Sipo Costa, Guedes & Bernils 2021 Hamdan & Fernandes 2015   Chironius exoletus Green Sipo Costa, Guedes & Bernils 2021 Dixon, Wiest & Cei 1993 Franca & Silva Braz 2013 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Chironius flavolineatus Yellow-striped Sipo Albuquerque & Freitas 2011 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Dixon, Wiest & Cei 1993 Franca & Silva Braz 2013 Hamdan & Fernandes 2015 Hamdan, Scali & Fernandes 2014 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pinto, Fernandes & Marques 2008 Pinto, Marques & Fernandes 2010 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Chironius quadricarinatus Paraguayan Sipo Argolo 1998 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pinto, Fernandes & Marques 2008 Pinto, Marques & Fernandes 2010 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Clelia (Northern Snake Eaters)   Clelia plumbea Plumbeous Snake Eater Costa, Guedes & Bernils 2021 Moreira, Fenolio, Silva & Silva 2009 Genus Dipsas (Neotropical Snail Eaters)   Dipsas bucephala Bull Snail Eater Costa, Guedes & Bernils 2021   Dipsas mikanii Brazilian White-collared Snail Eater Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Dipsas ventrimaculata Spot-bellied Snail Eater Costa, Guedes & Bernils 2021 Genus Drymarchon (Indigo Snakes and Cribos)   Drymarchon corais Yellow-tailed Cribo Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Drymoluber (Glossy Racers)   Drymoluber brazili Southern Glossy Racer Caldeira-Costa, Ribeiro-Moura & Neves-Feio 2013 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Freitas, Franca & Verissimo 2012 Moreira, Fenolio, Silva & Silva 2009 Genus Dryophylax (Northern Mock Vipers)   Dryophylax hypoconia Southern Mock Viper Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Franco & Ferreira 2002 Trevine, Caicedo-Portilla, Hoogmoed, Thomas, Franco, Montingelli, Osorno-Munoz & Zaher 2021 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Dryophylax phoenix Petrolina Mock Viper Costa, Guedes & Bernils 2021 Franco, Trevine, Montingelli & Zaher 2017 Trevine, Caicedo-Portilla, Hoogmoed, Thomas, Franco, Montingelli, Osorno-Munoz & Zaher 2021 Genus Erythrolamprus (Neotropical Smooth Snakes)   Erythrolamprus aesculapii Southern Mock Coral Snake Costa, Guedes & Bernils 2021 Curcio, Scali & Rodrigues 2015 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Erythrolamprus almadensis Almada Smooth Snake Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009   Erythrolamprus frenatus Paraguayan Smooth Snake Costa, Guedes & Bernils 2021 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Erythrolamprus macrosomus Southern Royal Smooth Snake Ascenso, Costa & Prudente 2019 Costa, Guedes & Bernils 2021 Dixon 1989 Entiauspe-Neto, Abegg, Koch, Nunez, Azevedo, Moraes, Tiutenko, Bialves & Loebmann 2021 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Peters & Orejas-Miranda 1970 Valdujo & Nogueira 2001   Erythrolamprus maryellenae Goias Smooth Snake Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Dixon 1985 Dixon 1989 Franca & Silva Braz 2013 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Erythrolamprus poecilogyrus Yellow-bellied Smooth Snake Ascenso, Costa & Prudente 2019 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Peters & Orejas-Miranda 1970 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Erythrolamprus reginae Northern Royal Smooth Snake Ascenso, Costa & Prudente 2019 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Erythrolamprus typhlus Velvety Smooth Snake Costa, Guedes & Bernils 2021 Dixon 1987 Genus Helicops (South American Keelbacks)   Helicops angulatus Brown-banded Keelback Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Leao & Brandao 2012 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Roberto, Ribeiro, Sousa-Delfino & Oliveira-Almeida 2009 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Helicops gomesi Sao Paulo Keelback Costa, Guedes & Bernils 2021 Silveira, Prudente, Argolo, Abrahao, Nogueira, Strussmann, Loebmann, Barbo, Franco, Costa, Moura, Zaher, Borges-Martins, Martins, Oliveira, Hoogmoed, Marques, Passos, Bernils, Kawashita-Ribeiro, Sawaya & Costa 2019 Valdujo & Nogueira 2001   Helicops infrataeniatus Surinam Keelback Costa, Guedes & Bernils 2021 Moraes-da-Silva, Amaro, Sales Nunes, Rodrigues & Curcio 2021 Moraes-da-Silva, Amaro, Sales Nunes, Strussmann, Teixeira, Andrade, Sudre, Recoder, Rodrigues & Curcio 2019   Helicops modestus Brazilian Olive Keelback Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moraes-da-Silva, Amaro, Sales Nunes, Rodrigues & Curcio 2021 Moraes-da-Silva, Amaro, Sales Nunes, Strussmann, Teixeira, Andrade, Sudre, Recoder, Rodrigues & Curcio 2019 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Helicops polylepis South American Many-scaled Keelback Costa, Guedes & Bernils 2021   Helicops trivittatus Three-striped Keelback Costa, Guedes & Bernils 2021 Genus Hydrodynastes (False Water Cobras)   Hydrodynastes bicinctus Two-banded False Water Cobra Costa, Guedes & Bernils 2021 Franco, Fernandes & Bentim 2007 Silva & Franca e Souza 2002   Hydrodynastes gigas Giant False Water Cobra Costa, Guedes & Bernils 2021 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Imantodes (American Blunt-headed Tree Snakes)   Imantodes cenchoa Common Blunt-headed Tree Snake Campos Nogueira 2001 Costa, Guedes & Bernils 2021 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Leptodeira (Typical Cat-eyed Snakes)   Leptodeira annulata South American Banded Cat-eyed Snake Campos & Lage 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009   Leptodeira pulchriceps Bodoquena Banded Cat-eyed Snake Valdujo & Nogueira 2001 Genus Leptophis (Neotropical Parrot Snakes)   Leptophis ahaetulla Common Green Parrot Snake Albuquerque & Fernandes 2022 Albuquerque, Santos, Borges-Nojosa & Avila 2022 Costa, Guedes & Bernils 2021   Leptophis marginatus Paraguayan Parrot Snake Albuquerque & Fernandes 2022 Genus Lygophis (South American Striped Ground Snakes)   Lygophis dilepis Northern Striped Ground Snake Costa, Guedes & Bernils 2021   Lygophis meridionalis Paraguayan Striped Ground Snake Castro & Oliveira 2017 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Michaud & Dixon 1987 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Lygophis paucidens Cerrado Striped Ground Snake Costa, Guedes & Bernils 2021 Lema 1989 Michaud & Dixon 1987 Peters & Orejas-Miranda 1970 Serrano, Vieira-Alencar, Diaz-Ricaurte & Nogueira 2020 Valdujo & Nogueira 2001 Genus Mastigodryas (Neotropical Lizard Eaters)   Mastigodryas boddaerti Brown Lizard Eater Costa, Guedes & Bernils 2021 Montingelli 2009 Montingelli, Grazziotin, Battilana, Murphy, Zhang & Zaher 2019 Siqueira, Nascimento, Montingelli & Santos-Costa 2013 Souza & Silva 2001 Genus Mesotes (Southern Mock Vipers)   Mesotes rutilus Sao Paulo Mock Viper Costa, Guedes & Bernils 2021 Trevine, Caicedo-Portilla, Hoogmoed, Thomas, Franco, Montingelli, Osorno-Munoz & Zaher 2021 Genus Mussurana (Brazilian Snake Eaters)   Mussurana quimi South Brazilian Snake Eater Costa, Guedes & Bernils 2021 Entiauspe-Neto, Abegg, Quintela & Loebmann 2017 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Oxybelis (Neotropical Vine Snakes)   Oxybelis aeneus Brown Vine Snake Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Genus Oxyrhopus (Calico Snakes)   Oxyrhopus guibei Parana Calico Snake Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Zaher & Caramaschi 1992   Oxyrhopus petolarius Banded Calico Snake Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Oxyrhopus rhombifer Rhombic Calico Snake Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009 Valdujo & Nogueira 2001   Oxyrhopus trigeminus Northern Brazilian Calico Snake Campos & Lage 2013 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Zaher & Caramaschi 1992 Genus Palusophis (Eastern Lizard Eater)   Palusophis bifossatus Eastern Lizard Eater Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Marques & Muriel 2007 Montingelli 2009 Montingelli, Grazziotin, Battilana, Murphy, Zhang & Zaher 2019 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Phalotris (South American Collared Snakes)   Phalotris matogrossensis Mato Grosso Collared Snake Costa, Guedes & Bernils 2021 Lema, D'Agostini & Cappellari 2005   Phalotris mertensi Sao Paulo Reticulated Collared Snake Costa, Guedes & Bernils 2021 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Phalotris nasutus Snouted Collared Snake Costa, Guedes & Bernils 2021 Ferrarezzi 1993 Ferreira, Wang & Himmelstein 2005 Lema 2002 Valdujo & Nogueira 2001 Genus Philodryas (South American Bush Racers)   Philodryas aestiva Eastern Brazilian Green Bush Racer Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Philodryas livida Blue Bush Racer Costa, Guedes & Bernils 2021 Valdujo & Nogueira 1999   Philodryas mattogrossensis Mato Grosso Bush Racer Cacciali, Cabral, Ferreira & Koehler 2016 Costa, Guedes & Bernils 2021 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Philodryas nattereri Paraguay Bush Racer Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Smith, Scott, Cacciali, Atkinson & Pheasey 2013 Thomas 1976 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Philodryas olfersii Common Green Bush Racer Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Thomas 1976 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Philodryas psammophidea South American Striped Racer Costa, Guedes & Bernils 2021 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Genus Phimophis (South American Shovel-nosed Snakes)   Phimophis guerini Argentine Shovel-nosed Snake Costa, Guedes & Bernils 2021 Filho, Santana, Vieira, Alves, Montenegro & Freitas 2012 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Marques, Tinoco, Browne-Ribeiro & Fazolato 2012 Moreira, Fenolio, Silva & Silva 2009 Silva Lisboa, Araujo-Neto, Soares Tiburcio & Silva 2011 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Pseudablabes (Reticulated Bush Racers)   Pseudablabes agassizii Burrowing Night Snake Campos Nogueira 2001 Carvalho-Junior, Schettino Canelas & Cordeiro-Malta 2006 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Souza Filho & Plombon 2014   Pseudablabes patagoniensis Patagonian Bush Racer Campos Nogueira 2001 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Gouveia, Novelli, Vieira & Sousa 2017 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Pseudoboa (Neotropical Scarlet Snakes)   Pseudoboa coronata Black-headed Scarlet Snake Caldeira Costa, Cotta & MacCulloch 2015 Costa, Guedes & Bernils 2021   Pseudoboa nigra Blackish Scarlet Snake Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009 Orofino, Pizzatto & Marques 2010 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Zaher, Oliveira & Franco 2008 Genus Psomophis (Smooth Ground Snakes)   Psomophis joberti Brazilian Ground Snake Costa, Guedes & Bernils 2021 Moreira, Fenolio, Silva & Silva 2009 Myers & Cadle 1994 Genus Rhachidelus (Brazilian Bird Snake)   Rhachidelus brazili Brazilian Bird Snake Arruda, Avila, Morais & Kawashito-Ribeiro 2011 Costa, Guedes & Bernils 2021 Fernandes & Passos 2002 Lima Silveira 2004 Moreira, Fenolio, Silva & Silva 2009 Genus Rodriguesophis (Collared Sharp-nosed Snakes)   Rodriguesophis iglesiasi Piaui Sharp-nosed Snake Costa, Guedes & Bernils 2021 Genus Simophis (South American False Coral Snake)   Simophis rhinostomus South American False Coral Snake Cacciali, Scott, Gunther, Sawaya, Brusquetti & Bauer 2009 Costa, Guedes & Bernils 2021 Genus Siphlophis (Liana Snakes)   Siphlophis leucocephalus White-headed Liana Snake Costa, Guedes & Bernils 2021 Prudente, Silva, Santos Meireles & Puorto 2017   Siphlophis longicaudatus Long-tailed Liana Snake Costa, Guedes & Bernils 2021 Prudente, Silva, Santos Meireles & Puorto 2017 Genus Spilotes (Chicken Snakes)   Spilotes pullatus Chicken Snake Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Cunha, Moreira & Sousa-Silva 2010 Franca & Silva Braz 2013 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Tantilla (American Centipede Snakes)   Tantilla melanocephala South American Black-headed Centipede Snake Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Sawaya & Sazima 2003 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Xenodon (Northern Mock Lanceheads and South American Hognose Snakes)   Xenodon merremii Toad-eating Mock Lancehead Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Xenodon nattereri Banded Hognose Snake Argolo 2002 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lima Silveira, Ribeiro Costa & Salles 2004   Xenodon severus Giant Mock Lancehead Costa, Guedes & Bernils 2021 Genus Xenopholis (South American Flat-headed Snakes)   Xenopholis undulatus Southern Flat-headed Snake Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Loebmann 2009 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Micrurus (American Coral Snakes)   Micrurus brasiliensis Brazilian Coral Snake Campbell & Lamar 2004 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Pires, Silva, Feitosa, Prudente, Filho & Zaher 2014 Silva & Sites 1999   Micrurus carvalhoi Brazilian Ribbon Coral Snake Castro, Oliveira, Gonzalez, Curcio & Feitosa 2017 Costa, Guedes & Bernils 2021 Pires, Silva, Feitosa, Prudente, Filho & Zaher 2014 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 [as M. lemniscatus]   Micrurus frontalis Cerrado Coral Snake Campbell & Lamar 2004 Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Franca & Silva Braz 2013 Moreira, Fenolio, Silva & Silva 2009 Silva & Sites 1999   Micrurus surinamensis Amazonian Aquatic Coral Snake Costa, Guedes & Bernils 2021 Silva, Rodrigues Silva & Pessoa 2008  Family Leptotyphlopidae (Thread Snakes) Genus Epictia (Neotropical Thread Snakes)   Epictia clinorostris Flat-nosed Thread Snake Arredondo & Zaher 2010 Costa, Guedes & Bernils 2021 Francisco, Pinto & Fernandes 2012 Genus Trilepida (Three-scaled Thread Snakes)   Trilepida brasiliensis Bahia Thread Snake Costa, Guedes & Bernils 2021 Pinto & Curcio 2011   Trilepida fuliginosa Preto River Thread Snake Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Guedes, Queiroz, Sousa & Hamdan 2020 Martins, Koch, Pinto, Folly, Fouquet & Passos 2019 Moreira, Fenolio, Silva & Silva 2009 Passos, Caramaschi & Pinto 2006 Pinto & Fernandes 2012   Trilepida koppesi Terenos Thread Snake Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Filogonio & Canelas 2015 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Martins, Koch, Pinto, Folly, Fouquet & Passos 2019 Passos, Caramaschi & Pinto 2006 Pinto & Fernandes 2012 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Valdujo & Nogueira 2001 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007  Family Typhlopidae (Typical Blind Snakes) Genus Amerotyphlops (American Blind Snakes)   Amerotyphlops brongersmianus Bahia Blind Snake Campos & Lage 2013 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Dixon & Hendricks 1979 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007  Family Viperidae (Vipers) Genus Bothrops (American Lanceheads)   Bothrops alternatus Urutu Campbell & Lamar 2004 Costa, Guedes & Bernils 2021 Nogueira & Valdujo 2001   Bothrops itapetiningae Itapetininga Lancehead Campbell & Lamar 2004 Costa, Guedes & Bernils 2021   Bothrops jararaca Jararaca Costa, Guedes & Bernils 2021   Bothrops lutzi Cerrado Lancehead Campbell & Lamar 2004 Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Loebmann 2009 Silva & Rodrigues 2008   Bothrops marmoratus Marbled Lancehead Carrasco, Grazziotin, Farfan, Koch, Ochoa, Scrocchi, Leynaud & Chaparro 2019 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Silva & Rodrigues 2008   Bothrops mattogrossensis Mato Grosso Lancehead Campbell & Lamar 2004 Costa, Guedes & Bernils 2021 Silva & Rodrigues 2008   Bothrops moojeni Brazilian Lancehead Campbell & Lamar 2004 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Moreira, Fenolio, Silva & Silva 2009 Nogueira, Sawaya & Martins 2003 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007   Bothrops neuwiedi Eastern Brazilian Lancehead Campbell & Lamar 2004 Cintra, Silva & Silva 2009 Costa, Guedes & Bernils 2021 Hoge & Romano Hoge 1981 Moreira, Fenolio, Silva & Silva 2009 Silva & Rodrigues 2008   Bothrops pauloensis Black-faced Lancehead Campbell & Lamar 2004 Costa, Guedes & Bernils 2021 David, Dobiey & Vogel 2018 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Ramalho, Silva, Soares, Ferraz, Arruda & Prado 2018 Silva & Rodrigues 2008 Valdujo, Nogueira & Martins 2002 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Crotalus (Typical Rattlesnakes)   Crotalus durissus South American Rattlesnake Campbell & Lamar 2004 Campos & Lage 2013 Costa, Guedes & Bernils 2021 Franca & Silva Braz 2013 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Lachesis (Bushmasters)   Lachesis muta South American Bushmaster Costa, Guedes & Bernils 2021 Fernandes, Franco & Fernandes 2004

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Chelidae (Austro-American Side-necked Turtles) Genus Acanthochelys (South American Side-necked Swamp Turtles)   Acanthochelys spixii Black Spiny-necked Swamp Turtle Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chelus (Matamatas)   Chelus fimbriata Southern Matamata Costa, Guedes & Bernils 2021 Lima Silveira 2011 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Vargas-Ramirez, Caballero, Morales-Betancourt, Lasso, Amaya, Martinez, Silva Viana, Vogt, Pires Farias, Hrbek, Campbell & Fritz 2020 Genus Mesoclemmys (Hump-backed Toad-headed Turtles)   Mesoclemmys perplexa Piaui Toad-headed Turtle Campos & Lage 2013 Campos, Moraes & Pereira 2011 Costa, Guedes & Bernils 2021   Mesoclemmys vanderhaegei Tobati Toad-headed Turtle Brito, Strussmann, Kawashita-Ribeiro, Morais, Avila & Campos 2012 Costa, Guedes & Bernils 2021 Maffei, Nascimento, Moya & Donatelli 2016 Marques, Boehm, Brito, Cabrera & Verdade 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Souza 2004 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007 Genus Phrynops (Lesser Toad-headed Turtles)   Phrynops geoffroanus Brazilian Toad-headed Turtle Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Kinosternidae (American Mud and Musk Turtles) Genus Kinosternon (American Mud Turtles)   Kinosternon scorpioides Scorpion Mud Turtle Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Podocnemididae (Giant Side-necked Turtles) Genus Podocnemis (South American Side-necked River Turtles)   Podocnemis expansa Giant South American River Turtle Alves-Junior, Lustosa, Bosso, Balestra, Bastos, Miranda & Santos 2012 Costa, Guedes & Bernils 2021 Ferreira-Junior & Amorim Castro 2006 Ferreira-Junior & Castro 2005 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Podocnemis unifilis Yellow-spotted Amazon River Turtle Costa, Guedes & Bernils 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Chelonoidis (South American Tortoises)   Chelonoidis carbonarius South American Red-footed Tortoise Costa, Guedes & Bernils 2021 Lopes Santos, Pereira de Andrade, Victor-Junior & Vaz-Silva 2014 Pereira Ramalho, Fagundes Franca, Guerra, Marciano, Vale & Rodrigues Silva 2018 Vaz-Silva, Guimaraes-Guedes, Azevedo-Silva, Gontijo, Barbosa, Aloisio & Oliveira 2007