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Taxonomy of the family Lacertidae
Bibliography of the genus Eremias
Biodiversity of the family Lacertidae








Genus
Eremias

Desert and Steppe Runners, Eurasian Sand Lizards, Old World Racerunners, Desert Lacertids

Wüstenrenner, Buckelnasen

Ørkenløbere, Ørkenfirben

1834 Eremias Wiegmann (type species: Lacerta velox Pallas 1771; syn. ICZN (1985): Opinion 1318) (but see Jeremchenko 1999: type species: Lacerta variabilis
1834 Scapteira Wiegmann (type species: Lacerta grammica Lichtenstein 1823; syn. Arnold 1989)
1841 Aspidorhinus Eichwald (type species: Aspidorhinus gracilis Eichwald 1841; syn. Boulenger 1921)
1843 Eremioscopus Fitzinger (type species: Lacerta guttulata Lichtenstein 1823; syn. Arnold 1989)
1918 Macmahonia Boulenger (type species: Scaptira aporosceles Alcock & Finn 1896; syn. Smith 1935)
1928 Ommateremias Lantz (type species: Lacerta arguta Pallas 1773; syn. Arnold 1989)
1928 Rhabderemias Lantz (type species: Podarces scripta Strauch 1867; syn. Arnold 1989)
1973 Pareremias Szczerbak (type species: Eremias multiocellata Günther 1872; syn. Arnold 1989)
Contents: 41 species, of which 17 (41.5%) are endemic.
Remarks: Some of the nominate genera or subgenera listed in the synonymy above have been treated as separate genera (e.g., Welch 1983). Previously also included the genera Heliobolus, Mesalina, Pedioplanis, and Pseuderemias (e.g., Boulenger 1921).
Distribution: E. Europe, Middle East, C. and E. Asia.
Reported from: Afghanistan, Armenia, Azerbaijan, China (Anhui, Beijing, Gansu, Hebei, Heilongjiang, Henan, Inner Mongolia, Jiangsu, Jilin, Liaoning, Ningxia, Qinghai, Shandong, Shaanxi, Shanxi, Tibet, Xinjiang), Georgia, Iran, Kazakhstan, Kyrgyzstan, Moldova, Mongolia, North Korea, Pakistan, Romania, Russia, South Korea, Tajikistan, Turkey, Turkmenistan, Ukraine (incl. Crimea), Uzbekistan.

Eremias acutirostris

Lesser Reticulated Desert Runner, Point-snouted Desert Runner

Kleiner Netz-Wüstenrenner

Lille Netørkenløber

1887 Scapteira acutirostris Boulenger
Eremias acutirostris Lantz 1928

Remarks: Previously included aporosceles.
Distribution: Afghanistan, Iran, Pakistan.


Eremias afghanistanica

Afghan Desert Runner

Afghanischer Wüstenrenner

Afghansk Ørkenløber

1991 Eremias afghanistanica Böhme & Szczerbak

Distribution: Afghanistan.


Eremias andersoni

Striped Kavir Desert Runner

Gestreifter Kavir-Wüstenrenner

Stribet Kavir-ørkenløber

1978 Eremias andersoni Darevsky & Shcherbak
Rhabderemias andersoni Welch 1983

Distribution: Iran.


Eremias aporosceles

Greater Reticulated Desert Runner

Großer Netz-Wüstenrenner

Stor Netørkenløber

1896 Scapteira aporosceles Alcock & Finn
Macmahonia aporosceles Boulenger 1921
Eremias aporosceles Smith 1935

Remarks: Previously regarded as a synonym of acutirostris (e.g., Szczerbak 1974).
Distribution: Pakistan.


Eremias argus

Mongolian Desert Runner

Mongolischer Wüstenrenner

Mongolsk Ørkenløber

1869 Eremias argus Peters
Podarces argus Strauch 1876
1925 Eremias barbouri Schmidt (Szczerbak 1974)

Distribution: China (Anhui, Beijing, Gansu, Hebei, Heilongjiang, Henan, Inner Mongolia, Jiangsu, Jilin, Liaoning, Ningxia, Qinghai, Shandong, Shaanxi, Shanxi), Mongolia, North Korea, Russia, South Korea.


Eremias arguta

Kazakhstan Steppe Runner, Motley Lizard, (Steppe Runner)

Kasachischer Wüstenrenner

Kazakhstansk Steppeløber

1773 Lacerta arguta Pallas
Eremias arguta Gray 1845
Podarces arguta Strauch 1876
Ommateremias arguta Welch 1983
1789 Lacerta deserti Gmelin (Szczerbak 1974)
Ommateremias arguta deserti Welch 1983
Eremias arguta deserti Lantz 1922
1814 Lacerta variabilis Pallas (Szczerbak 1974)
Podarcis variabilis Ménétries 1832
Eremias variabilis Wiegmann 1834
1823 Lacerta leucosticta Lichtenstein (Szczerbak 1974)
1831 Lacerta pustulata Eichwald (Szczerbak 1974)
1832 Lacerta boristhenis Dwigubskij (Szczerbak 1974)
1832 Podarcis irritans Menetries (Szczerbak 1974)
1912 Eremias arguta potanini Bedriaga (Szczerbak 1974)
1923 Eremias arguta occidentalis Terentjev
1934 Eremias arguta uzbekistanica Chernov
1953 Eremias arguta transcaucasica Darevsky
Ommateremias arguta transcaucasica Welch 1983
1986 Eremias arguta darevskii Tsaruk

Distribution: Armenia, Azerbaijan, China (Gansu, Inner Mongolia, Xinjiang), Georgia, Iran, Kazakhstan, Kyrgyzstan, Moldova, Mongolia, Romania, Russia, Tajikistan, Turkmenistan, Ukraine (incl. Crimea), Uzbekistan.


Eremias aria

Jalalabad Desert Runner

Jalalabad-Wüstenrenner

Jalalabad-ørkenløber

1967 Eremias aria Anderson & Leviton
Ommateremias aria Welch 1983

Distribution: Afghanistan.


Eremias brenchleyi

Ordos Desert Runner

Ordos-Wüstenrenner

Ordos-ørkenløber

1872 Eremias brenchleyi Günther
Podarces brenchleyi Strauch 1876
Eremias argus brenchleyi Boettger 1918

Remarks: Records from Russia and Mongolia are regarded as erroneous (Zhao & Adler 1993).
Distribution: China (Anhui, Beijing, Hebei, Henan, Inner Mongolia, Jiangsu, Shandong, Shanxi).


Eremias buechneri

Kaschar Desert Runner

Kaschar-Wüstenrenner

Kaschar-ørkenløber

1906 Eremias buechneri Bedriaga 1905 [1906]

Remarks: Records from Kyrgyzstan refer to kokshaaliensis (Sindaco & Jeremchenko 2008). Not mentioned for China by Dai, Chen, Zhang & Wang (2006).
Distribution: China (Xinjiang).


Eremias cholistanica

Cholistan Desert Runner

Cholistan-Wüstenrenner

Cholistan-ørkenløber

2006 Eremias cholistanica Baig & Masroor

Distribution: Pakistan.


Eremias dzungarica

Dzungarian Desert Runner

Dsungarischer Wüstenrenner

Dzungarisk Ørkenløber

2017 Eremias dzungarica Orlova, Polyarkov, Chirikova, Nazarov, Munkhbaatar, Munkhbayar & Terbish

Distribution: Kazakhstan, Mongolia.


Eremias fahimii

Markazi Desert Runner, (Fahimi's Racerunner)

Markazi-Wüstenrenner

Markazi-ørkenløber

2020 Eremias fahimii Mozaffari, Ahmadzadeh & Pirooz

Distribution: Iran.





Eremias fasciata

Kerman Striped Desert Runner

Kerman-Wüstenrenner

Kerman-ørkenløber

1874 Eremias fasciata Blanford
Podarces fasciata Strauch 1876
Rhabderemias fasciata Welch 1983

Remarks: Old records from Turkmenistan have not been confirmed recently (Sindaco & Jeremchenko 2008).
Distribution: Afghanistan, Iran, Pakistan.


Eremias grammica

Karakumy Reticulated Desert Runner, (Reticulated Desert Runner)

Karakumy-Netz-Wüstenrenner

Karakumy-ørkenløber

1823 Lacerta grammica Lichtenstein
Podarcis grammica Wagler 1830
Scapteira grammica Wiegmann 1834
Eremias grammica Lantz 1928
1928 Eremias zarudnyi Lantz [substitute name for Scapteira persica Nikolski 1900]

Distribution: Afghanistan, China (Xinjiang), Iran, Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan.


Eremias intermedia

Aral-Caspian Desert Runner

Aral-Kaspischer Wüstenrenner

Aral-ørkenløber

1876 Podarces intermedia Strauch
Eremias intermedius Bogdanov 1882
1905 Eremias transcaspica Nikolski [not Scapteira transcaspica Nikolski 1905] (Szczerbak 1974)
1912 Eremias intermedia oxyrrhina Bedriaga (Szczerbak 1974)

Remarks: See under nigrocellata.
Distribution: Afghanistan, Iran, Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan.


Eremias isfahanica

Isfahan Desert Runner

Isfahan-Wüstenrenner

Isfahan-ørkenløber

2016 Eremias isfahanica Rastegar-Pouyani, Hosseinian, Rafiee, Kami, Rajabizadeh & Wink

Distribution: Iran.


Eremias kakari

Balochistan Desert Runner

Belutschistan-Wüstenrenner

Balochistan-ørkenløber

2020 Eremias kakari Masroor, Khisroon, Khan & Jablonski

Distribution: Pakistan.





Eremias kavirensis

Barred Kavir Desert Runner

Gebänderter Kavir-Wüstenrenner

Båndet Kavir-ørkenløber

2007 Eremias kavirensis Mozaffari & Parham

Distribution: Iran.


Eremias kokshaaliensis

Kokshaal Desert Runner

Kokshaal-Wüstenrenner

Kokshaal-ørkenløber

1999 Eremias kokshaaliensis Eremchenko & Panfilov

Remarks: This species was previously regarded as populations of buechneri (Sindaco & Jeremchenko 2008). Not mentioned for China by Dai, Chen, Zhang & Wang (2006).
Distribution: China (Xinjiang), Kyrgyzstan.


Eremias kopetdaghica

Kopet Dagh Desert Runner

Kopet-Dagh-Wüstenrenner

Kopet Dagh-ørkenløber

1972 Eremias strauchi kopetdaghica Shcherbak
Eremias kopetdaghica Smid, Moravec, Kodym, Kratochvil, Yousefkhani, Rastegar-Pouyani & Frynta 2014

Distribution: Iran, Turkmenistan.


Eremias lalezharica

Mount Lalezhar Desert Runner

Lalezhar-Wüstenrenner

Lalezhar-ørkenløber

1994 Eremias lalezharica Moravec

Distribution: Iran.


Eremias lineolata

Khorasan Striped Desert Runner

Khorasan-Wüstenrenner

Khorasan-ørkenløber

1896 Scapteira lineolata Nikolsky
Eremias lineolata Lantz 1928
Rhabderemias lineolata Welch 1983
1905 Scapteira grum-grzimailoi Bedriaga (Szczerbak 1974)

Distribution: Afghanistan, Iran, Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan.


Eremias montana

Kermanshah Desert Runner

Kermanshah-Wüstenrenner

Kermanshah-ørkenløber

2001 Eremias montanus Rastegar-Pouyani & Rastegar-Pouyani
2008 Eremias novo Rastegar-Pouyani & Rastegar-Pouyani (Bafti, Sharifi & Papenfuss 2009)
Eremias nova [justified emendation] Sindaco, Venchi & Grieco 2013

Remarks: There appears to be some confusion concerning the name Eremias novo. This was a temporary name for some populations subsequently identified as Eremias montana inhabiting the alpine meadows of Alvand Mountains in Hamedan Province (Bafti, Sharifi & Papenfuss 2009, citing pers. comm. with N. Rastegar-Pouyani, September 2008). Nevertheless, Eremias novo has subsequently been listed as a valid species by some authors (e.g.. Sindaco, Venchi & Grieco 2013; Smid, Moravec, Kodym, Kratochvil, Yousefkhani, Rastegar-Pouyani & Frynta 2014).
Distribution: Iran.


Eremias multiocellata

Multiocellated Desert Runner

Kleinfleckiger Wüstenrenner

Småplettet Ørkenløber

1872 Eremias multiocellata Günther
Podarces multiocellata Strauch 1876
1876 Podarces planiceps Strauch (Szczerbak 1974)
1906 Eremias multiocellata koslowi Bedriaga
1912 Eremias multiocellata reticulata Bedriaga (Szczerbak 1974)
1973 Eremias multiocellata bannikowi Shcherbak
2010 Eremias multiocellata tsaganbogdensis Munkhbayar & Borkin

Remarks: Previously included stummeri and yarkandensis. Some older records of multiocellata may represent confusions with stummeri or yarkandensis. Records from Kyrgyzstan and Uzbekistan are herein tentatively assigned to stummeri, although Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov (2006) reported multiocellata (but also stummeri and yarkandensis) from Kyrgyzstan. However, according to Sindaco & Jeremchenko (2008), the westernmost record of multiocellata is in far eastern Kazakhstan.
Distribution: China (Gansu, Inner Mongolia, Liaoning, Ningxia, Qinghai, Shaanxi, Tibet, Xinjiang), Kazakhstan, Mongolia, Russia.


Eremias nigrocellata

Black-ocellated Desert Runner

Schwarzfleckiger Wüstenrenner

Sortplettet Ørkenløber

1896 Eremias nigrocellata Nikolsky
Eremias intermedia nigrocellata Nikolsky 1899
Ommateremias nigrocellata Welch 1983

Distribution: Afghanistan, Iran, Tajikistan, Turkmenistan, Uzbekistan.


Eremias nikolskii

Kyrgyz Desert Runner

Kirgisischer Wüstenrenner

Kirgisisk Ørkenløber

1905 Eremias nikolskii Bedriaga

Distribution: Kyrgyzstan, Tajikistan, Uzbekistan.


Eremias papenfussi

Alborz Mountains Desert Runner

Alborz-Wüstenrenner

Alborz-ørkenløber

2011 Eremias papenfussi Mozaffari, Ahmadzadeh & Parham

Distribution: Iran.


Eremias persica

Persian Desert Runner

Persischer Wüstenrenner

Persisk Ørkenløber

1874 Eremias persica Blanford
Podarces persica Strauch 1876
Eremias velox persica Boettger 1886
1998 Eremias nigrolateralis Rastegar-Pouyani & Nilson (Rastegar-Pouyani, Rastegar-Pouyani, Noureini, Joger & Wink 2010)

Other common names:
nigrolateralis: Black-sided Desert Runner
Remarks: Presence in Iraq needs confirmation (Leviton, Anderson, Adler & Minton 1992), although reported by Khalaf (1959). Safaei-Mahroo & al. (2015) listed nigrolateralis as a valid species, but without comments.
Distribution: Afghanistan, Azerbaijan, Iran, Pakistan, Turkmenistan.


Eremias pleskei

Transcaucasian Desert Runner

Transkaukasischer Wüstenrenner

Transkaukasisk Ørkenløber

1907 Eremias pleskei Bedriaga
Eremias fasciata pleskei Nikolsky 1913
Rhabderemias pleskei Welch 1983

Distribution: Armenia, Azerbaijan, Iran, Turkey.


Eremias przewalskii

Gobi Desert Runner

Gobi-Wüstenrenner

Gobi-ørkenløber

1876 Podarces przewalskii Strauch
Eremias przewalskii Boulenger 1887
1876 Podarces brachydactyla Strauch (Szczerbak 1974)
1876 Podarces kessleri Strauch (Szczerbak 1974)
1970 Eremias przewalskii tuvensis Shcherbak

Remarks: Dai, Chen, Zhang & Wang (2006) doubted the validity of przewalskii and suggested that it could be a synonym of multiocellata.
Distribution: China (Gansu, Inner Mongolia, Ningxia, Qinghai, Xinjiang), Mongolia, Russia.


Eremias quadrifrons

Alashan Desert Runner

Alashan-Wüstenrenner

Alashan-ørkenløber

1876 Podarces quadrifrons Strauch
Eremias quadrifrons Boulenger 1887

Remarks: Dai, Chen, Zhang & Wang (2006) did not consider quadrifrons a valid species.
Distribution: China (Inner Mongolia).


Eremias regeli

Tajik Desert Runner

Tadschikischer Wüstenrenner

Tadsjikisk Ørkenløber

1905 Eremias regeli Bedriaga
1911 Eremias bedriagai Nikolski (Szczerbak 1974)

Distribution: Afghanistan, Tajikistan, Turkmenistan, Uzbekistan.


Eremias roborowskii

Turpan Steppe Runner

Turpan-Wüstenrenner

Turpan-ørkenløber

1907 Eremias velox var. roborowskii Bedriaga
Eremias roborowskii Liu, Guo, Chen, Li, Yue & Zeng 2018

Remarks: Previously regarded as a subspecies of velox. Raised to species status by Liu, Guo, Chen, Li, Yue & Zeng (2018).
Distribution: China (Xinjiang).





Eremias scripta

Caspian Desert Runner, Sand Racerunner, (Vermiculate Sand Lacerta)

Kaspischer Wüstenrenner

Kaspisk Ørkenløber

1867 Podarces scripta Strauch
Scapteira scripta Severzow 1873
Eremias scripta Lantz 1928
Rhabderemias scripta Welch 1983
1905 Scapteira bilkewitschi Nikolski (Szczerbak 1974)
1918 Scapteira lasdini Tzarewsky (Szczerbak 1974)
Eremias scripta lasdini Vedmederya, Zinenko & Barabanov 2009
1973 Eremias scripta pherganensis Shcherbak & Vashetko

Distribution: Afghanistan, Kazakhstan, Pakistan, Tajikistan, Turkmenistan, Uzbekistan.


Eremias strauchi

Armenian Desert Runner, (Strauch's Desert Runner)

Strauchs Wüstenrenner

Armensk Ørkenløber

1878 Eremias strauchi Kessler
Eremias velox strauchi Nikolsky 1913

Remarks: Previously included kopetdaghica.
Distribution: Armenia, Azerbaijan, Iran, Turkey.


Eremias stummeri

Tien Shan Desert Runner

Tien Shan-Wüstenrenner

Tien Shan-ørkenløber

1940 Eremias velox stummeri Wettstein
Eremias multiocellata stummeri Bischoff 1993
Eremias stummeri Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006

Remarks: Regarded as a synonym of multiocellata by Szczerbak (1974, 2003) and the Reptile Database (viewed 16 April 2013), but as a valid species by others (e.g., Sindaco & Jeremschenko 2008; Dujsebayeva 2010). Validity confirmed by Orlova, Polyarkov, Chirikova, Nazarov, Munkhbaatar, Munkhbayar & Terbish (2017). Not mentioned for China by Dai, Chen, Zhang & Wang (2006).
Distribution: China (Xinjiang), Kazakhstan, Kyrgyzstan, Uzbekistan.


Eremias suphani

Anatolian Desert Runner

Anatolischer Wüstenrenner

Anatolsk Ørkenløber

1968 Eremias velox suphani Basoglu & Hellmich
Eremias suphani Bischoff & Böhme 1980

Distribution: Iran, Turkey.


Eremias szczerbaki

Arpa Valley Desert Runner, (Szczerbak's Desert Runner)

Arpa-Wüstenrenner

Arpa-ørkenløber

1992 Eremias szczerbaki Eremchenko & Panfilov
Eremias multiocellata szczerbaki Bischoff 1993

Remarks: Revalidated as a separate species by Orlova, Polyarkov, Chirikova, Nazarov, Munkhbaatar, Munkhbayar & Terbish (2017).
Distribution: Kyrgyzstan.


Eremias velox

Rapid Steppe Runner, Rapid Racerunner

Schneller Wüstenrenner

Hurtig Ørkenløber

1771 Lacerta velox Pallas
Podarcis velox Wagler 1830
Eremias velox Wiegmann 1834
1771 Lacerta cruenta Pallas (Szczerbak 1974)
1802 Scincus cruentatus Daudin [substitute name for Lacerta cruenta Pallas 1771] (Szczerbak 1974)
1820 Lacerta coccinea Merrem [substitute name for Lacerta cruenta Pallas 1771] (Szczerbak 1974)
1820 Lacerta tigrina Kuhl (Szczerbak 1974)
1831 Lacerta argulus Eichwald (Szczerbak 1974)
1831 Lacerta gracilis Eichwald (Szczerbak 1974)
1834 Lacerta vittata Eversmann (Szczerbak 1974)
1873 Eremias erythrurus Sewerzow (Szczerbak 1974)
1928 Eremias velox caucasia Lantz
1999 Eremias velox borkini Jeremchenko & Panfilov

Remarks: Previously included persica, roborowskii, strauchi, and suphani.
Distribution: Afghanistan, Azerbaijan, China (Gansu, Inner Mongolia, Xinjiang), Georgia, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Russia, Tajikistan, Turkmenistan, Uzbekistan.


Eremias vermiculata

Central Asian Desert Runner

Zentralasiatischer Wüstenrenner

Centralasiatisk Ørkenløber

1875 Eremias vermiculata Blanford
1876 Podarces pylzowi Strauch (Szczerbak 1974)

Distribution: China (Gansu, Inner Mongolia, Ningxia, Xinjiang), Kazakhstan, Mongolia.


Eremias yarkandensis

Yarkand Desert Runner

Yarkand-Wüstenrenner

Yarkand-ørkenløber

1875 Eremias yarkandensis Blanford
Eremias multiocellata yarkandensis Boulenger 1921
1875 Eremias yarkandensis saturata Blanford (Szczerbak 1974)

Remarks: Revalidated as a separate species by Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov (2006), supported by others (e.g., Dai, Chen, Zhang & Wang 2006; Orlova, Polyarkov, Chirikova, Nazarov, Munkhbaatar, Munkhbayar & Terbish 2017). Restricted to western China and Kyrgyzstan (Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006; Sindaco & Jeremchenko 2008). Following these authors, Zhao & Adler's (1993) report from Kazakhstan is herein tentatively referred to stummeri.
Distribution: China (Xinjiang), Kyrgyzstan.