Reptiles of
Georgia 

 

Bibliography of the reptiles of Georgia Taxonomic history of the reptiles of Georgia Introduced species
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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist
General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial.
Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:
Lizards (Sauria)
= endemicEX = extinct or assumed extinct

Species
References
 Family Agamidae (Agamas and Dragons)
Genus Laudakia (Asian Rock Agamas)
 
Laudakia caucasia

Caucasian Rock Agama

Ananjeva & Tuniev 1994 Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bosch & Bischoff 2004 Orlowa in Boehme (ed.) 1981 Sindaco & Jeremchenko 2008 Szczerbak 2003 Yousefkhani, Ficetola, Rastegar-Pouyani, Ananjeva, Rastegar-Pouyani & Masroor 2013
 Family Anguidae (Slowworms, etc.)
Genus Anguis (Slowworms)
 
Anguis colchica

Eastern Slowworm

Bakradze & Chkhikvadze 1993 [as A. fragilis] Bischoff 1978 [as A. fragilis] Bosch & Bischoff 2004 Dely in Boehme (ed.) 1981 [as A. fragilis] Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010
Genus Pseudopus (Eurasian Glass Lizard)
 
Pseudopus apodus

Eurasian Glass Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bosch & Bischoff 2004 Obst in Boehme (ed.) 1981
 Family Gekkonidae (Typical Geckos)
Genus Tenuidactylus (Central Asian Thin-toed Geckos)
 
Tenuidactylus caspius

Caspian Thin-toed Gecko

Bakradze & Chkhikvadze 1993 Bischoff 1978 Bosch & Bischoff 2004 Szczerbak & Golubev 1996
 Family Lacertidae (Typical Lizards)
Genus Darevskia (Caucasian Rock Lizards)
 
Darevskia adjarica

Northern Red-bellied Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arribas, Ilgaz & Kumlutas 2018 Bakradze & Chkhikvadze 1993 Bischoff 1978 [as parvula] Bosch & Bischoff 2004 Darevsky & Eiselt 1980 Fu, Murphy & Darevsky 1997 [as parvula] Sindaco & Jeremchenko 2008 Szczerbak 2003 Tuniyev, Ananjeva, Agasyan, Orlov & Tuniyev 2009 Uzzell & Darevsky 1975 [as parvula]
 
Darevskia alpina

Alpine Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008
 
Darevskia armeniaca

Armenian Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975
 
Darevskia brauneri

Western Caucasian Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2001 Bosch & Bischoff 2004 Darevsky in Boehme (ed.) 1984 Doronin, Doronina & Bekoshvili 2019 Sindaco & Jeremchenko 2008 Szczerbak 2003
 
Darevskia caucasica

Common Caucasian Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2001 Bischoff 2003 Bosch & Bischoff 2004 Darevsky in Boehme (ed.) 1984 Sindaco & Jeremchenko 2008
 
Darevskia clarkorum

Borcka Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Darevsky & Tuniyev 1997 Sindaco & Jeremchenko 2008
 
Darevskia daghestanica

Dagestan Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008
 
Darevskia dahli

Shagali Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2001 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975
 
Darevskia derjugini

Artvin Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Fu, Murphy & Darevsky 1997 Orlowa & Bischoff in Boehme (ed.) 1984 Sindaco & Jeremchenko 2008 Szczerbak 2003 Terentjev & Chernov 1965 [1949]
  *
Darevskia dryada

Charnali Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Darevsky & Tuniyev 1997
  *
Darevskia mixta

Adjar Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Fu, Murphy & Darevsky 1997 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975
 
Darevskia nairensis

Nairi Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003
 
Darevskia obscura

Borjomi Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arribas, Candan, Kornilios, Ayaz, Kumlutas, Gul, Yilmaz, Caynak & Ilgaz 2022 Bakradze & Chkhikvadze 1993 Bischoff 1978 Boehme & Bischoff in Boehme (ed.) 1984 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003
 
Darevskia portschinskii

River Kura Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975
 
Darevskia praticola

Caucasian Meadow Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arakelyan, Danielyan, Corti, Sindaco & Leviton 2011 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Stugren in Boehme (ed.) 1984
 
Darevskia raddei

Azerbaijan Rock Lizard

Arakelyan, Danielyan, Corti, Sindaco & Leviton 2011
 
Darevskia rudis

Spiny-tailed Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Boehme & Bischoff in Boehme (ed.) 1984 Bosch & Bischoff 2004 Fu, Murphy & Darevsky 1997 Sindaco & Jeremchenko 2008 Szczerbak 2003 Terentjev & Chernov 1965 [1949]
 
Darevskia saxicola

Russian Rock Lizard

Darevsky in Boehme (ed.) 1984
 
Darevskia szczerbaki

Anapa Rock Lizard

Milto, Pestov & Bezman-Moseyko 2010
 
Darevskia unisexualis

White-bellied Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Szczerbak 2003
 
Darevskia valentini

Basarchay Rock Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arribas, Candan, Kornilios, Ayaz, Kumlutas, Gul, Yilmaz, Caynak & Ilgaz 2022 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003
Genus Eremias (Desert and Steppe Runners)
 
Eremias arguta

Kazakhstan Steppe Runner

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Szczerbak 1974
 
Eremias velox

Rapid Steppe Runner

Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 1974
Genus Lacerta (Eurasian Green Lizards)
 
Lacerta agilis

Northern Eurasian Sand Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bischoff in Boehme (ed.) 1984 Bosch & Bischoff 2004 Doronin, Doronina & Bekoshvili 2019 Sindaco & Jeremchenko 2008 Szczerbak 2003
 
Lacerta media

Levant Emerald Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Nettmann & Rykena in Boehme (ed.) 1984 Sindaco & Jeremchenko 2008 Szczerbak 2003 Terentjev & Chernov 1965 [1949]
 
Lacerta strigata

Caspian Green Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Darevsky in Boehme (ed.) 1984 Doronin, Doronina & Bekoshvili 2019 Sindaco & Jeremchenko 2008 Szczerbak 2003
Genus Ophisops (Snake-eyed Lizards)
 
Ophisops elegans

Elegant Snake-eyed Lizard

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Darevsky & Beutler in Boehme (ed.) 1981 Szczerbak 2003 Terentjev & Chernov 1965 [1949]
 Family Scincidae (Skinks)
Genus Ablepharus (Eurasian Snake-eyed Skinks)
 
Ablepharus pannonicus

Southwest Asian Snake-eyed Skink

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bosch & Bischoff 2004 Grossmann, Kowalski & Schillert 2013 Szczerbak 2003
Genus Eumeces (Afro-Asian Long-legged Skinks)
 
Eumeces princeps

Southwest Asian Long-legged Skink

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bosch & Bischoff 2004 Terentjev & Chernov 1965 [1949]

Snakes (Serpentes)
= endemicEX = extinct or assumed extinct

Species
References
 Family Boidae (Typical Boas)
Genus Eryx (Sand Boas)
 
Eryx jaculus

Western Sand Boa

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Kudrjavtsev & Mamet 1989 Stimson 1969
 Family Colubridae (Typical Snakes or Colubrids)
Genus Coronella (Palearctic Smooth Snakes)
 
Coronella austriaca

Northern Smooth Snake

Bosch & Bischoff 2004 Engelmann in Boehme (ed.) 1993 Geniez 2018 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Steward 1971 Terentjev & Chernov 1965 [1949]
Genus Dolichophis (Greater Whip Snakes)
 
Dolichophis schmidti

Red-bellied Whip Snake

Arakelyan, Danielyan, Corti, Sindaco & Leviton 2011 Bosch & Bischoff 2004 Geniez 2018 Steward 1971
Genus Eirenis (Middle Eastern Dwarf Snakes)
 
Eirenis collaris

Collared Dwarf Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Kudrjavtsev & Mamet 1989 Mahlow, Tillack, Schmidtler & Muller 2013 Szczerbak 2003 Terentjev & Chernov 1965 [1949]
 
Eirenis modestus

Ring-headed Dwarf Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Mahlow, Tillack, Schmidtler & Muller 2013 Schmidtler & Baran in Boehme (ed.) 1993 Sikmashvili 1975 Szczerbak 2003 Terentjev & Chernov 1965 [1949]
Genus Elaphe (Palearctic-Oriental Rat Snakes)
 
Elaphe dione

Dione Rat Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Obst & Szczerbak in Boehme (ed.) 1993 Schulz 1996 Steward 1971 Terentjev & Chernov 1965 [1949]
 
Elaphe urartica

Urartian Rat Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 [as E. sauromates] Boehme & Szczerbak in Boehme (ed.) 1993 [as E. quatuorlineata] Bosch & Bischoff 2004 [as E. sauromates] Geniez 2018 [as E. sauromates] Jablonski, Kukushkin, Avci, Bunyatova, Kumlutas, Ilgaz, Polyakova, Shiryaev, Tuniyev & Jandzik 2019 Kudrjavtsev & Mamet 1989 [as E. sauromates] Schulz 1996 [as E. sauromates] Szczerbak 2003 [as E. sauromates]
Genus Hemorrhois (Palearctic Whip Snakes)
 
Hemorrhois ravergieri

Mountain Whip Snake

Bosch & Bischoff 2004 Geniez 2018 Kudrjavtsev & Mamet 1989 Terentjev & Chernov 1965 [1949]
Genus Natrix (Eurasian Keelbacks)
 
Natrix natrix

Common European Grass Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 [as megalocephala] Anonymous 2005 Bosch & Bischoff 2004 Geniez 2018 Gruber 1989 [as megalocephala] Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Orlov & Tuniyev 1992 [as megalocephala] Orlov & Tuniyev in Boehme (ed.) 1999 [as megalocephala]
 
Natrix tessellata

Dice Snake

Anonymous 2005 Bosch & Bischoff 2004 Frotzler, Davitashvili & Mebert 2011 Gruschwitz, Lenz, Mebert & Lanka in Boehme (ed.) 1999 Tuniyev, Tuniyev, Kirschey & Mebert 2011
Genus Platyceps (Flat-headed Whip Snakes)
 
Platyceps najadum

European Slender Whip Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Darevsky & Szczerbak in Boehme (ed.) 1993 Geniez 2018
Genus Telescopus (Afro-Asian Cat Snakes)
 
Telescopus fallax

European Cat Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1999 Kudrjavtsev & Mamet 1989 Szczerbak 2003 Terentjev & Chernov 1965 [1949]
Genus Zamenis (Western Palearctic Rat Snakes)
 
Zamenis hohenackeri

Transcaucasian Rat Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bischoff in Boehme (ed.) 1993 Bosch & Bischoff 2004 Geniez 2018 Hofmann, Mebert, Schulz, Helfenberger, Goecmen & Boehme 2018 Kudrjavtsev & Mamet 1989 Nilson & Andren 1984 Schulz 1996 Sikmashvili 1975 Szczerbak 2003
 
Zamenis longissimus

Common Aesculapian Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Boehme in Boehme (ed.) 1993 Bosch & Bischoff 2004 Geniez 2018 Nilson & Andren 1984 Schulz 1996 Sindaco, Venchi & Grieco 2013 Terentjev & Chernov 1965 [1949]
 Family Lamprophiidae (House Snakes, etc.)
Genus Malpolon (Montpellier Snakes)
 
Malpolon insignitus

Eastern Montpellier Snake

Bosch & Bischoff 2004 Geniez 2018 Terentjev & Chernov 1965 [1949] [as M. monspessulanus]
 Family Typhlopidae (Typical Blind Snakes)
Genus Xerotyphlops (Desert Blind Snakes)
 
Xerotyphlops vermicularis

European Worm Snake

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1993 Kudrjavtsev & Mamet 1989 Szczerbak 2003
 Family Viperidae (Vipers)
Genus Macrovipera (Giant Vipers)
 
Macrovipera lebetinus

Blunt-nosed Viper

Bosch & Bischoff 2004 David & Ineich 1999 Geniez 2018 McDiarmid, Campbell & Toure 1999
Genus Vipera (Typical Vipers or Adders)
 
Vipera ammodytes

European Nose-horned Viper

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 David & Vogel 2010 Geniez 2018 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 Kudrjavtsev & Mamet 1989 McDiarmid, Campbell & Toure 1999 Sindaco, Venchi & Grieco 2013 Szczerbak 2003
 
Vipera darevskii

South Caucasian Viper

Tuniyev, Avci, Ilgaz, Olgun, Petrova, Bodrov, Geniez & Teynie 2018 Tuniyev, Iremashvili, Heras & Tuniyev 2014
 
Vipera dinniki

West Caucasian Viper

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 David & Ineich 1999 Geniez 2018 McDiarmid, Campbell & Toure 1999 Orlov & Tuniyev 2005 Sindaco, Venchi & Grieco 2013 Sindaco, Venchi, Carpaneto & Bologna 2000 Tuniyev 2016
 
Vipera kaznakovi

Caucasian Red Viper

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 David & Ineich 1999 David & Vogel 2010 Geniez 2018 Kudrjavtsev & Mamet 1989 McDiarmid, Campbell & Toure 1999 Orlov & Tuniyev 2005 Sindaco, Venchi & Grieco 2013 Sindaco, Venchi, Carpaneto & Bologna 2000 Szczerbak 2003 Tuniyev 2016
 
Vipera lotievi

Caucasian Meadow Viper

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 David & Ineich 1999 McDiarmid, Campbell & Toure 1999 Nilson & Andren 2001 [Russia/Azerbaijan border] Tuniyev 2016 Venchi & Sindaco 2007
 
Vipera shemakhensis

Shemakhan Steppe Viper

Tuniyev 2016 [as schemakhensis [sic]] Tuniyev, Iremashvili, Petrova & Kravchenko 2018
  *
Vipera tuniyevi

Georgian Viper

Ananjeva, Gabaev, Iremashvili, Lotiev & Petrova 2021

Turtles (Testudines)
= endemicEX = extinct or assumed extinct

Species
References
 Family Emydidae (Western Pond Turtles)
Genus Emys (European Pond Turtles)
 
Emys orbicularis

Common European Pond Turtle

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Fritz & Havas 2006 Fritz in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021
 Family Geoemydidae (Eastern Pond Turtles)
Genus Mauremys (Stripe-necked Turtles)
 
Mauremys caspica

Caspian Turtle

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 Leviton, Anderson, Adler & Minton 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Terentjev & Chernov 1965 [1949] Wischuf & Fritz in Fritz (ed.) 2001
 Family Testudinidae (Tortoises)
Genus Testudo (Palearctic Tortoises)
 
Testudo graeca

Spur-thighed Tortoise

Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bonin, Devaux & Dupre 2006 Bosch & Bischoff 2004 Buskirk, Keller & Andreu in Fritz (ed.) 2001 Fritz & Havas 2006 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021