Related bibliographies:
Reptiles
 Lizards
 Helodermatidae
Central America
North America
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Bibliography of the genus
Heloderma (Beaded Lizards)
(Reptilia: Sauria: Helodermatidae)
Note:
In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.
Heloderma in general
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Applegate, R.W. 1999. Heloderma suspectum, horridum. The horrible, suspicious, venomous lizards of the New World. Reptilia (GB) 7: 28-33.
Arlington, O.N. 1930. Notes on the two poisonous lizards with special reference to Heloderma suspectum. Bulletin of the Antivenin Institute of America 4: 29-35.
Ast, J.C. 2001. Mitochondrial DNA evidence and evolution in Varanoidea (squamata). Cladistics 17(3): 211-226.
Beaman, K.R.; Beck, D.D.; McGurty, B.M. 2006. The Beaded Lizard (Heloderma horridum) and Gila Monster (Heloderma suspectum): a bibliography of the family Helodermatidae. Smithsonian Herpetological Information Service 136: 1-66.
Beck, D.D. 2005. Biology of Gila Monsters and Beaded Lizards. University of California Press, Berkeley, Los Angeles & London. 211 pp.
Beck, D.D.; Dohm, M.R.; Garland, T.; Ramirez-Bautista, A.; Lowe, C.H. 1995. Locomotor performance and activity energetics of helodermatid lizards. Copeia 1995(3): 577-585.
Beck, D.D.; Lowe, C.H. 1994. Resting metabolism of helodermatid lizards: allometric and ecological relationships. Journal of Comparative Physiology B Biochemical Systemic and Environmental Physiology 164(2): 124-129.
Beck, D.D.; Martin, B.E.; Lowe, C.H. 2005. Population ecology. pp. 107-117. In: Beck, D.D. Biology of gila monsters and beaded lizards. University of California Press, Berkeley, Los Angeles & London. 211 pp.
Bhullar, B.A.S.; Smith, K.T. 2008. Helodermatid lizard from the Miocene of Florida, the evolution of the dentary in Helodermatidae, and comments on dentary morphology in Varanoidea. Journal of Herpetology 42(2): 286-302.
Bissinger, B.E.; Simon, C.A. 1979. Comparison of tongue extrusions in representatives of six families of lizards. Journal of Herpetology 13(2): 133-139.
Bogert, C.M. 1956. The world's only venomous lizards. Animal Kingdom 59: 105-109.
Bogert, C.M.; Martin del Campo, R. 1952. Geoclines and ontoclines in lizards of the genus Heloderma. Annu. Meet. Amer. Soc. Ichthyol. Herpetol. 32nd., Austin, Texas.
Bogert, C.M.; Martin del Campo, R. 1956. The Gila Monster and its allies. The relationships, habits, and behavior of the lizards of the Family Helodermatidae. Bulletin of the American Museum of Natural History 109: 1-238.
Böhme, W.; Ziegler, T. 2009. A review of iguanian and anguimorph lizard genitalia (Squamata: Chamaeleonidae; Varanoidea, Shinisauridae, Xenosauridae, Anguidae) and their phylogenetic significance: comparisons with molecular data sets. Journal of Zoological Systematics and Evolutionary Research 47(2): 189-202.
Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2. Iguanidae, Xenosauridae, Zonuridae, Anguidae, Anniellidae, Helodermatidae, Varanidae, Xantusiidae, Teiidae, Amphisbaenidae. British Museum (Natural History), London. 497 pp.
Boulenger, G.A. 1891. Notes on the osteology of Heloderma horridum and H. suspectum, with remarks on the systematic position of the Helodermatidae and on the vertebrae of the Lacertilia. Proceedings of the Zoological Society of London 1891(1): 109-118.
Boyer, D.M. 1995. Venomous reptiles: an overview of families, handling, restraint techniques, and emergency protocol. Proceedings of the Association of Reptilian & Amphibian Veterinarians 1995: 83-90.
Branch, W.R. 1982. Hemipeneal morphology of platynotan lizards. Journal of Herpetology 16(1): 16-38.
Brennan, G.A. 1924. A case of death from Heloderma bite. Copeia 1924(129): 45.
Brouwer, K. 1992. The 1991 EEP annual reports. IZN (International Zoo News) 39(6): 8-17.
Brown, D.E.; Carmony, N.B. 1991. Gila monster. Facts and folklore of America's Aztec lizard. High-Lonesome Books, Silver City. 127 pp.
Campbell, J.A.; Lamar, W.W. 1989. The venomous reptiles of Latin America. Cornell University Press, Ithaca, New York. 425 pp.
Campbell, J.A.; Lamar, W.W. 2004. The venomous reptiles of the Western Hemisphere. Vol. 1-2. Comstock Publishing Associates, Ithaca, N.Y. 870 pp.
Caras, R.A. 1978. Dangerous to man: the definitive story of wildlife's reputed dangers. Barrie & Jenkins, London. 327 pp.
Conrad, J.L.; Ast, J.C.; Montanari, S.; Norell, M.A. 2011. A combined evidence phylogenetic analysis of Anguimorpha (Reptilia: Squamata). Cladistics 27(3): 230-277.
Cooper, W.E.; Arnett, J. 2001. Absence of discriminatory tongue-flicking responses to plant chemicals by helodermatid lizards. Southwestern Naturalist 46(3): 405-409.
Cooper, W.E.; Arnett, J. 2003. Correspondence between diet and chemosensory responsiveness by helodermatid lizards. Amphibia-Reptilia 24(1): 86-91.
Duellman, W.E. 1950. A case of Heloderma poisoning. Copeia 1950: 151.
Dufton, M.J. 1993. Kill and cure: the promising future for venom research. Endeavour (Oxford) 17(3): 138-140.
Eidenmüller, B. 1993. Erganzende Bemerkungen zu einem Nachtzuchtbericht von Heloderma suspectum. Salamandra 29(3-4): 258-260.
Eidenmüller, B.; Philippen, H.D. 2008. Varanoid lizards. (Terralog vol. 6). Edition Chimaira, Frankfurt am Main / Rodgau. 148 pp.
Eidenmüller, B.; Reisinger, M. 2011. Beaded lizards & Gila Monsters: captive care & husbandry. Edition Chimaira, Frankfurt am Main. 157 pp.
Elliott, W.B. 1978. Chemistry and immunology of reptilian venoms. pp. 163-436. In: Gans, C. & Gans, K.A. (eds.). Biology of the Reptilia. Vol. 8. Physiology B. Academic Press, London & New York. 782 pp.
Flores-Villela, O.A.; Rios-Muñoz, C.A.; Schwenk, K.; Zamudio-Varela, G.; Magana-Cota, G. 2010. An unpublished manuscript of Alfredo Duges related to the classification of lizards according to tongue morphology, c. 1898. Archives of Natural History 37(2): 246-254.
Gadd, L. 1980. Deadly beautiful: the world's most poisonous animals and plants. Macmillan, London & New York. 208 pp.
Gervais, P. 1873. Du Moloch et de l'Heloderme. Journal of Zoology (London) 1873: 451.
Gienger, C.M.; Beck, D.D. 2007. Heads or tails? Sexual dimorphism in helodermatid lizards. Canadian Journal of Zoology 85(1): 92-98.
Good, D.A.; Bauer, A.M.; Günther, R. 1993. An annotated type catalogue of the anguimorph lizards (Squamata: Anguidae, Helodermatidae, Varanidae, Xenosauridae) in the Zoological Museum, Berlin. Mitteilungen aus dem Zoologischen Museum in Berlin 69(1): 45-56.
Grossman, A.B. 1955. Review of "The systematic position of Lanthonotus and the affinities of the anguinomorphan lizards". Quarterly Review of Biology 30: 400.
Habermehl, G.G. 1981. Venomous animals and their toxins. Springer, Berlin, Heidelberg & New York. 195 pp. (Based on the translation of the 2nd German edition published 1977).
Hensley, M.M. 1949. Mammal diet of Heloderma. Herpetologica 5(6): 152.
Herrel, A.; Vree, F. de 1999. The cervical musculature in helodermatid lizards. Belgian Journal of Zoology 129(1): 175-185.
Hoser, R.T. 2013. Not in Heloderma. A revised taxonomy and new genus for the Gila Monster. Australasian Journal of Herpetology 21: 37-40.
Jennings, M.R. 1984. Longevity records for lizards of the family Helodermatidae. Bulletin of the Maryland Herpetological Society 20(1): 22-23.
Kaup, J.J. 1865. Einige Nachträge zur Gattung Heloderma horridum (Wiegm.). Wiegm. Arch. 1865: 33-40.
Knauer, F. 1908. Giftechsen. Natw. Wochenschr., Jena 23: 393-396.
Kochva, E. 1978. Oral glands of the Reptilia. pp. 43-161. In: Gans, C. & Gans, K.A. (eds.). Biology of the Reptilia. Vol. 8. Physiology B. Academic Press, London & New York. 782 pp.
McCurry, M.R.; Mahony, M.; Clausen, P.D.; Quayle, M.R.; Walmsley, C.W.; Jessop, T.S.; Wroe, S.; Richards, H.; McHenry, C.R. 2015. The relationship between cranial structure, biomechanical performance and ecological diversity in varanoid lizards. PLoS ONE 10(6): e0130625.
Mebs, D. 1978. Pharmacology of reptilian venoms. pp. 437-560. In: Gans, C. & Gans, K.A. (eds.). Biology of the Reptilia. Vol. 8. Physiology B. Academic Press, London & New York. 782 pp.
Mertens, R. 1963. Liste der rezenten Amphibien und Reptilien. Helodermatidae, Varanidae, Lanthanotidae. Das Tierreich 79: 1-26.
Minton, S.A.; Minton, M.R. 1980. Venomous reptiles. Scribners, New York. 308 pp.
Molnar, R.E. 2004. The long and honorable history of monitors and their kin. pp. 10-67. In: Pianka, E.R.; King, D.R.; King, R.A. (eds.). Varanoid lizards of the world. Indiana University Press, Bloomington. 588 pp.
Molnar, R.E.; Pianka, E.R. 2004. Biogeography and phylogeny of varanoids. pp. 68-76. In: Pianka, E.R.; King, D.R.; King, R.A. (eds.). Varanoid lizards of the world. Indiana University Press, Bloomington. 588 pp.
Morris, P.J.; Alberts, A.C. 1996. Determination of sex in white-throated monitors (Varanus albigularis), gila monsters (Heloderma suspectum), and beaded lizards (H. horridum) using two-dimensional ultrasound imaging. Journal of Zoo and Wildlife Medicine 27(3): 371-377.
Morris, P.J.; Henderson, C. 1998. Gender determination in mature gila monsters, Heloderma suspectum, and Mexican beaded lizards, Heloderma horridum, by ultrasound imaging of the ventral tail. Bulletin of the Association of Reptilian and Amphibian Veterinarians 8(4): 4-5.
Murphy, J.B. 2014. Studies on venomous reptiles in zoos and aquariums: Part II. True vipers, Fea's viper, mole vipers, pitvipers, venomous lizards, conclusion. Herpetological Review 45(2): 346-364.
Norris, R.L. 2004. Venom poisoning by North American reptiles. pp. 683-708. In: Campbell, J.A.; Lamar, W.W. The venomous reptiles of the Western Hemisphere. Vol. 2. Comstock Publishing Associates, Ithaca & London. pp. 477-870.
Odermatt, C. 1940. Beiträge zur Kenntnis des Gebisses von Heloderma. Vjschr. naturf. Ges. Zurich 85(3-4): 98-141.
Olson, R.E.; Marx, B.; Rome, R. 1986. Descriptive dentition morphology of lizards of Middle & North America. 1: Scincidae, Teiidae, and Helodermatidae. Bulletin of the Maryland Herpetological Society 22(3): 97-124.
Pianka, E.R.; King, D.R.; King, R.A. (eds.). 2004. Varanoid lizards of the world. Indiana University Press, Bloomington. 588 pp.
Poulin, S.; Ivanyi, C.S. 2003. A technique for manual restraint of helodermatid lizards. Herpetological Review 34(1): 43.
Pregill, G.K.; Gauthier, J.A.; Greene, H.W. 1986. The evolution of helodermatid squamates, with the description of a new taxon and an overview of Varanoidea. Transactions of the San Diego Society of Natural History 21(11): 167-202.
Rieppel, O. 1980. The phylogeny of anguinomorph lizards. Denkschriften der Schweizerischen Naturforschenden Gesellschaft 94: 1-86.
Schmidt, W.J. 1912. Studien am Integument der Reptilien. II. Die Hautverknöcherungen von Heloderma. Zoologische Jahrbücher (Suppl.) (Jena) 15(Bd 2): 219-228.
Schneider, K.M. 1941. Über Fettlager im Schwanz der Krusten-(Heloderma Wiegm.) und Stutz-Echse (Trachydosaurus Gray). Zoologische Garten N.F. (Leipzig) 13: 236-247.
Schuett, G.W.; Reiserer, R.S. 2017. Sampling localities and corresponding haplotype data for helodermatid lizards analyzed in Douglas et al. (2010). Herpetological Review 48(4): 798-801.
Sprackland, R.G. 1993. Venomous lizards. Tropical Fish Hobbyist 41(10): 138-140, 142, 144-145.
Steel, R. 1996. Living dragons: a natural history of the world's monitor lizards. Blandford, London. 160 pp.
Storer, T.I. 1931. Heloderma poisoning in man. Bulletin of the Antivenin Institute of America 5(1): 12-15.
Sullivan, B.K.; Nowak, E.M.; Kwiatkowski, M.A. 2015. Problems with mitigation translocation of herpetofauna. Conservation Biology 29(1): 12-18.
Sullivan, J.B.; Wingert, W.A. 1989. Reptile bites. pp. 479-511. In: Auerbach, P.S. & Geehr, E.D. (eds.). Management of wilderness and environmental emergencies. C.V. Mosby Company, St Louis, Baltimore & Toronto. 2nd edition. 1068 pp.
Thireau, M. 1995. Mors. 10. Les helodermes. pp. 227-229. In: Goyffon, M. & Heurtault, J. La fonction venimeuse. Masson, Paris, Milano & Barcelona. 284 pp.
Trutnau, L. 1970. Ein beitrag zur behandlung erkrankter krustenechsen (Heloderma horridum und Heloderma suspectum). DATZ (Die Aquarien- und Terrarien-Zeitschrift) 23(8): 251-253.
Tu, A.T. (ed.) 1991. Handbook of natural toxins. Vol. 5. Reptile venoms and toxins. Marcel Dekker, Inc., New York, Basel & Hong Kong. 827 pp.
Wermuth, H. 1958. Liste der rezenten Amphibien und Reptilien. Helodermatidae. Das Tierreich 72: 1-16.
Wilms, T. 2006. Krustenechsen - die Gattung Heloderma. Reptilia (D) 11(58): 18-25.
Woodson, W.D. 1947. Toxicity of Heloderma venom. Herpetologica 4(1): 31-33.
Woodson, W.D. 1949. Summary of Heloderma's food habits. Herpetologica 5(4): 91-92.
Wright, K.; Pugh, C.; Walker, L. 1995. Ultrasonographic sexing of helodermatid lizards. pp. 239-242. In: Junge, R.E. (ed.). Proceedings Joint Conference American Association of Zoo Veterinarians, Wildlife Disease Association, American Association of Wildlife Veterinarians, East Lansing, Michigan, August 12-17, 1995. American Association of Zoo Veterinarians, American Association of Wildlife Veterinarians & Wildlide Disease Association, Place of publication not given. 546 pp.
Zima, J. (ed.). 2002. 2001 Gila Monster (Heloderma suspectum) and Beaded Lizard (Heloderma horidum). EEP studbook. Zoological Garden, Usti nad Labem. 33 pp.
Zimmermann, H. 1992. DGHT-Arbeitsgruppe Warane und Krustenechsen. Herpetofauna (Weinstadt) 14(78): 21.
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Heloderma alvarezi
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Reiserer, R.S.; Schuett, G.W.; Beck, D.D. 2013. Taxonomic reassessment and conservation status of the beaded lizard, Heloderma horridum (Squamata: Helodermatidae). Amphibian and Reptile Conservation 7(1): 74-96.
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Heloderma charlesbogerti
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Anzueto-V., R.; Campbell, J.A. 2010. Guatemalan Beaded Lizard (Heloderma horridum charlesbogerti) on the Pacific versant of Guatemala. Southwestern Naturalist 55(3): 453-454.
Ariano-Sanchez, D. 2006. The Guatemalan Beaded Lizard: endangered inhabitant of a unique ecosystem. Iguana (West Hartford) 13(3): 178-184.
Ariano-Sánchez, D.; Salazar, G. 2012. Natural history notes: Heloderma horridum charlesbogerti (Guatemalan Beaded Lizard). Shelter use. Herpetological Review 43(4): 645-646.
Ariano-Sánchez, D.; Salazar, G. 2013. Natural history notes: Heloderma horridum charlesbogerti (Guatemalan Beaded Lizard). Wild reproductive ecology. Herpetological Review 44(2): 324.
Ariano-Sánchez, D.; Salazar, G. 2015. Spatial ecology of the endangered Guatemalan Beaded Lizard, Heloderma charlesbogerti (Sauria: Helodermatidae), in a tropical dry forest of the Motagua Valley, Guatemala. Mesoamerican Herpetology 2(1): 64–74.
Hill, R.L.; Lock, B.; Brock, J.L.; Brothers, D.J.; Wade, C. 2014. Heloderma charlesbogerti (Guatemalan beaded lizard). Cannibalism. Herpetological Review 45(3): 455.
Kunz, K. 2006. Giftige Riesen vor dem Aus? REPTILIA-Projekt zum Schutz der Guatemala-Skorpionskrustenechse. Reptilia (D) 11(58): 39-42.
Kunz, K. 2007. Fortschritte beim Projekt zum Schutz der Guatemala-Skorpionskrustenechse. ZGAP Mitteilungen 23(1): 14-15.
Melendez Garcia, L.S. 2006. Die Guatemala-Skorpionskrustenechse, Heloderma horridum charlesbogerti Campbell & Vannini, 1998. Reptilia (D) 11(58): 36-38.
Melendez Garcia, L.S. 2006. Die Guatemala-Skorpionskrustenechse. ZGAP Mitteilungen 22(1): 16-17.
Owens, T.C. 2006. Ex-situ: notes on reproduction and captive husbandry of the Guatemalan Beaded Lizard (Heloderma horridum charlesbogerti). Iguana (West Hartford) 13(3): 212-215.
Reiserer, R.S.; Schuett, G.W.; Beck, D.D. 2013. Taxonomic reassessment and conservation status of the beaded lizard, Heloderma horridum (Squamata: Helodermatidae). Amphibian and Reptile Conservation 7(1): 74-96.
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Heloderma exasperatum
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Lemos-Espinal, J.A.; Chiszar, D.; Smith, H.M. 2003. Presence of the Rio Fuerte Beaded Lizard (Heloderma horridum exasperatum) in western Chihuahua. Bulletin of the Maryland Herpetological Society 39(2): 47-51.
Radovanovic, A. 2014. Captive husbandry and management of the Rio Fuerte Beaded Lizard Heloderma exasperatum. Herpetological Bulletin 130: 6-8.
Reiserer, R.S.; Schuett, G.W.; Beck, D.D. 2013. Taxonomic reassessment and conservation status of the beaded lizard, Heloderma horridum (Squamata: Helodermatidae). Amphibian and Reptile Conservation 7(1): 74-96.
Reisinger, M. 2001. Haltung und Nachzucht von Heloderma horridum exasperatum. Elaphe 9(2): 18-21.
Reisinger, M. 2006. Die erfolgreiche und kontinuierliche Haltung und Nachzucht der Skorpions-Krustenechse Heloderma horridum exasperatum. Reptilia (D) 11(58): 26-35.
Reisinger, M. 2006. Successful long-term keeping and breeding of the Rio Fuerte Beaded Lizard. Heloderma horridum exasperatum. Reptilia (GB) 47: 25-33.
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Heloderma horridum
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Anonymous. 1984. Rare lizard shines at zoo. Chicago Herpetological Society Newsletter 1984(April): 1 p. (Unpaginated. Reprinted from Chicago Sun Times, March 8, 1984).
Aguilar-Miguel, X.; Monroy-Vilchis, O.; Casas-Andreu, G. 2003. Geographic distribution: Heloderma horridum horridum (Mexican Beaded Lizard). Herpetological Review 34(4): 384.
Ahumada-Carrillo, I.T.; Hernández-Jiménez, C.A.; García-Vázquez, U.O. 2011. Geographic distribution: Heloderma horridum (Mexican Beaded Lizard). Herpetological Review 42(1): 111.
Anstanding, L.M. 1984. The captive breeding and rearing of the Mexican bearded lizard, Heloderma horridum, at the Detroit Zoo. Annual Reptile Symposium on Captive Propagation and Husbandry Proceedings 7: 64-73.
Ariano-Sánchez, D.; Muñoz-Alonso, A.; Marquez, L.C.; Acevedo, M. 2014. Heloderma horridum. The IUCN Red List of Threatened Species 2014: e.T9864A3152367. http://dx.doi.org/10.2305/IUCN.UK.2014-1.RLTS.T9864A3152367.en.
Avila-Villegas, H. 2007. Geographic distribution: Heloderma horridum horridum (Mexican Beaded Lizard). Mexico: Zacatecas. Herpetological Review 38(2): 218.
Balderas-Valdivia, C.J. 2004. Reconocimiento diferencial de los depredadores y variación del comportamiento defensivo de Heloderma horridum en una población de la selva decidua de Jalisco, México. Boletin de la Sociedad Herpetologica Mexicana 12(2): 59.
Balderas-Valdivia, C.J. 2005. Respuesta congénita y variación del comportamiento aversivo en Heloderma horridum. Boletin de la Sociedad Herpetologica Mexicana 13(1): 1-9.
Balderas-Valdivia, C.J.; Ramirez-Bautista, A. 2005. Aversive behavior of Beaded Lizard, Heloderma horridum, to sympatric and allopatric predator snakes. Southwestern Naturalist 50(1): 24-31.
Bañuelos-Alamillo, J.A.; Carbajal-Márquez, R.A.; Quintero-Díaz, G.E.; Moreno-Ochoa, G. 2015. Geographic distribution: Heloderma horridum (Mexican Beaded Lizard). Herpetological Review 46(3): 383.
Beck, D.D.; Lowe, C.H. 1991. Ecology of the beaded lizard, Heloderma horridum, in a tropical dry forest in Jalisco, Mexico. Journal of Herpetology 25(4): 395-406.
Beck, D.D.; Ramirez-Bautista, A. 1991. Combat behavior of the beaded lizard, Heloderma h. horridum, in Jalisco, Mexico. Journal of Herpetology 25(4): 481-484.
Camarillo-Rangel, J.L.; Aguilar-Cortes, R.; Gonzalez-Ruiz, A. 1985. Distributional records of amphibians and reptiles from the State of Mexico. Herpetological Review 16(3): 85.
Campbell, J.A.; Vannini, J.P. 1988. A new subspecies of beaded lizard, Heloderma horridum, from the Motagua Valley of Guatemala. Journal of Herpetology 22(4): 457-468.
Cochran, D.M. 1944. Dangerous reptiles (popular). Rep. Smithson. Instn. 1943: 275-325.
Conant, R. 1965. Old-timers among the reptiles. America's First Zoo 17: 32-34.
Conners, J.S. 1993. A long-term breeding programme for the beaded lizard Heloderma horridum at Detroit Zoo. International Zoo Yearbook 32: 184-188.
Cuesta Terron, C. 1934. El escorpion. Heloderma horridum Wiegmann. 2nd ed. revised by Rafel Martin del Campo. pp. 1-14. Folletos Divulgacion Client., Inst. Biol., Univ. Nac. Mexico.
Curtis, L. 1949. Notes on the eggs of Heloderma horridum. Herpetologica 5(6): 148.
Davis, W.B.; Dixon, J.R. 1961. Reptiles (exclusive of snakes) of the Chilpancingo region, Mexico. Proceedings of the Biological Society of Washington 74: 37-56.
Domiguez-Vega, H.; Monroy-Vilchis, O.; Balderas-Valdivia, C.J.; Gienger, C.M.; Ariano-Sánchez, D. 2012. Predicting the potential distribution of the beaded lizard and identification of priority areas for conservation. Journal for Nature Conservation 20(4): 247-253.
Douglas, M.E.; Douglas, M.R.; Schuett, G.W.; Beck, D.D.; Sullivan, B.K. 2010. Conservation phylogenetics of helodermatid lizards using multiple molecular markers and a supertree approach. Molecular Phylogenetics and Evolution 55(1): 153-167.
Duges, A. 1897. Sur l'Heloderma horridum, Wiegm. Act. Soc. Chili 7: 113-117.
Furrer, S. 2004. Die erfolgreiche Nachzucht der Skorpionskrustenechse (Heloderma horridum exasperatum) im Zoo Zürich, Schweiz. Reptilia (D) 9(49): 76-78.
Gienger, C.M.; Walker-Johnson, G.; McMillan, M.; Sheldon, S.; Tracy, C.R. 2005. Timing of hatching in Beaded Lizards (Heloderma horridum). Sonoran Herpetologist 18(8): 93-94.
Goldberg, S.R.; Beck, D.D. 2001. Natural history notes: Heloderma horridum (Beaded Lizard). Reproduction. Herpetological Review 32(4): 255-256.
Gonzalez-Ruiz, A.; Godinez-Cano, E.; Rojas-Gonzalez, I. 1996. Captive reproduction of the Mexican acaltetepon, Heloderma horridum. Herpetological Review 27(4): 192.
Herrel, A.; Wauters, I.; Aerts, P.; Vree, F. de 1997. The mechanics of ovophagy in the beaded lizard (Heloderma horridum). Journal of Herpetology 31(3): 383-393.
Honegger, R.E. 1998. Beitrag zu Haltung und Zucht der Skorpionskrustenechse, Heloderma horridum, im Zoo Zürich. Zoologische Garten 68(5): 287-299.
Loc-Barragán, J.A.; Ramírez-Silva, J.P.; Woolrich-Piña, G.A. 2016. Distribution notes: Heloderma horridum (Weigmann, 1829). Mesoamerican Herpetology 3(3): 766-767.
Monroy-Vilchis, O.; Hernández-Gallegos, O.; Rodriguez-Romero, F. 2005. Natural history notes: Heloderma horridum horridum (Mexican Beaded Lizard). Unusual habitat. Herpetological Review 36(4): 450.
Muniz-Martinez, R.; Rojas-Perez, M.A. 2009. Registro nuevo del escorpion mexicano Heloderma horridum (Reptilia: Helodermidae) en Durango, Mexico. Revista Mexicana de Biodiversidad 80(3): 871-873.
Ottley, J.R. 1981. Geographic distribution: Heloderma horridum exasperatum. Herpetological Review 12(2): 64.
Philippen, G. 1989. Neue Unterart der Krusteneidechse aus dem Motagua-Tal in Guatemala. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 42(10): 588.
Reiserer, R.S.; Schuett, G.W.; Beck, D.D. 2013. Taxonomic reassessment and conservation status of the beaded lizard, Heloderma horridum (Squamata: Helodermatidae). Amphibian and Reptile Conservation 7(1): 74-96.
Sánchez-de la Vega, G.; Buenrostro-Silva, A.; García-Grajales, J.; Mata-Silva, V. 2012. Geographic distribution: Heloderma horridum (Mexican Beaded Lizard). Herpetological Review 43(1): 102.
Shaw, C.E. 1964. Beaded lizards - dreaded, but seldom deadly. Zoonooz 37(3): 10-15.
Taub, A.M. 1963. On the longevity and fecundity of Heloderma horridum horridum. Herpetologica 19: 149.
Trutnau, L. 1970. Die Skorpion-Krustenechse-Heloderma horridum (Wiegmann). Aquarien Terrarien 17: 228-231.
Trutnau, L. 1976. Zum Fortpflanzungsverhalten der Skorpions - Krustenechse (Heloderma h. horridum). DATZ (Die Aquarien- und Terrarien-Zeitschrift) 29(12): 424-428.
Zweifel, R.G.; Norris, K.S. 1955. Contribution to the herpetology of Sonora, Mexico: descriptions of new subspecies of snakes (Micruroides euryxanthus and Lampropeltis getulus) and miscellaneous collecting notes. American Midland Naturalist 54: 230-249.
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Heloderma suspectum
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