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Taxonomy of the genus Sauromalus Biodiversity of the family Iguanidae

Bibliography of the genus
Sauromalus (North American Chuckwallas)

(Reptilia: Sauria: Iguanidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Sauromalus in general

Avery, D.F.; Tanner, W.W. 1964. The osteology and myology of the head and thorax regions of the obesus group of the genus Sauromalus Dumeril (Iguanidae). Brigham Young University Science Bulletin Biol. 5(3): 1-30.

Bustard, R. 1961. Collared lizards and chuckawallas. Aquarist and Pondkeeper 26: 134-135.

Dziewa, T. 1963. Mealworms in the diet of the chuckwalla. Bulletin of the Philadelphia Herpetological Society 11(3-4): 57.

Etheridge, R.E. 1982. Checklist of the iguanine and Malagasy iguanid lizards. pp. 7-37. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Hollingsworth, B.D. 1998. The systematics of Chuckwallas (Sauromalus) with a phylogenetic analysis of other iguanid lizards. Herpetological Monographs 12: 38-191.

Hollingsworth, B.D. 1999. Erratum: The systematics of Chuckwallas (Sauromalus) with a phylogenetic analysis of other iguanid lizards. Herpetological Monographs 13: 195.

Hollingsworth, B.D. 2004. Understanding the diversity of Chuckwallas (genus Sauromalus) and a debate over names. Iguana (West Hartford) 11(2): 78-86.

Köhler, G.; Zorn, H. 2014. Chuckwallas. Herpeton, Offenbach. 142 pp.

Littell, D. 1984. Weathering the seasons chuckwalla style. Zoonooz 57(5): 9-13.

Montanucci, R.R. 2004. A note on the identity of chuckwallas inhabiting Isla Danzante, Baja California Sur. Herpetological Review 35(3): 223-224.

Norris, K.S.; Dawson, W.R. 1964. Obserevations on the water economy and electrolyte excretion of chuckwallas (Lacertilia, Sauromalus). Copeia 1964: 638-646.

Petren, K.; Case, T.J. 1997. A phylogenetic analysis of body size evolution and biogeography in Chuckwallas (Sauromalus) and other iguanines. Evolution 51(1): 206-219.

Petren, K.; Case, T.J. 2002. Updated mtDNA phylogeny for Sauromalus and implications for the evolution of gigantism. Appendix 8.1. pp. 574-579. In: Case, T.J.; Cody, M.L.; Ezcurra, E. (eds.). A new island biogeography of the Sea of Cortes. Oxford University Press, Oxford, New York, etc. 669 pp.

Robinson, M.D. 1972. Chromosomes, protein polymorphism, and systematics of insular chuckwalla lizards (genus Sauromalus) in the Gulf of California, Mexico. Dissertation Abstracts International (B) 33(2): 965.

Robinson, M.D. 1974. Chromosomes of the insular species of chuckwalla lizards (genus Sauromalus) in the Gulf of California, Mexico. Herpetologica 30(2): 162-167.

Ryan, M.J. 1982. Variation in iguanine social organization: mating systems in chuckawallas (Sauromalus). pp. 380-390. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Schimmelpfennig, U.; Schimmelpfennig, J. 2013. Der Chuckwalla – Sonnenanbeter in der Felswüste. Reptilia (D) 18(102): 60-66.

Shaw, C.E. 1945. The chuckwallas, genus Sauromalus. Transactions of the San Diego Society of Natural History 10(15): 269-306.

Werning, H. 2002. Die Könige der nordamerikanischen Wüste: Chuckwallas. Draco 3(2) (10): 66-73.

Werning, H. 2015. Chuckwalla-Anekdoten. Teil 1. Reptilia (D) 20(112): ?

Wolfel, H. 1999. Chuckwallas. Die Gattung Sauromalus sowie Haltung und Vermehrung von Sauromalus obesus. Reptilia (D) 4(17): 51-58.

Sauromalus ater

Abts, M.L. 1987. Environment and variation in life history traits of the chuckwalla, Sauromalus obesus. Ecological Monographs 57(3): 215-232.

Abts, M.L. 1988. Reproduction in the saxicolous desert lizard, Sauromalus obesus: the female reproductive cycle. Copeia 1988(2): 382-393.

Abts, M.L. 1988. Reproduction in the saxicolous desert lizard, Sauromalus obesus: the male reproductive cycle. Herpetologica 44(4): 404-415.

Aucone, B. 2004. Captive husbandry of the Chuckwalla (Sauromalus ater). Iguana (West Hartford) 11(2): 114-118.

Barrows, C.W. 2011. Sensitivity to climate change for two reptiles at the Mojave-Sonoran Desert interface. Journal of Arid Environments 75(7): 629-635.

Baumer, M.; Drew, F.C. 2011. Using livestock markers to individually identify communally-housed Chuckwallas (Sauromalus ater) at the Bronx Zoo. Herpetological Review 42(1): 51-52.

Baumer, M.; Drew, F.C.; Casey, B.; Titus, V. 2012. Successful incubation of Common Chuckwalla (Sauromalus ater) eggs at the Bronx Zoo using suspended incubation method. Herpetological Review 43(4): 597-599.

Beaman, K.R.; Toohey, D.R.; Crandall, G.T. 2003. Natural history notes: Sauromalus obesus (Common Chuckwalla). Albinism. Herpetological Review 34(3): 246.

Berry, K.H. 1974. The ecology and social behavior of the chuckwalla, Sauromalus obesus obesus Baird. University of California Publications in Zoology 101: 1-60.

Blodgett, N.E.; Grandmaison, D.D. 2011. Natural history notes: Sauromalus ater (Common Chuckwalla). Entrapment. Herpetological Review 42(4): 608.

Boarman, W.I.; Berry, K.H. 1995. Common ravens in the southwestern United States, 1968-92. pp. 73-75. In: LaRoe, E.T.; Farris, G.S.; Puckett, C.E.; Doran, P.D. & Mac, M.J. (eds.). Our living resources: a report to the nation on the distribution, abundance, and health of U.S. plants, animals and ecosystems. U.S. Department of the Interior, National Biological Service, Washington, D.C. 530 pp.

Boundy, J.; Balgooyen, T.G. 1988. Record lengths for some amphibians and reptiles from the western United States. Herpetological Review 19(2): 26-27.

Brattstrom, B.H. 2001. Comments on the proposed precedence of the specific name of Euphryne obesus Baird, 1858 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Squamata). Bulletin of Zoological Nomenclature 58(4): 307.

Brown, L.E. 2001. Comments on the proposed precedence of the specific name of Euphryne obesus Baird, 1858 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Squamata). Bulletin of Zoological Nomenclature 58(4): 307.

Case, T.J. 1972. Thermoregulation and evaporative cooling in the chuckwalla, Sauromalus obesus. Copeia 1972(1): 145-150.

Case, T.J. 1975. Ecological and physiological studies of desert lizard. Dissertation Abstracts International (B) 35(9): 4330.

Case, T.J. 1976. Body size differences between populations of the chuckwalla, Sauromalus obesus. Ecology 57(2): 313-323.

Case, T.J. 1976. Seasonal aspects of thermoregulatory behaviour in the chuckawalla, Sauromalus obesus (Reptilia, Lacertilia, Iguanidae). Journal of Herpetology 10(2): 85-95.

Clark, H.O. 2008. Common Chuckwalla (Sauromalus ater): behavior. Sonoran Herpetologist 21(5): 54.

Clause, A.G.; Condon, K. 2016. Geographic distribution: Sauromalus ater (Common Chuckwalla). Herpetological Review 47(2): 260.

Cliff, F.S. 1958. A new species of Sauromalus from Mexico. Copeia 259-261.

Cooper, W.E.; Flowers, M. 2001. Plant chemical discriminations by an herbivorous iguanid lizard, Sauromalus ater. Amphibia-Reptilia 22(1): 69-80.

Crampton, B. 1972. The desert chuckwalla. Animals 14(8): 346-347.

Crawford, E.C.; Kampe, G. 1971. Physiological responses of the lizard Sauromalus obesus to changes in ambient temperature. American Journal of Physiology 220: 1256-1260.

Crooks, B. 1983. The care and breeding of Sauromalus obesus obesus in captivity. British Herpetological Society Bulletin 7: 68-71.

Crooks, W. 1980. The chuckwalla (Sauromalus obesus). Herptile 25(4): 29-31.

Deban, S.M.; O'Reilly, J.C.; Theimer, T. 1994. Mechanism of defensive inflation in the Chuckwalla, Sauromalus obesus. Journal of Experimental Zoology 270(5): 451-459.

Dugan, E.A.; Daniels, B.; Torre, G. de la 2017. Natural history notes: Crotalus ruber (Red Diamond Rattlesnake). Diet. Herpetological Review 48(1): 210.

Espinoza, R.E.; Carreno, C.A.; Tracy, C.R.; Tracy, C.R. 1998. Geographic distribution: Sauromalus obesus obesus (Western Chuckwalla). Herpetological Review 29(1): 51-52.

Firth, B.T.; Turner, J.S.; Ralph, C.L. 1989. Thermoregulatory behaviour in two species of iquanid lizards (Crotaphytus collaris and Sauromalus obesus): diel variation and the effect of pinealectomy. Journal of Comparative Physiology B Biochemical Systemic and Environmental Physiology 159(1): 13-20.

Gates, G.O. 1968. Geographical distribution and character analysis of the iguanid lizard Sauromalus obesus in Baja California, Mexico. Herpetologica 24: 285-288.

Gaulden, R. 1998. Life on the rocks: Sauromalus obesus. Reptile & Amphibian Magazine 56: 62-69.

Glaudas, X.; Heindl, A.L. 2007. Natural history notes: Crotaphytus bicinctores (Great Basin Collared Lizard). Diet. Herpetological Review 38(4): 453.

Goode, M.J.; Horrace, W.C.; Sredl, M.J.; Howland, J.M. 2005. Habitat destruction by collectors associated with decreased abundance of rock-dwelling lizards. Biological Conservation 125(1): 47-54.

Grismer, L.L.; McGuire, J.A.; Hollingsworth, B.D. 1994. A report on the herpetofauna of the Vizcaino Peninsula, Baja California, Mexico, with a discussion of its biogeographic and taxonomic implications. Bulletin of the Southern California Academy of Sciences 93(2): 45-80.

Hague, M.T.J.; Routman, E.J. 2016. Does population size affect genetic diversity? A test with sympatric lizard species. Heredity 116(1): 92-98.

Hansen, R.M. 1974. Dietary of the chuckwalla Sauromalus obesus, determined by dung analysis. Herpetologica 30(2): 120-123.

Hollingsworth, B.D.; Grismer, L.L. 1996. Geographic distribution: Sauromalus ater ater (Espiritu Santo Chuckwalla). Herpetological Review 27(1): 33.

Hollingsworth, B.D.; Grismer, L.L.; Wong, H.; Cryder, M.R. 1997. Geographic distribution: Sauromalus ater ater (Espiritu Santo Chuckwalla). Herpetological Review 28(1): 51.

International Commission on Zoological Nomenclature. 2004. Opinion 2072 (Case 3143). Euphryne obesa Baird, 1858 (currently Sauromalus obesus Reptilia, Squamata): proposal to give the specific name precedence over that of Sauromalus ater Dumeril, 1856 not approved. Bulletin of Zoological Nomenclature 61(1): 74-75.

Johnson, S.R. 1965. An ecological study of the chuckwalla, Sauromalus obesus Baird in the western Mojave Desert. American Midland Naturalist 73: 1-29.

Koleska, D.; Svobodova, V.; Husák, T.; Kulma, M.; Jablonski, D. 2017. Tail bifurcation recorded in Sauromalus ater. Herpetology Notes 10: 363-364.

Krysko, K.L. 2001. Comments on the proposed precedence of the specific name of Euphryne obesus Baird, 1858 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Sauria). Bulletin of Zoological Nomenclature 58(3): 229.

Kwiatkowski, M.A. 2003. Variation in conspicuousness among populations of an iguanid lizard, Sauromalus obesus (=ater). Copeia 2003(3): 481-492.

Kwiatkowski, M.A.; Sullivan, B.K. 2002. Geographic variation in sexual selection among populations of an iguanid lizard, Sauromalus obesus (=ater). Evolution 56(10): 2039-2051.

Kwiatkowski, M.A.; Sullivan, B.K. 2002. Mating system structure and population density in a polygynous lizard, Sauromalus obesus (= ater). Behavioral Ecology 13(2): 201-208.

Lamb, T.; Jones, T.R.; Avise, J.C. 1992. Phylogeographic histories of representative herpetofauna of the southwestern U.S.: mitochondrial DNA variation in the desert iguana (Dipsosaurus dorsalis) and the Chuckwalla (Sauromalus obesus). Journal of Evolutionary Biology 5(3): 465-480.

Lappin, A.K.; Hamilton, P.S.; Sullivan, B.K. 2006. Bite-force performance and head shape in a sexually dimorphic crevice-dwelling lizard, the Common Chuckwalla (Sauromalus ater (= obesus)). Biological Journal of the Linnean Society 88(2): 215-222.

Mayhew, W.W. 1963. Some food preferences of captive Sauromalus obesus. Herpetologica 19: 10-18.

McDiarmid, R.W.; Queiroz, K. de; Beaman, K.; Crother, B.; Etheridge, R.E.; Flores-Villela, O.A.; Frost, D.; Grismer, L.L.; Hollingsworth, B.D.; Kearney, M.; McGuire, J.A.; Wright, J.; Zug, G.R. 2002. Comment on the proposed precedence of the specific name of Euphryne obesus Baird, 1859 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Squamata). Bulletin of Zoological Nomenclature 59(1): 45-48.

McGinnis, S.M.; Falkenstein, M. 1971. Thermoregulatory behaviour in three sympatric species of iguanid lizards. Copeia 1971: 552-554.

Montanucci, R.R. 1997. Captive management, behavior and conservation of Chuckwallas, Sauromalus obesus (Lacertilia: Iguanidae). Bulletin of the Chicago Herpetological Society 32(6): 121-137.

Montanucci, R.R. 1999. Preferential coprophagy in neonate Chuckwallas (Sauromalus, Lacertilia, Iguanidae). Herpetological Review 30(4): 221-222.

Montanucci, R.R. 2000. Comments on the restriction of the type locality and nomenclature of the chuckwalla, Sauromalus ater Dumeril, 1856. Herpetological Review 31(3): 138-142.

Montanucci, R.R. 2001. A clarification of the type locality and collector of the holotype of Sauromalus obesus (Baird) 1858. Herpetological Review 32(1): 17-20.

Montanucci, R.R. 2008. Historical evidence for the type locality of Sauromalus ater Dumeril, 1856. Herpetological Review 39(3): 326-328.

Montanucci, R.R.; Smith, H.M.; Adler, K.; Auth, D.L.; Axtell, R.W.; Case, T.J.; Chiszar, D.; Collins, J.T.; Conant, R.; Murphy, R.; Petren, K.; Stebbins, R.C. 2001. Case 3143. Euphryne obesus Baird, 1858 (Reptilia, Squamata): proposed precedence of the specific name over that of Sauromalus ater Dumeril, 1856. Bulletin of Zoological Nomenclature 58(1): 37-40.

Muchlinski, A.E.; Hogan, J.M.; Stoutenburgh, R.J. 1990. Body temperature regulation in a desert lizard, Sauromalus obesus, under undisturbed field conditions. Comparative Biochemistry and Physiology A Comparative Physiology 95(4): 579-583.

Nagy, K.A. 1973. Behavior, diet and reproduction in a desert lizard, Sauromalus obesus. Copeia 1973(1): 93-102.

Nagy, K.A. 1988. Seasonal patterns of water and energy balance in desert vertebrates. Journal of Arid Environments 14(2): 201-210.

Nagy, K.A. 2002. Comment on the proposed precedence of the specific name of Euphryne obesus Baird, 1858 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Squamata). Bulletin of Zoological Nomenclature 59(3): 205.

Niemarkt, J. 2003. Zur Haltung und Nachzucht des Chuckwallas (Sauromalus obesus obesus). Iguana Rundschreiben 16(1): 17-22.

Porter, W.P.; Sabo, J.L.; Tracy, C.R.; Reichman, O.J.; Ramankutty, N. 2002. Physiology on a landscape scale: plant-animal interactions. Integrative and Comparative Biology 42(3): 431-453.

Prieto, A.A. 1972. The ethology of the Arizona chuckwalla (Sauromalus obesus tumidus.). Dissertation Abstracts International (B) 32(9): 5540.

Prieto, A.A.; Ryan, M.J. 1978. Some observations of the social behavior of the Arizona chuckwalla, Sauromalus obesus tumidus (Reptilia, Lacertilia, Iguanidae). Journal of Herpetology 12(3): 327-336.

Prieto, A.A.; Sorenson, M.W. 1975. Food preferences of the Arizona chuckwalla (Sauromalus obesus tumidus). Bulletin New Jersey Academy of Science 20(1): 8-11.

Prieto, A.A.; Sorenson, M.W. 1975. Predator-prey relationships of the Arizona chuckwalla (Sauromalus obesus tumidus). Bulletin New Jersey Academy of Science 20(1): 12-13.

Prieto, A.A.; Sorenson, M.W. 1977. Reproduction in the Arizona chuckwalla, Sauromalus obesus tumidus (Shaw). American Midland Naturalist 98(2): 463-469.

Repp, R. 1998. Wintertime observations on five species of reptiles in the Tucson area: sheltersite selections/fidelity to sheltersites/notes on behavior. Bulletin of the Chicago Herpetological Society 33(3): 49-56.

Rorabaugh, J.C. 2003. Apparent accidental death chuckwalla meets agave. Sonoran Herpetologist 16(4): 44-45.

Ruppert, R.M. 1980. Comparative assimilation efficiencies of two lizards. Comparative Biochemistry and Physiology A Comparative Physiology 67(3): 491-496.

Sahner, A.; Geyer, B.; Selzer, D. 2007. Anorexie bei einer Echse. Tieraerztliche Praxis Ausgabe K Kleintiere Heimtiere 35(3): 213, 219-221.

Salt, G.W. 1943. The lungs and inflation mechanism of Sauromalus obesus. Copeia 1943: 193.

Sanborn, S.R. 1972. Food habits of Sauromalus obesus obesus on the Nevada test site. Journal of Herpetology 6(2): 142-144.

Savage, J.M. 2001. Comments on the proposed precedence of the specific name of Euphryne obesus Baird, 1858 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Squamata). Bulletin of Zoological Nomenclature 58(4): 307-308.

Schneider, H. 1988. Das Naturerlebnis 'chuckwalla'. Aquaria (St. Gallen) 35(1): 10-14.

Shaw, C.E. 1939. Food habits of the chuckwalla, Sauromalus obesus. Herpetologica 1939: 153.

Shaw, C.E. 1952. Notes on the eggs and young of some United States and Mexican lizards. 1. Herpetologica 8(3): 71-79.

Shaw, C.E. 1969. The obese vegetarians. Zoonooz 42: 6-11.

Shaw, C.E. 2004. Food habits of the Chuckwalla, Sauromalus obesus. Iguana (West Hartford) 11(2): 124.

Shuttleworth, T.J.; Thompson, J.L.; Dantzler, W.H. 1987. Potassium secretion by nasal salt glands of desert lizard Sauromalus obesus. American Journal of Physiology 253: 83-90.

Smith, H.M.; Montanucci, R.R. 2001. Comments on the proposed precedence of the specific name of Euphryne obesus Baird, 1858 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Squamata). Bulletin of Zoological Nomenclature 58(4): 308.

Smith, P.W.; Hensley, M.M. 1959. Notes on a small collection of amphibians and reptiles from the vicinity of the Pinacate lava cap in north-western Sonora, Mexico. Transactions of the Kansas Academy of Sciences 61: 64-76.

Smits, A.W.; Yorke, C.D. 1980. Winter activity and mortality in juvenile chuckwallas (Sauromalus obesus). Journal of Herpetology 14(1): 100-101.

Sullivan, B.K.; Flowers, M.A. 1998. Large iguanid lizards of urban mountain preserves in northern Phoenix, Arizona. Herpetological Natural History 6(1): 13-22.

Sullivan, B.K.; Kwiatkowski, M.A.; Hamilton, P.S. 2004. Growth in Sonoran Desert populations of the Common Chuckwalla (Sauromalus obesus). Western North American Naturalist 64(1): 137-140.

Sullivan, B.K.; Sullivan, K.O. 2008. Common Chuckwalla (Sauromalus ater) populations in the Phoenix metropolitan area: stability in urban preserves. Herpetological Conservation and Biology 3(2): 149-154.

Sullivan, B.K.; Sullivan, K.O. 2012. Common Chuckwalla (Sauromalus ater) in an urban preserve: persistence of a small population and estimation of longevity. Herpetological Conservation and Biology 7(3): 437-441.

Sullivan, B.K.; Williams, R.E. 2010. Common Chuckwallas (Sauromalus ater) in urban preserves: do food plants or crevice retreats influence abundance? Herpetological Conservation and Biology 5(1): 102-110.

Switak, K.H. 1977. Leben in der Wüste. Der Chuckwalla. Aquarium (Bornheim) 11(93): 128-131.

Tanner, W.W. 1958. Herpetology of Glen Canyon of the Upper Colorado River Basin. Herpetologica 14: 193-193.

Tanner, W.W.; Avery, D.F. 1964. A new Sauromalus obesus from the upper Colorada Basin of Utah. Herpetologica 20: 38-42.

Taylor, H.L. 2001. Comments on the proposed precedence of the specific name of Euphryne obesus Baird, 1858 over that of Sauromalus ater Dumeril, 1856 (Reptilia, Sauria). Bulletin of Zoological Nomenclature 58(3): 229.

Templeton, J.R. 1967. Panting and pulmonary inflation, two mutually exclusive responses in the chuckwalla, Sauromalus obesus. Copeia 1967: 224-225.

Tracy, C.R. 1999. Differences in body size among Chuckwalla (Sauromalus obesus) populations. Ecology (Washington, D.C.) 80(1): 259-271.

Tracy, C.R. 2004. Environmental influences on body size of two species of herbivorous desert lizards. pp. 158-175. In: Alberts, A.C.; Carter, R.L.; Hayes, W.K.; Martins, E.P. (eds.). Iguanas: biology and conservation. University of California Press, Berkeley, Los Angeles & London. 356 pp.

Tracy, C.R.; Diamond, J. 2005. Regulation of gut function varies with life-history traits in chuckwallas (Sauromalus obesus: Iguanidae). Physiological and Biochemical Zoology 78(4): 469-481.

Tracy, C.R.; Tracy, C.R.; Turner, J.S. 1992. Contrasting physiological abilities for heating and cooling in an amphibian (Rana pipiens) and reptile (Sauromalus obesus). Herpetologica 48(1): 57-60.

Ustach, P.C. 1997. Geographic distribution: Sauromalus obesus (Chuckwalla). Herpetological Review 28(3): 158.

Ventura-Trejo, J.; Valdez-Villavicencio, J.H.; Hollingsworth, B.D. 2008. Geographic distribution: Sauromalus ater (Common Chuckwalla). Herpetological Review 39(2): 239-240.

Werman, S.D. 1982. Notes on the ecology of the chuckwalla, Sauromalus obesus near Baker, California. Journal of Herpetology 16(4): 417-418.

Werning, H. 1998. Erfahrungen mit der Haltung und Vermehrung von Chuckwallas (Sauromalus obesus obesus). Iguana Rundschreiben 11(2)(20): 16-21.

Wölfel, H. 2001. Die "echten Leguane" der Wüste - Chuckwallas (Sauromalus obesus) und Wüstenleguane (Dipsosaurus dorsalis). Iguana Rundschreiben 14(1): 15-22.

Wölfel, H. 2006. Chuckwallas: the genus Sauromalus: with notes on keeping and breeding Sauromalus ater. Reptilia (GB) 48: 35-42.

Sauromalus hispidus

Carl, G.; Jones, J.P. 1979. The eggs and hatchlings of Sauromalus hispidus (Reptilia, Lacertilia, Iguanidae). Journal of Herpetology 13(3): 293-296.

Case, T.J. 1982. Ecology and evolution of the insular gigantic chuckawallas, Sauromalus hispidus and Sauromalus varius. pp. 184-212. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Grismer, L.L.; Beaman, K.R.; Lawler, H.E. 1995. Sauromalus hispidus. Catalogue of American Amphibians and Reptiles 615: 1-4.

Reynoso, V.H.; Monteverde, M.J.; Martínez-González, C.; Zarza, E.; Cerdá-Ardura, A. 2017. Geographic distribution: Sauromalus hispidus (Spiny Chuckwalla). Herpetological Review 48(1): 126.

Shaw, C.E. 1946. A new locality for the spiny chuekwalla, Sauromalus hispidus. Copeia 1946(4): 254.

Smits, A.W. 1985. Behavioral and dietary responses to aridity in the chuckwalla, Sauromalus hispidus. Journal of Herpetology 19(4): 441-449.

Smits, A.W.; Ward, J.; Lillywhite, H. 1986. Effects of hyperkalemia on thermoregulatory and feeding behaviors of the lizard, Sauromalus hispidus. Copeia 1986(2): 518-520.

Sylber, C.K. 1985. Eggs and hatchlings of the yellow giant chuckwalla in captivity. Herpetological Review 16(1): 18-21.

Sylber, C.K. 1988. Feeding habits of the lizards Sauromalus varius and S. hispidus in the Gulf of California. Journal of Herpetology 22(4): 413-424.

Velarde, E.; Hollingsworth, B.D.; Rebman, J.P. 2008. Geographic distribution: Sauromalus hispidus (Spiny Chuckwalla). Herpetological Review 39(3): 368.

Werning, H. 2002. Der Stachelige Riesenchuckwalla Sauromalus hispidus Stejneger, 1877: biologie, Freilandbeobachtungen und Europäische Erstnachzucht. Salamandra 38(1): 27-44.

Sauromalus klauberi

Shaw, C.E. 1941. A new chuckwalla from Santa Catalina Island, Gulf of California, Mexico. Transactions of the San Diego Society of Natural History 9(28): 285-288.

Sauromalus varius

Case, T.J. 1982. Ecology and evolution of the insular gigantic chuckawallas, Sauromalus hispidus and Sauromalus varius. pp. 184-212. In: Burghardt, G.M. & Rand, A.S. (eds.). Iguanas of the world: their behaviour, ecology and conservation. Noyes Publications, Park Ridge, New Jersey. 472 pp.

Densmore, L.D.; Pliler, E.D.; Lawler, H.E. 1994. A molecular approach for determining genetic variation in captive and natural populations of the piebald Chuckwalla (Sauromalus varius). pp. 343-351. In: Murphy, J.B.; Adler, K. & Collins, J.T. (eds.). Captive management and conservation of amphibians and reptiles. SSAR (Society for the Study of Amphibians and Reptiles). 408 pp. (Contributions to Herpetology no. 11).

Finnley, D. (ed.) 1979. Rulemaking actions. November 1979. Five foreign reptiles proposed as endangered. Endangered Species Technical Bulletin 4(12): 8.

Finnley, D. (ed.) 1979. San Esteban Island chuckwalla under review. Endangered Species Technical Bulletin 4(3): 6.

Finnley, D. (ed.) 1980. Rulemaking actions. March 1980. Five foreign reptiles listed as endangered. Endangered Species Technical Bulletin 5(4): 13.

Gray, R.L. 1995. San Esteban Chuckwallas Sauromalus varius. An Arizona-Sonora Desert Museum delight. Vivarium (Lakeside) 6(4): 52-54.

Hollingsworth, B.D.; Mahrdt, C.R.; Grismer, L.L.; Banta, B.H.; Sylber, C.K. 1997. The occurrence of Sauromalus varius on a satellite islet of Isla Salsipuedes, Gulf of California, Mexico. Herpetological Review 28(1): 26-28.

Lawler, H.E.; Beaman, K.R.; Grismer, L.L. 1995. Sauromalus varius. Catalogue of American Amphibians and Reptiles 616: 1-4.

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