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Taxonomy of the genus Agkistrodon Biodiversity of the family Viperidae

Bibliography of the genus
Agkistrodon (American Copperheads and Moccasins)

(Reptilia: Serpentes: Viperidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Agkistrodon in general

Aldridge, R.D.; Duvall, D. 2002. Evolution of the mating season in the pitvipers of North America. Herpetological Monographs 16: 1-25.

Bushar, L.M.; Maliga, M.; Reinert, H.K. 2001. Cross-species amplification of Crotalus horridus microsatellites and their application in phylogenetic analysis. Journal of Herpetology 35(3): 532-537.

Chiasson, R.B. 1982. The apical pits of Agkistrodon (Reptilia: Serpentes). Journal of the Arizona-Nevada Academy of Science 16(3) [1981]: 69-73.

Conant, R. 1986. Phylogeny and zoogeography of the genus Agkistrodon in North America. pp. 89-92. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European herpetological meeting (3rd ordinary general meeting of the Societas Europaea Herpetologica), Prague 1985. Charles University, Prague. 754 pp.

Conant, R. 1992. Comments on the survival status of members of the Agkistrodon complex. pp. 29-33. In: Strimple, P.D. & Strimple, J.L. (eds.). Contributions in herpetology. Greater Cincinnati Herpetological Society, Cincinnati. 111 pp.

Ernst, C.H. 1965. Fang length comparisons of American Agkistrodon. Transactions of the Kentucky Academy of Science 26: 12-18.

Ernst, C.H. 1982. A study of the fangs of snakes belonging to the Agkistrodon-complex. Journal of Herpetology 16(1): 72-80.

Ernst, G.H. 1964. A study of sexual dimorphism in American Agkistrodon fang lengths. Herpetologica 20: 214.

Fernandez, P.; Gutierrez, J.M. 2008. Mortality due to snakebite envenomation in Costa Rica (1993-2006). Toxicon 52(3): 530-533.

Ferrier, W.; Ferrier, U. 1999. Agkistrodon. New World pit vipers. Reptilia (GB) 8: 58-63.

Gloyd, H.K.; Conant, R. 1943. A synopsis of the American forms of Agkistrodon (copperheads and moccasins). Bull. Chioago Acad. Sci. 7(2): 147-170.

Isogawa, K. 1995. Comparative observations on dorsal scales of shed skins of the genus Agkistrodon (Viperidae, Crotalinae) from Far East Asia. The Snake 27(1): 7-11.

Isogawa, K. 1997. Classification of the Old World Agkistrodon (Viperidae; Crotalinae) with comments on some problems in its taxa. Honyurui Kagaku 37(1): 47-53.

Klauber, L.M. 1957. Agkistrodon or Ancistrodon? Copeia 1956(4) [1957]: 258-259.

Knight, A.; Densmore, L.D.; Rael, E.D. 1992. Molecular systematics of the Agkistrodon complex. pp. 49-70. In: Campbell, J.A. & Brodie, E.D. (eds.). Biology of the pitvipers. Selva, Tyler, Texas. 467 pp.

Newman, D.J.; Lee, J.R. 2013. Natural history notes: Lampropeltis nigra (Black Kingsnake). Diet. Herpetological Review 44(3): 523.

Parker, H.W. 1966. Ancistrodon not Agkistrodon (Reptilia-Serpentes) Z.N. (S.) 671. Bulletin of Zoological Nomenclature 22: 300-302.

Rannala, B.; Yang, Z. 2017. Efficient Bayesian species tree inference under the multispecies coalescent. Systematic Biology 66(5): 823-842

Sabrosky, C.W. 1967. Ancistrodon versus Agkistrodon, Z.N. (S.) 671. Bulletin of Zoological Nomenclature 24: 203.

Savitzky, A.H. 1992. Embryonic development of the maxillary and prefrontal bones of crotaline snakes. pp. 119-141. In: Campbell, J.A. & Brodie, E.D. (eds.). Biology of the pitvipers. Selva, Tyler, Texas. 467 pp.

Smith, H.M.; Gloyd, H.K. 1964. Nomenclatural notes on the snake names Scytale, Boa scytale and Agkistrodon mokasen. Herpetologica 19: 280-282.

Toriba, M. 1999. Comments on the taxonomy of Agkistrodon listed in Stejneger (1907). (In Japanese). Bulletin of the Herpetological Society of Japan 1: 3-5.

Tu, A.T.; Adams, B.L. 1968. Phylogenetic relationships among venomous snakes of the genus Agkistrodon from Asia and the north American continent. Nature (London) 217: 760-762.

Wittner, D. 1978. A discussion of venomous snakes of North America. Distribution and occurrence of North American snakes of the genera Crotalus, Sistrurus, and Agkistrodon. Herp 14(1): 12-17.

Agkistrodon bilineatus

Allen, E.R. 1949. Observations of the feeding habits of the juvenile cantil. Copeia 1949(3): 225-226.

Arenas-Monroy, J.C.; Ahumada-Carrillo, I.T. 2015. Distribution notes: Agkistrodon bilineatus Günther, 1863 (Squamata: Viperidae): confirmation of an inland locality for central Jalisco, Mexico. Mesoamerican Herpetology 2(3): 371-374.

Babb, R.D.; Dugan, E.A. 2008. Geographic distribution: Agkistrodon bilineatus (Cantil). Herpetological Review 39(1): 110.

Bartlett, R.D. 1995. Copperheads, masters of camouflage (with mention of Cottonmouths, Cantils and Gaboon Vipers). Tropical Fish Hobbyist 44(4): 152, 154, 156, 158, 160-163, 165-168, 170-171.

Blair, K.B.; Killebrew, F.C.; Smith, H.M.; Chiszar, D. 1997. Mexican amphibians and reptiles in the West Texas A & M University Natural History Museum. Bulletin of the Chicago Herpetological Society 32(8): 174-177.

Boerema, H. 1989. The keeping and breeding of Agkistrodon bilineatus bilineatus. Litteratura Serpentium (English Edition) 9(6): 267-268.

Bolanos, R. 1968. Agkistrodon bilineatus Gunther from Costa Rica. Revista de Biologia Tropical 16: 277-278.

Bryson, R.W.; Mendoza-Quijano, F. 2007. Cantils of Hidalgo and Veracruz, Mexico, with comments on the validity of Agkistrodon bilineatus lemosespinali. Journal of Herpetology 41(3): 536-539.

Canseco-Marquez, L.; Nolasco-Velez, A.L. 2008. Geographic distribution: Agkistrodon bilineatus (Cantil). Herpetological Review 39(3): 369.

Cruz, G.A.; Wilson, L.D.; Espinosa, J. 1979. Two additions to the reptile fauna of Honduras, Eumeces managuae Dunn and Agkistrodon bilineatus (Gunther), with comment on Pelamis platurus (Linnaeus). Herpetological Review 10(1): 26-27.

DeSantis, D.L.; Mata-Silva, V.; García-Padilla, E.; Wilson, L.D.; Ramírez-Bautista, A. 2015. Distribution notes: Agkistrodon bilineatus (Günther, 1863). Mesoamerican Herpetology 2(4): 552-553.

Drent, J. 1991. Agkistrodon bilineatus bilineatus. Litteratura Serpentium (English Edition) 11(6): 145.

Dugan, E.A.; Meyer, S.A. 2007. Natural history notes: Agkistrodon bilineatus (Cantil). Reproduction. Herpetological Review 38(1): 85-86.

Furiani, M. 1989. The reproduction of Agkistrodon bilineatus (Gunther, 1888) in captivity. Litteratura Serpentium (English Edition) 9(4): 142-144.

García-Grajales, J.; Buenrostro-Silva, A. 2011. Ampliación del área de distribución gepgráfica de Oxybelis fulgidus (Serpentes: Colubridae) y Agkistrodon bilineatus (Serpentes: Viperidae) en la planicie costera central de Oaxaca, México [Geographic distribution enlargment of Oxybelis fulgidus (Serpentes: Colubridae) and Agkistrodon bilineatus (Serpentes: Viperidae) from the costal central plain of Oaxaca, Mexico]. (In Spanish). Acta Zoologica Mexicana Nueva Serie 27(2): 491-495.

Gloyd, H.K. 1957. Distribution of the Mexican moccasin: a correction. Copeia 1956(4) [1957]: 259.

Henderson, R.W. 1978. Notes on Agkistrodon bilineatus (Reptilia, Serpentes, Viperidae) in Belize. Journal of Herpetology 12(3): 412-413.

Lemos-Espinal, J.A.; Smith, H.M.; Chiszar, D. 2006. Agkistrodon b. bilineatus in Western Chihuahua. Bulletin of the Maryland Herpetological Society 42(4): 173-175.

Loc-Barragán, J.A.; Carbajal-Márquez, R.A. 2016. Nature notes: Agkistrodon bilineatus Günther, 1863. Diet. Mesoamerican Herpetology 3(3): 733-735.

Loveridge, A. 1928. Note on Agkistrodon bilineatus Gunther. Bulletin of the Antivenin Institute of America 2: 52.

Macchiavelli, G. 1990. Reproduction of Agkistrodon bilineatus bilineatus. Litteratura Serpentium (English Edition) 10(5): 221-224.

Neill, W.T. 1960. The caudal lure of various juvenile snakes. Quarterly Journal of the Florida Academy of Sciences 23: 173-200.

Porras, L.W.; Wilson, L.D.; Schuett, G.W.; Reiserer, R.S. 2013. A taxonomic reevaluation and conservation assessment of the common cantil, Agkistrodon bilineatus (Squamata: Viperidae): a race against time. Amphibian and Reptile Conservation 7(1): 48-73.

Smetsers, P. 1993. Agkistrodon bilineatus - Gunther, the tropical moccasin. Litteratura Serpentium (English Edition) 13(2): 38-40.

Smith, H.M.; Chiszar, D. 2001. A new subspecies of Cantil (Agkistrodon bilineatus) from central Veracruz, Mexico (Reptilia: Serpentes). Bulletin of the Maryland Herpetological Society 37(4): 130-136.

Strimple, P. 1992. Caudal movements of an adult Mexican cantil, Agkistrodon bilineatus bilineatus Gunther. pp. 55-56. In: Strimple, P.D. & Strimple, J.L. (eds.). Contributions in herpetology. Greater Cincinnati Herpetological Society, Cincinnati. 111 pp.

Strimple, P. 1995. Comments on caudal luring in snakes with observations on this behaviour in two subspecies of cantils, Agkistrodon bilineatus ssp. Litteratura Serpentium (English Edition) 15(3): 74-77.

Trutnau, L. 2001. Einige Bemerkungen zur Biologie, Pflege und Nachzucht der mexikanischen Giftschlangenart Agkistrodon bilineatus Günther, 1863. Herpetofauna (Weinstadt) 23(131): 5-14.

Zweifel, R.G. 1959. The provenance of reptiles and amphibians collected in western Mexico by J.J. Major. American Museum Novitates 1949: 1-9.

Agkistrodon conanti

Allen, E.R.; Swindell, D. 1948. Cottonmouth Moccasin of Florida. Herpetologica 4: 1-15.

Enge, K.M. 2013. Geographic distribution: Agkistrodon piscivorus conanti (Florida Cottonmouth). Herpetological Review 44(3): 475.

Enge, K.M.; Mays, J.D.; Harrison, B.B. 2016. Geographic distribution: Agkistrodon piscivorus conanti (Florida Cottonmouth). Herpetological Review 47(1): 82.

Friers, J.; Flaherty, J.P. 2015. Natural history notes: Agkistrodon piscivorus conanti (Florida cottonmouth). Diet. Herpetological Review 46(4): 636-637.

Gloyd, H.K. 1968. Two additional subspecies of North American crotalid snakes, genus Agkistrodon. Proceedings of the Biological Society of Washington 82: 219-232.

Grajal-Puche, A.; Josimovich, J.; Falk, B.; Reed, R.N. 2016. Natural history notes: Agkistrodon piscivorus conanti (Florida Cottonmouth). Diet. Herpetological Review 47(2): 307.

Hecht-Kardasz, K.A.; Drayer, A.N.; Himes, J.G. 2012. Geographic distribution: Agkistrodon piscivorus (Cottonmouth). USA: Florida: Bay Co. Herpetological Review 43(3): 446-447.

Heinrich, G.; Studenroth, K.R. 1996. Natural history notes: Agkistrodon piscivorus conanti (Florida Cottonmouth). Diet. Herpetological Review 27(1): 22.

Lillywhite, H.B.; Brischoux, F. 2012. Is it better in the moonlight? Nocturnal activity of insular Cottonmouth snakes increases with lunar light levels. Journal of Zoology (London) 286(3): 194-199.

Lillywhite, H.B.; McCleary, R.J.R. 2008. Trophic ecology of insular Cottonmouth snakes: review and perspective. South American Journal of Herpetology 3(2): 175-185.

Lillywhite, H.B.; Sheehy, C.M.; McCue, M.D. 2002. Scavenging behaviors of Cottonmouth snakes at island bird rookeries. Herpetological Review 33(4): 259-261.

Lillywhite, H.B.; Sheehy, C.M.; Zaidan, F. 2008. Pitviper scavenging at the intertidal zone: an evolutionary scenario for invasion of the sea. Bioscience 58(10): 947-955.

May, P.G.; Farrell, T.M. 1998. Florida's flatwoods. Cottonmouths: observations of feeding behavior. Reptile & Amphibian Magazine 56: 18-24.

Palis, J.G. 1993. Life history notes: Agkistrodon piscivorus conanti (Florida Cottonmouth). Prey. Herpetological Review 24(2): 59, 62.

Sheehy, C.M.; Sandfoss, M.R.; Lillywhite, H.B. 2017. Cannibalism and changing food resources in insular cottonmouth snakes. Herpetological Review 48(2): 310-312.

Studenroth, K.R. 1991. Life history notes: Agkistrodon piscivorus conanti (Florida Cottonmouth). Foraging behavior. Herpetological Review 22(2): 60.

Tetzlaff, S.J.; Tetzlaff, D.L.; Schoenbeck, M.; Clark, N. 2014. Natural history notes: Agkistrodon piscivorus conanti (Florida Cottonmouth). Diet and phenotype. Herpetological Review 45(3): 511-512.

Walters, A.C.; Card, W. 1996. Natural history notes: Agkistrodon piscivorus conanti (Florida Cottonmouth). Brood defense. Herpetological Review 27(4): 203.

Wharton, C.H. 1966. Reproduction and growth in the cottonmouths, Agkistrodon piscivorus Lacepede, of Cedar Keys, Florida. Copeia 1966: 149-161.

Wharton, C.H. 1969. The Cottonmouth moccasin on Sea Horse Key, Florida. Bulletin of the Florida State Museum Biological Sciences 14: 227-272.

Young, B.A.; Aguiar, A.; Lillywhite, H.B. 2008. Foraging cues used by insular Florida Cottonmouths, Agkistrodon piscivorus conanti. South American Journal of Herpetology 3(2): 135-144.

Agkistrodon contortrix

Abbuhl, B. 2008. Natural history notes: Agkistrodon contortrix mokeson (Northern Copperhead). Mid-winter thermoregulation. Herpetological Review 39(4): 465.

Allen, E.R. 1938. The Copperhead in northern Florida. Copeia 1938: 50.

Allen, E.R.; Neill, W.T. 1957. The copperhead. Florida Wild Life 1950 [1957]: 18-19.

Arndt, R.G. 1971. The copperhead in New York. Conservationist 25(5): 18-19.

Arndt, R.G. 1985. Distributional notes on some Delaware snakes, with two first records for the state. Bulletin of the Maryland Herpetological Society 21(2): 67-73.

Babcock, H.L. 1926. The copperhead. Bull. Boston Soc. Nat. Hist. 41: 3-6.

Bailey, R.M. 1940. The northern copperhead in Iowa. Iowa State Coll. J. Sc. 15(1): 1-2.

Barbour, R.W. 1962. An aggregation of copperheads, Agkistrodon contortrix. Copeia 1962: 640.

Barrass, G.T.; Barrass, A.N. 2011. Geographic distribution: Agkistrodon contortrix (Copperhead). Herpetological Review 42(2): 242.

Bartlett, R.D. 1995. Copperheads, masters of camouflage (with mention of Cottonmouths, Cantils and Gaboon Vipers). Tropical Fish Hobbyist 44(4): 152, 154, 156, 158, 160-163, 165-168, 170-171.

Barton, A.J. 1949. Ophiophagy by a juvenile copperhead. Copeia 1949(3): 232.

Beane, J.C.; Gerwin, J.A. 2013. Natural history notes: Agkistrodon contortrix (Copperhead). Diet and accidental mortality. Herpetological Review 44(1): 150-151.

Beaupre, S.J.; Roberts, K.G. 2001. Natural history notes: Agkistrodon contortrix contortrix (Southern Copperhead). Chemotaxis, arboreality and diet. Herpetological Review 32(1): 44-45.

Bender, M.J.; Duffus, A.L.J. 2014. Geographic distribution: Agkistrodon contortrix (Copperhead). Herpetological Review 45(1): 93.

Boerema, H. 1990. Agkistrodon contortrix mukeson. Litteratura Serpentium (English Edition) 10(3): 148.

Boerema, H. 1990. Breeding results. Agkistrodon contortix contortix. Litteratura Serpentium (English Edition) 10(1): 54.

Boerema, H. 1991. Agkistrodon contortrix contortrix - Copperhead. Litteratura Serpentium (English Edition) 11(5): 114.

Boerema, H. 1991. Agkistrodon contortrix mokeson - Copperhead. Litteratura Serpentium (English Edition) 11(5): 114.

Booth, W.; Schuett, G.W. 2011. Molecular genetic evidence for alternative reproductive strategies in North American pitvipers (Serpentes: Viperidae): long-term sperm storage and facultative parthenogenesis. Biological Journal of the Linnean Society 104(4): 934-942.

Booth, W.; Smith, C.F.; Eskridge, P.H.; Hoss, S.K.; Mendelson, J.R.; Schuett, G.W. 2012. Facultative parthenogenesis discovered in wild vertebrates. Biology Letters 8(6): 983-985.

Boyer, D.A. 1933. A case report on the potency of the bite of a young copperhead. Copeia 1933: 97.

Brooks, G.C.; Graham, S.P. 2012. Geographic distribution: Agkistrodon contortrix contortrix (Southern Copperhead). USA: Alabama: Covington Co. Herpetological Review 43(4): 620.

Browder, R. 2005. Agkistrodon contortrix mokasen (Northern Copperhead). Catesbeiana 25(1): 31-32.

Broxson, M.D. 2011. Geographic distribution: Agkistrodon contortrix (Copperhead). Herpetological Review 42(1): 113.

Brumwell, M.J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. American Midland Naturalist 45(1): 187-231.

Burbrink, F.T.; Guiher, T.J. 2015. Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus Agkistrodon. Zoological Journal of the Linnean Society 173(2): 505-526.

Calmonte, A. 1973. Nördlicher Kupferkopf, Agkistrodon contortrix mokeson (Daudin, 1803). Aquarium Aqua Terra 7(54): 508-510.

Cameron, M.L.; Barclay, S.F.; McAllister, C.T. 2004. Geographic distribution: Agkistrodon contortrix (Copperhead). Herpetological Review 35(4): 409.

Carpenter, C.C. 1958. Reproduction, young, eggs and food of Oklahoma snakes. Herpetologica 14: 113-115.

Carr, W.H. 1926. The fasting of a copperhead. Copeia 1926(150): 104.

Carter, E.T.; Attum, O.; Eads, B.C.; Hoffman, A.S.; Kingsbury, B.A. 2014. Reducing the potential for human-snake encounters in a recreational park. Human-Wildlife Interactions 8(2): 158-167.

Cassano, E.; Smith, C.; Eskridge, P. 2012. Natural history notes: Agkistrodon contortrix contortrix (Southern Copperhead). Maximum captive length. Herpetological Review 43(2): 265.

Chenoweth, W.L. 1948. Birth and behavior of young copperheads. Herpetologica 4(5): 162.

Cochran, D.M. 1944. Dangerous reptiles (popular). Rep. Smithson. Instn. 1943: 275-325.

Cochran, P.A. 2008. A Cottonmouth (Agkistrodon piscivorus) in Minnesota, and historical reports of other pit vipers unexpected in the upper Midwest. Northeastern Naturalist 15(3): 461-464.

Collins, J.D.; Collins, J.T. 2003. Agkistrodon contortrix (Copperhead). Journal of Kansas Herpetology 8: 19.

Colvin, R.; Dennison, J. 2013. Geographic distribution: Agkistrodon contortrix (Copperhead). USA: Tennessee: Weakley Co. Herpetological Review 44(1): 108.

Connior, M.B. 2008. Geographic distribution: Agkistrodon contortrix contortrix (Southern Copperhead). USA: Arkansas. Herpetological Review 39(4): 484.

Cross, C.L.; Petersen, C.E. 2001. Modeling snake microhabitat from radiotelemetry studies using polytomous logistic regression. Journal of Herpetology 35(4): 590-597.

Cupp, P.V. 2018. Natural history notes: Agkistrodon contortrix (Copperhead). Diet / ophidiophagy. Herpetological Review 49(4): 747.

Dolley, J.S. 1939. An anomalous pregnancy in the copperhead. Copeia 1939: 170.

Douglas, M.E.; Douglas, M.R.; Schuett, G.W.; Porras, L.W. 2009. Climate change and evolution of the New World pitviper genus Agkistrodon (Viperidae). Journal of Biogeography 36(6): 1164-1180.

Dowling, H.G. 1956. Geographic relations of Ozarkian amphibians and reptiles. Southwestern Naturalist 1(4): 174-189.

Dowling, H.G. 1959. Rattles and hisses and rasping scales. Animal Kingdom 62: 140-142.

Durso, A.M. 2017. Natural history notes: Agkistrodon contortrix (Copperhead). Diet. Herpetological Review 48(1): 203-204.

Durso, A.M.; Morris, K.A. 2015. Natural history notes: Agkistrodon contortrix (copperhead). Diet. Herpetological Review 46(4): 636.

Dyke, J.U. van; Grace, M.S. 2010. The role of thermal contrast in infrared-based defensive targeting by the Copperhead, Agkistrodon contortrix. Animal Behaviour 79(5): 993-999.

Ferner, J.W.; Bramlage, M.; Bryant, W.S. 1999. Geographic distribution: Agkistrodon contortrix mokasen (Northern Copperhead). Herpetological Review 30(2): 112.

Finneran, L.C. 1953. Aggregation behavior of the female copperhead, Agkistrodon contortrix mokeson, during gestation. Copeia 1953: 61-62.

Fitch, H.S. 1959. A patternless phase of the copperhead. Herpetologica 15: 21-24.

Fitch, H.S. 1960. Autecology of the copperhead. University of Kansas Publications of the Museum of Natural History 13: 85-288.

Fitch, H.S. 2004. Observations on Osage Copperheads in northeastern Kansas. Journal of Kansas Herpetology 12: 20.

Fitch, H.S. 2004. The effect of female size on number of eggs or young in snakes. Journal of Kansas Herpetology 9: 11-12.

Fitch, H.S. 2005. Observations on wandering of juvenile snakes in northeastern Kansas. Journal of Kansas Herpetology 13: 11-12.

Fitch, H.S.; Clarke, A.L. 2002. An exceptionally large natural assemblage of female Copperheads (Agkistrodon contortrix). Herpetological Review 33(2): 94-95.

Flaherty, J.P.; Fulbright, M.C.; Settle, S.P.; Ross, A.T.; Gienger, C.M. 2014. Geographic distribution: Agkistrodon contortrix (Copperhead). Herpetological Review 45(1): 93.

Fogell, D.D.; Leonard, T.J.; Fawcett, J.D. 2002. Natural history notes: Agkistrodon contortrix phaeogaster (Osage Copperhead). Breeding. Herpetological Review 33(3): 209-210.

Folt, B.P.; Laurencio, D. 2013. Natural history notes: Agkistrodon contortrix (Copperhead). Diet. Herpetological Review 44(3): 517.

Freeman, H.W. 1955. Part V. The amphibia and reptilia of the Savannah river project area. 2. Chelonia. 3. Crocodilia, Sauria and Serpentes. U.S. Cur. Publ. Biol. 1(4): 239-244, 275-291.

Garton, J.S.; Dimmick, R.W. 1969. Food habits of the copperhead in middle Tennessee. Journal of the Tennessee Academy of Science 44: 113-117.

Gibson, J.D. 2003. Agkistrodon contortrix mokasen (Northern Copperhead). Catesbeiana 23(2): 66.

Gloyd, H.K. 1934. Studies on the breeding habits and young of the copperhead, Agkistrodon mokasen Beauvois. Papers of the Michigan Academy of Science, Arts and Letters 19(33): 587-604.

Gloyd, H.K. 1968. Two additional subspecies of North American crotalid snakes, genus Agkistrodon. Proceedings of the Biological Society of Washington 82: 219-232.

Gloyd, H.K.; Conant, R. 1938. The subspecies of the copperhead Agkistrodon mokasen Beauvois. Bull. Acad. Sci. Chicago 5(7): 163-166.

Godard, R.D.; Wilson, C.M.; Rock, C.M.; Cash, W.B. 2006. Natural history notes: Agkistrodon contortrix (Copperhead). Diet. Herpetological Review 37(4): 476.

Goodman, J.D. 1948. A report on the reptiles collected by J.M. Shaffer from the Keokuk Area, 1863-1895. Proceedings of the Iowa Academy of Science 55: 365-366.

Goodwin, O.K.; Wood, J.T. 1956. Distribution of poisonous snakes on the York-James Peninsula. A zoogeographic mystery. Va. J. Sci. N.S. 7(1): 17-21.

Govorchin, R.E. 1965. America's poisonous snakes. Frontiers 29: 70-73.

Graves, G.R. 2002. Copperhead preys on star-nosed mole in the Great Dismal Swamp. Banisteria 20: 70.

Greding, E.J. 1964. Food of Ancistrodon c. contortrix in Houston and Trinity counties, Texas. Southwestern Naturalist 9: 105.

Greenbaum, E. 2004. The influence of prey-scent stimuli on predatory behavior of the North American Copperhead Agkistrodon contortrix (Serpentes: Viperidae). Behavioral Ecology 15(2): 345-350.

Greenbaum, E.; Jorgensen, M. 2004. Envenomated-invertebrate prey preference of the viperid Agkistrodon contortrix during strike-induced chemosensory searching. Amphibia-Reptilia 25(2): 165-172.

Grogan, W.L. 1994. New herpetological distribution records from Maryland's eastern shore. Bulletin of the Maryland Herpetological Society 30(1): 27-32.

Gross, I.P.; Ewers, C.M.; Wang, Y.; Schweitzer, C.J. 2016. Natural history notes: Agkistrodon contortrix (Copperhead). Tunnel-blocking behavior. Herpetological Review 47(2): 306-307.

Groves, J.D. 1977. Aquatic behavior in the northern copperhead, Agkistrodon contortrix mokasen. Bulletin of the Maryland Herpetological Society 13(2): 114-115.

Guiher, T.J.; Burbrink, F.T. 2008. Demographic and phylogeographic histories of two venomous North American snakes of the genus Agkistrodon. Molecular Phylogenetics and Evolution 48(2): 543-553.

Hamilton, W.J.; Pollack, J.A. 1955. The food of some crotalid snakes from Fort Benning, Georgia. Natural History Miscellanea 140: 1-4.

Heatwole, H.; King, P.; Levine, S.G. 2007. Laterality in coiling behaviour of snakes: another interpretation. Laterality 12(6): 536-542.

Heatwole, H.; Poran, N.; King, P. 1999. Ontogenetic changes in the resistance of Bullfrogs (Rana catesbeiana) to the venom of Copperheads (Agkistrodon contortrix contortrix) and Cottonmouths (Agkistrodon piscivorus piscivorus). Copeia 1999(3): 808-814.

Irwin, K.J.; Irwin, L.K. 2002. Geographic distribution: Agkistrodon contortrix (Copperhead). Herpetological Review 33(2): 149.

Jensen, J.B. 1999. Natural history notes: Terrapene carolina carolina (Eastern Box Turtle). Diet. Herpetological Review 30(2): 95.

Johnson, J.E. 1948. Copperhead in a tree. Herpetologica 4(6): 214.

Johnston, B.L. 2004. Geographic distribution: Agkistrodon contortrix (Copperhead). Herpetological Review 35(1): 82.

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Agkistrodon howardgloydi

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Agkistrodon laticinctus

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Agkistrodon piscivorus

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Cypert, E. 1961. The effect of fires in the Okefenokee Swamp in 1954 and 1955. American Midland Naturalist 66: 485-503.

Davis, B.; Birkhead, R.D. 2013. Natural history notes: Terrapene carolina (Eastern Box Turtle). Diet. Herpetological Review 44(4): 659.

Davis, J.B. 2002. Natural history notes: Agkistrodon piscivorus (Cottonmouth). Predation. Herpetological Review 33(2): 136-137.

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Dowling, H.G. 1959. Rattles and hisses and rasping scales. Animal Kingdom 62: 140-142.

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Dunson, W.A.; Freda, J. 1985. Water permeability of the skin of the amphibious snake, Agkistrodon piscivorus. Journal of Herpetology 19(1): 93-98.

Durso, A.M.; Willson, J.D.; Winne, C.T. 2011. Needles in haystacks: estimating detection probability and occupancy of rare and cryptic snakes. Biological Conservation 144(5): 1508-1515.

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