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Pacific Ocean
Ecuador

Reptiles
Lizards
Iguanidae
Phyllodactylidae
Tropiduridae
Snakes
Colubridae
Elapidae
[marine species]

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Bibliography of the reptiles of
Galapagos Islands  [Ecuador]

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Anonymous. 1970. Tortoises v. rats and goats in the Galapagos. Oryx 10: 298-301.

Anonymous. 1973. Cambridge and London Galapagos expedition 1972. Cambridge Exped. J. 1973: 25-31.

Anonymous. 1974. Cambridge and London Galapagos expedition 1973. Cambridge Exped. J. 1974: 3-4.

Anonymous. 1986. News from Academy Bay. Noticias de Galapagos 43: 2-9.

Agustin, H. 1958. Dos Iguanas de Galapagoes. Div. Cient. Oc. Mus. Ecuador, Guayaquil. 4 pp.

Ali, J.R.; Aitchison, J.C. 2014. Exploring the combined role of eustasy and oceanic island thermal subsidence in shaping biodiversity on the Galapagos. Journal of Biogeography 41(7): 1227-1241.

Bailey, A.M. 1970. Narrative of the 1960 field trip of the Denver Museum of Natural History to the Galapagos Islands. The enchanted isles of Ecuador. Museum Pictor. Denver Mus. Nat. Hist. 19: 1-86.

Barlow, T. 1969. Galapagos Group. No. 155. Rare Galapagos species-status survey throughout the Galapagos Islands. Yearbook World Wildlife Fund 1968: 159-160, 242-243.

Barnett, L. 1958-1959. The enchanted Isles, Darwin's World of Nature. Life (Sept. 1958) (Part II); Life (Jan. 1959) (Part IV), Life (March 1959) (Part V); Life (June 1959) (Part VI); Life (July 1959) (Part VII).

Beebe, W. 1923. The Williams Galapagos Expedition. Bulletin of the New York Zoological Society 26: 51-61.

Beebe, W. 1924. Galapagos: World's End. New York & London. 433 pp.

Beebe, W. 1926. The Arcturus adventure. G.P. Putnam's Sons, New York & London. Unpaginated.

Bergen, T. 2013. Decline and rise of Galápagos tortoises. IRCF Reptiles & Amphibians 20(1): 23–29.

Bisconti, M.; Landini, W.; Bianucci, G.; Cantalamessa, G.; Carnevale, G.; Ragaini, L.; Valleri, G. 2001. Biogeographic relationships of the Galapagos terrestrial biota: Parsimony analyses of endemicity based on reptiles, land birds and Scalesia land plants. Journal of Biogeography 28(4): 495-510.

Blomberg, R. 1951. Strange reptiles of the Galapagos. Natural History (New York) 60: 234-238.

Brosset, A. 1963. Statut actuel de la faune aux iles Galapagos. Noticias de Galápagos 1: 5-8.

Calgua, A. 1993. Die Leguane der Galapagos-Inseln. Iguana Rundschreiben 6(1): 9-11.

Carpenter, C.C. 1964. Comparative behaviour of the lava lizards (Tropidurus) of the Galapagos Islands. American Zoologist 4: 274.

Carpenter, C.C. 1967. Some findings on Galapagos lizards. Noticias de Galapagos 9/10: 18-20.

Carrasco, M.A. 1998. Una expedicion cientifica multidisciplinar a las Islas Galapagos. Boletin de la Real Sociedad Espanola de Historia Natural Actas 95: 51-55.

Chase, I.R.S.; Hailman, J.P. 1963. Two views on conservation in the Galapagos. Bulletin of the Philadelphia Herpetological Society 11(3-4): 71-74.

Chiari, Y.; Hyseni, C.; Fritts, T.H.; Glaberman, S.; Marquez, C.; Gibbs, J.P.; Claude, J.; Caccone, A. 2009. Morphometrics parallel genetics in a newly discovered and endangered taxon of Galapagos tortoise. PLoS ONE 4(7): e6272, 1-11.

Christian, K.A. 1980. Cleaning/feeding symbiosis between birds and reptiles of the Galapagos Islands: new observations of inter-island variability. Auk 97(4): 887-889.

Ciofi, C.; Milinkovitch, M.C.; Gibbs, J.P.; Caccone, A.; Powell, J.R. 2002. Microsatellite analysis of genetic divergence among populations of giant Galapagos tortoises. Molecular Ecology 11(11): 2265-2283.

Denburgh, J. van 1912. Expedition of the California Academy of Sciences to the Galapagos Islands, 1905-06. 4. The snakes of the Galapagos Islands. Proceedings of the California Academy of Sciences (Ser. 4) 1: 323-374.

Denburgh, J. van 1912. Expedition of the California Academy of Sciences to the Galapagos Islands, 1905-06. 6. The geckos of the Galapagos Archipelago. Proceedings of the California Academy of Sciences (Ser. 4) 1: 405-430.

Denburgh, J. van; Slevin, J.R. 1913. Expedition of the California Academy of Sciences to the Galapagos Islands, 1905-06. 9. The Galapagoan lizards of the genus Tropidurus; with notes of the iguanas of the genera Conolophus and Amblyrhynchus. Proceedings of the California Academy of Sciences 2: 133-202.

Denburgh, J. van; Slevin, J.R. 1913. The Galapagoan lizards of the genus Tropidurus; with notes on the iguanas of the genera Conolophus and Amblyrhynchus. (Expedition of the California academy of Sciences to the Galapagos islands, 1905-1906, 9.). Proceedings of the California Academy of Sciences 2: 132-202.

Denkinger, J.; Parra, M.; Muńoz, J.P.; Carrasco, C.; Murillo, J.C.; Espinosa, E.; Rubianes, F.; Koch, V. 2013. Are boat strikes a threat to sea turtles in the Galapagos Marine Reserve. Ocean & Coastal Management 80: 29-35.

Dorst, J. 1959. Les îles Galapagos et leur monde étrange. 1. Les îles - Flore et faune - Les Reptiles. Nature (Paris) 3291: 289-296.

Dorst, J. 1961. The fate of wildlife in the Galapagos Islands. Oryx 6(1): 53-59.

Dowling, H.G. 1962. World's end revisited. The Zoological Society's Fifth Expedition to the Galapagos Islands. Animal Kingdom 65: 130-142.

Dowling, H.G. 1964. The case of the vanishing iguanas. Nat. Sci. N.Y. 2: 3-5.

Duffy, D.C. 1982. Seasnakes in the Galapagos. Noticias de Galapagos 35: 23.

Duffy, D.C. 1989. Galapagos gourmets and gourmands. Explorers Journal 67(4): 176-179.

Eckhardt, R.C. 1972. Introduced plants and animals in the Galápagos Islands. Bioscience 22(10): 585-590.

Edgar, G.J.; Banks, S.A.; Brandt, M.; Bustamante, R.H.; Chiriboga, A.; Earle, S.A.; Garske, L.E.; Glynn, P.W.; Grove, J.S.; Henderson, S.; Hickman, C.P.; Miller, K.A.; Rivera, F. 2010. El Nino, grazers and fisheries interact to greatly elevate extinction risk for Galapagos marine species. Global Change Biology 16(10): 2876-2890.

Eibl-Eibesfeldt, I. 1957. Bericht von einer Reise zu den Galapagos-Inseln unter besonderer Berücksichtigung verhaltenskundlicher, herpetologischer und ichthyologischer Beobachtungen. 3. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 10(3): 68-73.

Eibl-Eibesfeldt, I. 1960. Naturschutzprobleme auf den Galapagos-Inseln. Acta Tropica (Basel) 17(2): 97-137.

Eibl-Eibesfeldt, I. 1973. Galapagos. (Part 8). Eesti Loodus 16(4): 250-254.

Fowler de Neira, L.E.; Johnson, M.K. 1985. Diets of giant tortoises and feral burros on Volcan Alcedo, Galapagos. Journal of Wildlife Management 49(1): 165-169.

Fowler de Neira, L.E.; Roe, J.H. 1984. Emergence success of tortoise nests and effect of feral burros on nest success on Volcan Alcedo, Galapagos. Copeia 1984(3): 702-707.

Fowler, L. 1981. Feeding and population ecologies of the burros and tortoises of Volcan Alcedo, Isabela Island. Charles Darwin Research Station Annual Report 1980 [1981]: 112-114.

Fritts, T.H. 1981. Marine turtles of the Galapagos Islands and adjacent areas of the eastern Pacific on the basis of observations by J.R. Slevin 1905-1906. Journal of Herpetology 15(3): 293-301.

Fritts, T.H. 1984. Evolutionary divergence of giant tortoises in Galapagos. Biological Journal of the Linnean Society 21(1-2): 165-176.

Fritts, T.H.; Fritts, P.R. 1982. Race with extinction. Herpetological notes of J.R. Slevin's journey to the Galapagos 1905-1906. Herpetological Monographs 1: 1-98.

Garman, S. 1892. The reptiles of the Galapagos Islands. From the collections of Dr. Geo. Baur. Bull. Essex Inst. 24: 73-87.

Garrick, R.C.; Kajdacsi, B.; Russello, M.A.; Benavides, E.; Hyseni, C.; Gibbs, J.P.; Tapia, W.; Caccone, A. 2015. Naturally rare versus newly rare: demographic inferences on two timescales inform conservation of Galapagos giant tortoises. Ecology and Evolution 5(3): 676-694.

Gentile, G.; Ciambotta, M.; Tapia, W. 2013. Illegal wildlife trade in Galapagos: molecular tools help the taxonomic identification of confiscated iguanas and guide their rapid repatriation. Conservation Genetics Resources 5(3): 867-872.

Goding, F.W. 1922. Estudio sobre los reptiles encontrados en las Islas de Galapagos (Ecuador). Rev. Col. Nac. Vic. Roca Guayaquil 4(7): 79-90.

González-Pérez, F.; Cubero-Pardo, P. 2010. Efecto de actividades turisticas sobre el comportamiento de fauna representativa de las Islas Galapagos, Ecuador. Latin American Journal of Aquatic Research 38(3): 493-500.

Green, D. 1978. The east Pacific Green Sea Turtle in Galapagos. Noticias de Galápagos 28: 9-12.

Green, D. 1980. Population ecology of the east Pacific Green Turtle, Chelonia mydas agassizi, in the Galapagos Islands. Charles Darwin Research Station Annual Report 1979: 82-85.

Green, D. 1981. The Green Sea Turtle project in Galapagos: past, present & future. Noticias de Galapagos 33: 17-20.

Green, D. 1983. Galapagos Sea Turtles. Noticias de Galapagos 38: 22-25.

Green, D. 1984. Long-distance movements of Galapagos Green Turtles. Journal of Herpetology 18(2): 121-130.

Green, D. 1994. Galapagos Sea Turtles: an overview. NOAA Technical Memorandum NMFS-SEFSC 341: 65-68.

Green, D. 2003. Movements of Green Turtles within and without the Galapagos Archipelago, Ecuador. NOAA Technical Memorandum NMFS-SEFSC 503: 74-75.

Heleno, R.H.; Olesen, J.M.; Nogales, M.; Vargas, P.; Traveset, A. 2013. Seed dispersal networks in the Galapagos and the consequences of alien plant invasions. Proceedings of the Royal Society Biological Sciences Series B 280(1750): 1-9.

Heller, E. 1903. Papers from the Hopkins Stanford Galapagos Expedition, 1898-1899. XIV. Reptiles. Proceedings Washington Ac. 5: 39-98.

Henriksson, A.; Lundahl, G. 1982. [The Galapagos Islands - an island world for opportunistic animals]. (In Swedish). Fauna och Flora (Stockholm) 77(6): 315-320.

Herre, W. 1972. Galapagos-Archipel Laboratorium der Natur. Zeitschrift des Kölner Zoo 15(1): 17-25.

Hickin, N. 1979. Animal life of the Galapagos: an illustrated guide for visitors. Ferendune/Springwood, Faringdon, Oxon, England. 236 pp.

Highfield, A. 1991. Reptiles of the Galapagos Islands. Rephiberary 159: 3-4.

Honegger, R.E. 1964. Die letzten Riesen von Galapagos. 1. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 17: 275-278.

Honegger, R.E. 1964. Die letzten Riesen von Galapagos. 2. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 17: 309-312.

Honegger, R.E. 1972. Die Reptilien-Bestände auf den Galapagos-Inseln 1972. Natur und Museum (Frankfurt Am Main) 102(12): 437-454.

Houvenaghel, G.T.; Houvenaghel, N. 1974. Aspects écologiques de la zonation intertidale sur les côtes rocheuses des Îles Galapagos. Marine Biol., Berlin 26(2): 135-152.

Hurtado, G.M. 1980. Ecological study of the Green Turtle Chelonia mydas agassizi in the Galapagos Islands. Charles Darwin Research Station Annual Report 1979: 86-92.

Kramer, P. 1973. Wildlife conservation in the Galapagos Islands (Ecuador). Nature Resour. 9(4): 3-10.

Kramer, P. 1984. Man and other introduced organisms. Biological Journal of the Linnean Society 21(1-2): 253-258.

Krefft, G. 1953. Herpetologische Eindrücke einer Walfangreise. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 6: 100-103.

Kruuk, H. 1980. Ecology and control of feral dogs in Galapagos. Charles Darwin Research Station Annual Report 1979: 100-108.

Kutschera, U.; Kleinhans, S. 2013. Alfred Russel Wallace and the destruction of island life: the Iguana tragedy. Theory in Biosciences 342(4): 259-265.

Lanza, B. 1974. Le Isole Galapagos. Con la Spedizione Mares-G.R.S.T.S. all 'Arcipelago di Colombo'. 1. Universo (Florence) 54(3-4): 509-584.

Lanza, B. 1974. Le Isole Galapagos. Con la Spedizione Mares-G.R.S.T.S. all 'Arcipelago di Colombo'. 2. Universo (Florence) 54(6): 817-918.

Lanza, B. 1982. Notes on the natural history of some minor Galapagoan Islands. pp. 75-115. Museo Zoologico dell'Universita di Firenze, Galapagos. Studi e ricerche. Spedizione 'L. Mares - G.R.S.T.S.' Gruppo Ricerche Scientifische e Techniche Subacquee. 426 pp.

Lanza, B. 1982. On some Phyllodactylus from the Galapagos Islands (Reptilia Gekkonidae). pp. 149-182. Museo Zoologico dell'Universita di Firenze, Galapagos. Studi e ricerche. Spedizione 'L. Mares - G.R.S.T.S.' Gruppo Ricerche Scientifische e Techniche Subacquee. 426 pp.

Laurie, W.A. 1990. Effects of the 1982-83 El Nino-Southern Oscillation event on marine iguana (Amblyrhynchus cristatus Bell, 1925) populations on Galapagos. Elsevier Oceanography Series 52: 361-380.

Lévęque, R. 1963. Le statut actuel des vertébrés rares et menacés de l'Archipel des Galapagos. La Terre et la Vie (Paris) 110: 397-430.

Lewin, R. 1978. Galapagos: the endangered islands. New Scientist 79(1112): 168-171.

Lewin, R. 1978. Galapagos: the rise of optimism. New Scientist 79(1113): 261-263.

Louis, E. 1996. Darwin's legacy: conservation, the Galapagos tortoise (Geochelone nigra) ssp., and microsatellites. American Zoo and Aquarium Association Annual Conference Proceedings 1996: 221-224.

Mariscal, R.N. 1969. Charles Darwin and conservation in the Galapagos islands. Biological Conservation 2: 44-46.

Marquez, C.; Fritts, T.H.; Koster, F.; Rea, S.; Cepeda, F.; Llerena, F. 1995. Comportamiento de apareamiento al azar en tortugas gigantes juveniles en cautiverio en las Islas Galapagos. Revista de Ecologia LatinoAmericana 3(1-3): 13-18.

Marquez, C.; Gordillo, J.; Tupiza, A. 1994. The fire of 1994 and herpetofauna of southern Isabela. Noticias de Galapagos 54: 8-10.

Marquez, C.; Snell, H.; Snell, H. 2004. La Nina, El Nino, and the giant Galapagos tortoises of Isabela Geochelone nigra vandenburghi. Reptilia (GB) 37: 27-30.

Marquez, C.; Wiedenfeld, D.A.; Snell, H.; Fritts, T.; MacFarland, C.; Tapia, W.; Naranjo, S. 2004. Estado actual de las poblaciones de tortugas terrestres gigantes (Geochelone spp., Chelonia: Testudinae) en Las Islas Galapagos. Ecologia Aplicada 3(1-2): 98-111.

Martin, A.S. 1975. Las islas Galapagos. Zoo, Revta Parq. zool. Barcelona 22: 24-27.

Merlen, G. 1985. The 1982-83 El Nino: some of its consequences for Galapagos wildlife. Noticias de Galapagos 41: 8-15.

Mertens, R. 1960. Über die Schlangen der Galapagos. Senckenbergiana Biologica 41: 133-141.

Miller, H.F.; Miller, M.B.; Thomson, K.S.; Brandt, J.G. 1977. In the footsteps of Darwin: the Peabody Museum associates in Ecuador and the Galapagos Islands. Discovery (New Haven) 12(3): 17-30.

Moore, T.R. 1982. To eat and be eaten. Pacific Discovery 35(3): 11-20.

Niven, C. 1969. A chance of a lifetime - a trip to the Galapagos Islands. Bokmakierie 21(2): 33-35.

Parker, H.W. 1934. On the need for the preservation of the Galapagos fauna. Proceedings of the Linnean Society of London 2 [1933-34]: 80-84.

Patton, J.L. 1984. Genetical processes in the Galapagos. Biological Journal of the Linnean Society 21(1-2): 97-111.

Pauli, D.; Williams, E.E.; Hall, W.P. 1976. Lizard karyotypes from the Galapagos Islands: chromosomes in phylogeny and evolution. Breviora 441: 1-31.

Perry, R. 1982. Darwin's islands today. Country Life (London) 171(4423): 1542-1544.

Peterson, R.T. 1967. The Galapágos, eerie cradle of new species. National Geographic Magazine 131: 540-585.

Pritchard, P.C.H. 1971. Galapagos Sea Turtles - preliminary findings. Journal of Herpetology 5: 1-9.

Pritchard, P.C.H. 1973. Sea Turtles in the Galapagos Island. pp. 34-37. In: Marine turtles. Proceedings of the 2nd working meeting of marine turtle specialists. Organized by the Survival Service Commission, IUCN, 8-10 March 1971, at Morges, Switzerland. 109 pp.

Pritchard, P.C.H. 1997. Galapagos tortoise nomenclature: a reply. Chelonian Conservation and Biology 2(4): 619-621.

Quezada, M.Y. 2006. Development of the inter-American convention for the protection and conservation of Sea Turtles and its implementation in the Galapagos Islands, Ecuador. NOAA Technical Memorandum NMFS-SEFSC 536: 103-106.

Rassmann, K. 1996. Molecular genetics and conservation in the Galapagos. Noticias de Galapagos 56: 22-26.

Reynolds, R.P. 1982. Programa de herpetologia. Estacion Cientifica Charles Darwin Informe Anual 1981: 51-55.

Robinson, G.; Pino, E.M. del (eds.) 1985. El Nino en las Islas Galapagos: el evento de 1982-1983. Charles Darwin Foundation for the Galapagos Islands, Quito. 534 pp.

Roy Moore, T. de 1978. To the brink and beyond. Audubon 80(6): 75-91.

Roy, T. de; Jones, M. 1991. Portraits of Galapagos. Imprenta Mariscal, Quito, Ecuador. 115 pp.

Rueda, D.; Campbell, K.J.; Fisher, P.; Cunninghame, F.; Ponder, J.B. 2016. Biologically significant residual persistence of brodifacoum in reptiles following invasive rodent eradication, Galapagos Islands, Ecuador. Conservation Evidence 13: 38.

Schauenberg, P. 1970. Les Îles Galapagos. Revue Mens. Mus. Genčve 102: 5-10.

Schauenberg, P. 1973. L'Archipel des Galapagos: laboratoire de l'évolution. Pisciculture fr. 33: 49-58.

Scheer, G. 1957. Die Galapagos-Gruppe im Hessischen Landesmuseum zu Darmstadt. Natur und Volk 87: 134-137.

Schmitt, W.L. 1934. Hancock Galapagos Expedition, 1934. Exp. St Field-Work Smith. Inst. 1934: 17-32.

Scott, P. 1960. BBC/IUC Darwin Centenary Expedition to the British Virgin Islands, Trinidad, Panama, Ecuador and the Galapagos Islands. January-March 1959. Rep. Wildf. Tr. 11(1958-59): 61-76.

Serventy, V. 1987. The enchanted isles. Australian Natural History 22(5): 218-219.

Sitwell, N. 2011. Galapagos: a guide to the animals and plants. Wilmot Books, Chichester. 129 pp.

Slevin, J.R. 1931. Log of the Schooner "Academy". On a voyage of scientific research to the Galapagos Islands, 1905-1906. San Francisco. 162 pp.

Slevin, J.R. 1935. An account of the reptiles inhabiting the Galapagos Islands. Bulletin of the New York Zoological Society 88: 1-24.

Slevin, J.R. 1959. The Galápagos Islands. A history of their exploration. Occasional Papers of the California Academy of Sciences 25: 1-150.

Snell, H.L.; Fritts, T.H. 1983. The significance of diurnal terrestrial emergence of Green Turtles (Chelonia mydas) in the Galapagos Archipelago. Biotropica 15(4): 285-291.

Snow, D.W. 1964. Research station in the Galapagos. Oryx 7: 275-276.

Steindachner, F. 1876. Die Schlangen und Eidechsen der Galapagos Inseln. Festschrift Zool.-Bot. Ges. Wien 1876: 303-330.

Stone, C.P.; Loope, L.L.; Smith, C.W. 1988. Conservation biology in the Galapagos Archipelago: perspectives from Hawaii. Elepaio 48(1): 1-8.

Strijbos, J.P. 1977. [The reptiles of the Galapagos Islands]. (In Dutch). Levende Nat. 80(10): 233-240.

Sulloway, F.J. 1984. Darwin and the Galapagos. Biological Journal of the Linnean Society 21(1-2): 29-59.

Sulloway, F.J. 1987. Darwin and the Galapagos: three myths. Oceanus 30(2): 79-85.

Swash, A.; Still, R. 2000. Birds, mammals and reptiles of the Galapagos Islands: an identification guide. Pica Press, Robertsbridge. 168 pp.

Swash, A.; Still, R. 2005. Birds, mammals and reptiles of the Galapagos Islands: an identification guide. 2nd ed. Yale University Press, New Haven. 168 pp.

Switak, K.H. 1996. Reptiles of the Galapagos Archipelago. Monitor (Journal of the Victorian Herpetological Society) 7(3): 137-165

Thomas, R. 1997. Galápagos terrestrial snakes: biogeography and systematics. Herpetological Natural History 5(1): 19-40.

Thomas, R.A. 1997. Galapagos terrestrial snakes: biogeography and systematics. Herpetological Natural History 5(1): 19-40.

Topilow, A.A. 1994. The flora, fauna, and fishes of the Galapagos Islands. Tropical Fish Hobbyist 43(2): 211-229.

Townsend, C.H. 1928. The Galapagos Islands revisited. Bulletin of the New York Zoological Society 31(5): 148-169.

Townsend, C.H. 1930. The Astor Expedition to the Galapagos Islands. Bulletin of the New York Zoological Society 33: 135-155.

Vanzolini, P.E. 1965. On the Gonatodes of the Galapagos Islands (Sauria, Gekkonidae). Papeis Dep. Zool. Sao Paulo 17: 17-19.

Vries, T. de; Black, J. 1983. Of men, goats and guava - problems caused by introduced species in the Galapagos. Noticias de Galapagos 38: 18-21.

Wallner, L.M. 1995. The Charles Darwin Research Station: Darwins's legacy. Vivarium (Lakeside) 6(5): 26-29.

Werner, F. 2003. Iguanas of the Galápagos Islands. Iguana (West Hartford) 10(4): 145-149.

Werning, H. 2010. Auf den spuren von Darwin durch Südamerika Teil 4: Galapagos - der Darwin-Archipel. Reptilia (D) 15(81): 52-62.

West, W.R. 1989. Darwin and the Galapagos Islands. Carolina Tips 52(11): 41-44.

Wood, G.C. 1939. Results of the Pinchot South Sea Expedition. III. Galapagos reptiles. Not. Nat. Philadelphia 15: 1-4.

Wood, G.C. 1939. Zoological results of the George Vanderbilt South Pacific Expedition of 1937. IV. Galapagos reptiles. Notulae Naturae (Philadelphia) 16: 1-2.

Woram, J.M. 1991. Who killed the iguanas? Noticias de Galapagos 50: 12-17.

Wright, J.W. 1983. The evolution and biogeography of the lizards of the Galapagos Archipelago: evolutionary genetics of Phyllodactylus and Tropidurus populations. pp. 123-155. In: Bowman, R.I.; Berson, M. & Leviton, A.E. (eds.). Patterns of evolution in Galapagos organisms. Pacific Division AAAS, San Francisco. 568 pp.

Zug, G.R. 1997. Galapagos tortoise nomenclature: still unresolved. Chelonian Conservation and Biology 2(4): 618-619.

Zug, G.R. 2013. Reptiles and amphibians of the Pacific islands. A comprehensive guide. University of California Press, Berkeley & Los Angeles. 306 pp.