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Bibliography of the reptiles of
Sri Lanka

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Anonymous. 1937. Turtle oils from Ceylon. Bull. Imp. Inst. London 35(3): 316-318.

Anonymous. 1957. Cobra wins battle with viper. Loris 7(5): 421.

Anonymous. 1972. Ceylon protects all marine turtles. Marine Pollut. Bull. 3(10): 148.

Anonymous. 2000. Conservation assessment and management plan workshop for amphibians and reptiles of Sri Lanka. 26-30 November 1998, University of Peradeniya, Peradeniya, Sri Lanka. Frog Leg 4(2): 4-5.

Anonymous. 2000. Conservation Assessment and Management Plan Workshop for Amphibians and Reptiles of Sri Lanka. 26-30 November 1998, University of Peradeniya, Sri Lanka. Reptile Rap (Newletter of the Reptile Network of South Asia) 2(1): 2-5.

Anonymous. 2000. Status of endemic reptiles in Sri Lanka. Reptile Rap (Newletter of the Reptile Network of South Asia) 1(2): 2-3.

Anonymous. 2007. Sri Lanka. Tiere, Terrarien, Lebensräume. Draco 8(2) (30): 1-96.

Abayarathna, M.G.T.H.; Mahaulpatha, W.A.D.; Silva, A. de 2010. Reptile diversity of selected habitat types in the northern part of the Giritale Nature Reserve, Sri Lanka. Lyriocephalus 7(1-2): 211-221.

Abercromby, A.F. 1910. The snakes of Ceylon. Murray & Co., London. 89 pp.

Ahamad, R.; Dharmaretnam, M.; Silva, A. de 2004. Snakes from Batticaloa district: deposited in the zoology museum, Eastern University, Sri Lanka. Lyriocephalus 5(1-2): 130-134. (Special issue).

Ahlip, T.C. 1965. Our tank crocodiles. Loris 10: 242-245.

Amarakoon, A.M.R.K.; Nathanael, E.S.; Silva, A. de 2010. The pattern of reptiles killed by road traffic on the Anuradhapura-Mihintale Road, Sri Lanka. Lyriocephalus 7(1-2): 81-88.

Amarasinghe, A.A.T.; Bauer, A.M.; Ineich, I.; Rudge, J.; Bahir, M.M.; Gabadage, D.E. 2009. The original descriptions and figures of Sri Lankan gekkonid lizards (Squamata: Gekkonidae) of the 18th, 19th and 20th centuries. Taprobanica 1(2): 83-106.

Amarasinghe, A.A.T.; Manthey, U.; Stoeckli, E.; Ineich, I.; Kullander, S.O.; Tiedemann, F.; McCarthy, C.; Gabadage, D.E. 2009. The original descriptions and figures of Sri Lankan agamid lizards (Squamata: Agamidae) of the 18th and 19th centuries. Taprobanica 1(1): 2-15.

Amarasooriya, K.D. 2000. Classification of sea nesting beaches of Southern Sri Lanka. pp. 228-237. In: Pilcher, N. & Ismail, G. (eds.). Sea turtles of the Indo-Pacific: research management and conservation. Proceedings of the Second ASEAN Symposium and Workshop on Sea Turtle Biology and Conservation. ASEAN Academic Press, London. 361 pp.

Amarasooriya, K.D.; Jayathilaka, M.R.A. 2002. Marine Turtle nesting on the beaches of the north-western, western, and southern provinces of Sri Lanka. NOAA Technical Memorandum NMFS-SEFSC 477: 93-95.

Annandale, N. 1906. New and interesting lizards in the Colombo Museum. Spolia Zeylanica 3: 189-193.

Annandale, N. 1913. Some new and interesting batrachians and lizards from India, Ceylon and Borneo. Records of the Indian Museum 9(20): 301-307.

Bahir, M.M.; Surasinghe, T.D. 2005. A conservation assessment of the Sri Lankan Agamidae (Reptilia: Sauria). Raffles Bulletin of Zoology Supplement 12: 407-412.

Banks, J.; Banks, J. 1986. A selection of the animals of Sri Lanka. Lake House Investments Ltd, Colombo. 35pp. (Unpaginated).

Batuwita, S. 2000. Lizards of Mount Gannoruwa. Loris 22(3): 13-15.

Batuwita, S.; Maduwage, K.P.; Silva, A. 2004. Snakes of Mount Gannoruwa, central Sri Lanka. Lyriocephalus 5(1-2): 150. (Special issue).

Bauer, A.M. 1998. South Asian herpetological specimens of historical note in the Zoological Museum, Berlin. Hamadryad 23(2): 133-149.

Bauer, A.M.; Silva, A. de 2007. Return to Nilaveli: Edward Harrison Taylor's Sri Lankan herpetofaunal hotspot revisited. Herpetological Bulletin 102: 5-11.

Bauer, C.A.; Hoff, A.; Högstedt, G. 1971. Ceylon - några faunistiska intryck från en resa. Fauna Flora (Stockholm) 66: 149-158.

Becker, H.; Becker, R. 2015. Eine Reise zu den Schildkröten Sri Lankas. Marginata 45: ?

Berlioz, J. 1965. Aspects biogeographiques de Ceylan. Compte Rendu des Seances de la Societe de Biogeographie 365-370: 51-57.

Blanford, W.T. 1901. The distribution of vertebrate animals in India, Ceylon, and Burma. Phil. Trans. 194: 335-436.

Boake, B. 1866. On the Crocodiles of Ceylon. Journal of the Ceylon Branch of the Royal Asiatic Society 1866: 160-163.

Bossuyt, F.; Meegaskumbura, M.; Beenaerts, N.; Gower, D.J.; Pethiyagoda, R.; Roelants, K.; Mannaert, A.; Wilkinson, M.; Bahir, M.M.; Manamendra-Arachchi, K.; Ng, P.K.L.; Schneider, C.J.; Oommen, V.; Milinkovitch, M.C. 2004. Local endemism within the western Ghats-Sri Lanka biodiversity hotspot. Science (Washington, D.C.) 306(5695): 479-481.

Botejue, W.M.S.; Wattavidanage, J. 2011. Herpetofaunal diversity and distribution in Kalugala proposed forest reserve, Western province of Sri Lanka. Amphibian and Reptile Conservation 5(2): 65-80.

Brodie, J.; Sanjayan, M.; Corea, R.; Helmy, O.; Amarasin, C. 2008. Effects of the 2004 Indian Ocean tsunami on Sea Turtle populations in Sri Lanka. Chelonian Conservation and Biology 7(2): 249-251.

Burt, D.R.R. 1954. Lizards. Loris 6(5): 201-204.

Chalalochani, H.M.N.; Nathanael, E.S.; Silva, A. de 2010. Amphibians and reptiles of the Rajarata University, Campus Park, Mihintale. Lyriocephalus 7(1-2): 9-17.

Chalalochani, H.M.N.; Nathanael, E.S.; Silva, A. de 2010. Knowledge, attitude and practice towards amphibians and reptiles at Mihintale. Lyriocephalus 7(1-2): 99-103.

Chambers, M.R. 1980. Riddle of the red crocodiles at Yala National Park. Loris 15(3): 177.

Crusz, H. 1986. The vertebrates of Sri Lanka: endemism and other aspects. Report of the Society for Researches on Native Livestock 11: 65-80.

Das, I.; Silva, A. de 2005. A photographic guide to snakes and other reptiles of Sri Lanka. New Holland, London, etc. 144 pp.

Das, I.; Silva, A. de 2010. A Photographic Guide to Snakes and other Reptiles of Sri Lanka. 2nd ed. New Holland Publishers, London. 144 pp.

Das, I.; Silva, A. de; Jayantha, J. 2016. Illegal philatelic issues in the name of the Republic of Rwanda, depicting Sri Lankan snake images. Bibliotheca Herpetologica 12(1): 56-59.

Dattatri, S.; Samarajiva, D. 1982. Beach development in Sri Lanka. Marine Turtle Newsletter 22: 4-5.

Deraniyagala, P.E.P. 1930. The Crocodiles of Ceylon. Ceylon Journal of Science (B) 16: 89-95.

Deraniyagala, P.E.P. 1930. The Testudinata of Ceylon. Ceylon Journal of Science (B) 16: 43-88.

Deraniyagala, P.E.P. 1931. Some Ceylonese lizards. Spolia Zeylanica 16B: 139-180.

Deraniyagala, P.E.P. 1932. Them Gekkonoideae of Ceylon. Ceylon Journal of Science (B) 16: 291-310.

Deraniyagala, P.E.P. 1933. Cured marine products of Ceylon. Bull. Ceylon Fisheries Colombo 5: 49-78.

Deraniyagala, P.E.P. 1934. Some new fossorial skinks of Ceylon. Ceylon Journal of Science (B) 18: 231-233.

Deraniyagala, P.E.P. 1939. The tetrapod reptiles of Ceylon. Colombo Museum, Colombo. 412 pp.

Deraniyagala, P.E.P. 1949. Some cross-banded geckoes of India and Ceylon. (Abstract). Proceedings Indian Sci. Congr. 32nd(3) [1946]: 114.

Deraniyagala, P.E.P. 1949. Some vertebrate animals of Ceylon. 1. Publisher unknown. 118 pp.

Deraniyagala, P.E.P. 1953. A coloured atlas of some vertebrates from Ceylon. Volume 2: Tetrapod Reptilia. Ceylon Government Press, Colombo. 101 pp.

Deraniyagala, P.E.P. 1955. A coloured atlas of some vertebrates from Ceylon. 3. Serpentoid Reptilia. National Museum, Colombo. 121 pp.

Deraniyagala, P.E.P. 1960. Some southern temperature zone snakes, birds and whales that enter the Ceylon area. Spolia Zeylanica 29: 79-85.

Deraniyagala, P.E.P. 1961. Some of Linne's reptiles and a mammal from, Ceylon and elsewhere that are in Sweden. Spolia Zeylanica 29: 195.

Dissanayake, D.S.B.; Wellappuliarchchi, S.M. 2014. Knowledge and attitude on highly venomous snakes by questionnaire survey among the students of Rajarata University, Mihintale, Sri Lanka. International Research Journal of Biological Sciences 3(12): 67-72.

Dryden, G.L.; Wikramanayake, E.D. 1991. Space and time sharing by Varanus salvator and Varanus bengalensis in Sri Lanka. Mertensiella 2: 111-119.

Eisenberg, J.F.; Lockhart, M. 1972. An ecological reconnaissance of Wilpattu National Park, Ceylon. Smithsonian Contributions to Zoology 101: 1-118.

Ekanayake, E.M.L.; Kapurusinghe, T.; Saman, M.M.; Premakumara, M.G.C. 2002. Estimation of the number of leatherback (Dermochelys coriacea) nesting at Godavaya turtle rookery in southern Sri Lanka during the nesting season in the year 2001. Kachhapa Newsletter 2002: 6.

Ekanayake, E.M.L.; Ranawana, K.B. 2004. The average egg count for nesting Sea Turtles on Rekawa Beach in Sri Lanka. Lyriocephalus 5(1-2): 32-34. (Special issue).

Ekanayake, E.M.L.; Ranawana, K.B. 2006. Average egg counts, weights and diameters for five turtle species nesting in Rekawa, southern Sri Lanka. NOAA Technical Memorandum NMFS-SEFSC 536: 172-173.

Ekanayake, E.M.L.; Ranawana, K.B.; Kapurusinghe, T.; Premakumara, M.G.C.; Saman, M.M. 2002. Marine Turtle conservation in Rekawa turtle rookery in southern Sri Lanka. Ceylon Journal of Science Biological Sciences 30: 79-88.

Ekanayake, S.P.; Bambaradeniya, C.N.B.; Perera, W.P.N.; Perera, M.S.J.; Rodrigo, R.K.; Samarawickrama, V.A.P.; Peiris, T.N. 2005. A biodiversity status profile of Lunama - Kalametiya Wetland Sanctuary. Occasional Papers of IUCN Sri Lanka 8: 1-43.

Elzen, P. van den 1972. Results of the Austrian-Ceylonese hydrobiological mission, 1970 of the lst Zoological Institute of the University of Vienna (Austria) and the Department of Zoology of the Vidyalankara University of Ceylon, Kelaniya, (Sri Lanka) Ceylon. Bulletin Fish. Res. Stn Ceylon. 23(1-2): 113-125.

Ende, H.J. 1999. Meeresschildkröten. Schutz der Meeresschildkröten in Sri Lanka. Aquarium (Bornheim) 358: 50-53.

Environmental Investigation Agency 1995. Report on an investigation into threats to marine turtles in Sri Lanka and the Maldives, 26th November - 14th December 1994. Environmental Investigation Agency, London & Washington, D.C. 28 pp.

Erdelen, W. 1978. Distributional patterns of the genus Calotes (Sauria Agamidae) of Sri Lanka. Loris 14(6): 350-353.

Erdelen, W. 1984. The genus Calotes (Sauria, Agamidae) in Sri Lanka: distribution patterns. Journal of Biogeography 11(6): 515-525.

Erdelen, W. 1986. The genus Calotes (Sauria: Agamidae) in Sri Lanka: clutch sizes and reproductive seasonality of Calotes versicolor - preliminary results. Spixiana 9(1): 111-115.

Erdelen, W.R. 2011. Conservation of biodiversity in a hotspot: Sri Lanka's amphibians and reptiles. Amphibian and Reptile Conservation 5(2): 34-51.

Eschment, J.; Manthey, S.; Manthey, U. 1980. Für den Terrarianer eine Reste wert: Sri Lanka. 1. Der Sinharaja Forest. Aquarien Magazin 14(6): 296-305.

Farook, S.M.S. 1979. Our crocodiles... why protect them? Loris 15(1): 45-46.

Fernando, R. 1977. Turtle hatcheries in Sri Lanka. Marine Turtle Newsletter 3: 8.

Fletcher, T.B. 1908. Notes on snakes from Diyatalawa, Ceylon. Spolia Zeylanica 5: 98-101.

Gaulke, M.; Silva, A. de 1997. Monitor lizards of Sri Lanka: preliminary investigations on their population structure. Lyriocephalus 3(1): 1-5.

Grammentz, D. 2011. Gefährdung und "Schutz" des Sumpfkrokodils (Crocodylus palustris) und des Leistenkrokodils (Crocodylus porosus) auf Sri Lanka. Reptilia (D) 16(87): 36-45.

Haly, A. 1886. First report on the collection of lizards in the Colombo Museum. Colombo. 8 pp.

Haly, A. 1886. First report on the collection of snakes in the Colombo Museum. Colombo. 18 pp.

Haly, A. 1887. Second report on the collection of lizards in the Colombo Museum. Colombo. 4 pp.

Haly, A. 1892. Essay on the construction of zoological tables, with a tabular diagnosis of the snakes of Ceylon. Journal of the Ceylon Branch of the Royal Asiatic Society 11: 22-32, 183-232.

Haly, A. 1895. Notes on the species and varieties of Testudo in the Colombo Museum. Journal of the Ceylon Branch of the Royal Asiatic Society 13: 128-132.

Herath, N.D.; Rathnayake, N.D.; Hewamathes, K.K. 2001. Some observations on the amphibians in Udawattakele sanctuary, Kandy. Lyriocephalus 4(1-2): 116-120 (Special issue).

Hettiarachchi, D.K.; Wellapuliarachchi, S.M.; Nathanael, E.S.; Silva, A. de; Goonasekera, M.M. 2010. Knowledge, attitudes and traditional beliefs on geckoes at Mihintale. Lyriocephalus 7(1-2): 105-109.

Hettiarachchi, D.K.; Wellapuliarachchi, S.M.; Nathanael, E.S.; Silva, A. de; Goonasekera, M.M.; Bauer, A.M.; Rudge, J. 2010. The gecko diversity of the Mihintale Sanctuary, Sri Lanka. Lyriocephalus 7(1-2): 33-41.

Hewavisenthi, S. 1990. Exploitation of marine turtles in Sri Lanka: historic background and the present status. Marine Turtle Newsletter 48: 14-19.

Hewavisenthi, S. 1993. Turtle hatcheries in Sri Lanka: boon or bane? Marine Turtle Newsletter 60: 19-22.

Hewavisenthi, S. 1994. Bibliography of literature relating to marine turtles of Sri Lanka. Natural Resources, Energy and Science Authority of Sri Lanka, Colombo. I-iv, I-vi, 1-17.

Hewitt, J. 1929. On some Scincidae from South Africa, Madagascar and Ceylon. Annals of the Transvaal Museum 13: 1-8.

Holtzinger-Tenever, H. 1917. Verzeichnis der von Herrn. H. Schöde auf Ceylon und Sumatra gesammelten Reptilien. Mitteilungen aus dem Zoologischen Museum in Berlin 8: 425-454.

Ilangakoon, A. 2000. Marine Turtles: imperilled ancient ocean wanderers. Loris 22(3): 18-21.

IUCN 2008. A preliminary survey on the herpetofauna in the Anawilundawa wetland Sanctuary: the second Ramsar site of Sri Lanka. Tigerpaper (Bangkok) 35(3): 27-32.

Janzen, P. 2003. Sinharaja - Der Regenwald des Loewenkönigs. Draco 4(3) (15): 26-33.

Janzen, P.; Bopage, M. 2011. The herpetofauna of a small and unprotected patch of tropical rainforest in Morningside, Sri Lanka. Amphibian and Reptile Conservation 5(2): 1-13.

Janzen, P.; Klaas, P.; Ziesmann, S. 2007. Die Agamenfauna der Insel. Draco 8(2) (30): 24-33.

Janzen, P.; Klaas, P.; Ziesmann, S. 2007. Krokodile und Schildkröten auf Sri Lanka. Draco 8(2) (30): 45-49.

Janzen, P.; Klaas, P.; Ziesmann, S. 2007. Sri Lankas Schlangenfauna. Draco 8(2) (30): 56-64.

Jayewardene, J. 1982. The conservation of marine turtles in Sri Lanka. Tigerpaper (Bangkok) 9(2): 21-26.

Jirousek, V.T. 1983. Remarks on reptile protection in Sri Lanka and the Indian State of Tamil Nadu. Hamadryad 8(1): 18-19.

Jonklass, R. 1977. Crocodiles in Sri Lanka. Newsletter of the Madras Snake Park Trust 2(2): 2.

Kandamby, D.S.; Batuwita, S. 2004. Some reptiles from Galle district, southern Sri Lanka (Reptilia: Sauria & Ophidia). Lyriocephalus 5(1-2): 152-155. (Special issue).

Kapurusinghe, T. 2000. Community participation in turtle conservation in Sri Lanka. pp. 35-44. In: Pilcher, N. & Ismail, G. (eds.). Sea turtles of the Indo-Pacific: research management and conservation. Proceedings of the Second ASEAN Symposium and Workshop on Sea Turtle Biology and Conservation. ASEAN Academic Press, London. 361 pp.

Kapurusinghe, T. 2003. TCP, Sri Lanka initiates a new in-situ turtle nest protection programme. Kachhapa Newsletter 9: 8.

Kapurusinghe, T. 2006. Status and conservation of Marine Turtles in Sri Lanka. pp. 173-187. In: Shanker, K.; Choudhury, B.C. (eds.). Marine turtles of the Indian subcontinent. Universities Press, Hyderabad. 415 pp.

Kapurusinghe, T.; Ekanayake, L.; Saman, M.M.; Rathnakumara, D.S. 2008. Community-based Marine Turtle conservation in Kosgoda, Sri Lanka: nesting results from 2005 to 2007. Testudo 6(5): 40-47.

Karunarathna, D.M.S.S.; Amarasinghe, A.A.T. 2011. A preliminary survey of the reptile fauna in Nilgala Forest and its vicinity, Monaragala District, Sri Lanka. Taprobanica 3(2): 69-76.

Karunarathna, D.M.S.S.; Amarasinghe, A.A.T. 2012. Reptile diversity in Beraliya Mukalana proposed forest reserve, Galle District, Sri Lanka. Taprobanica 4(1): 20-26.

Karunarathna, D.M.S.S.; Amarasinghe, A.A.T.; Gabadage, D.E.; Bajir, M.M.; Harding, L.E. 2010. Current status of faunal diversity in Bellanwila-Attidiya Sanctuary, Colombo District - Sri Lanka. Taprobanica 2(1): 48-63.

Karunarathna, D.M.S.S.; Amarasinghe, A.A.T.; Karunarathna, D.M.G.N. 2011. Study of the reptilian faunal diversity of a fragmented forest patch in Kukulugala, Ratnapura district, Sri Lanka. Tigerpaper (Bangkok) 38(4): 19-27.

Karunarathna, D.M.S.S.; Amarasinghe, A.A.T.; Madawala, M.B.; Kandambi, H.K.D. 2012. Population status of two Varanus species (Reptilia: Sauria: Varanidae) in Sri Lanka's Puttalam Lagoon system, with notes on their diet and conservation status. Biawak 6(1): 22-33.

Karunarathna, S.; Ranwala, S.; Surasinghe, T.; Madawala, M. 2017. Impact of vehicular traffic on vertebrate fauna in Horton Plains and Yala National Parks of Sri Lanka: some implications for conservation and management. Journal of Threatened Taxa 9(3): 9928-9939.

Kularathne, S.A.M. 2013. Snakes, snakebite, and envenoming in Sri Lanka. A handbook on management of snakebite. 2nd edition. Gampola. 219 pp.

Kumari, M.D.I.P.; Nathanael, E.S.; Austin, C.C.; Silva, A. de 2010. The scincid lizards (Squamata: Scincidae) of Mihintale with special reference to Lankascincus fallax. Lyriocephalus 7(1-2): 57-64.

Kumari, M.D.I.P.; Nathanael, E.S.; Silva, A. de 2010. Knowledge and attitude towards scincid lizards (Squamata: Scincidae) by some inhabitants at Mihintale. Lyriocephalus 7(1-2): 111-113.

Leist, J.; Eschment, J. 1981. Für den Terrarianer eine Reise wert: Sri Lanka 3. Das Hochland der Gammaduwa/Elkadawa-Berggruppe. Aquarien Magazin 15(1): 23-27.

Maduwage, K.; Meegaskumbura, M.; Silva, A.; Pethiyagoda, R. 2008. Phylogenetic implications of hemipenial morphology in Sri Lankan agamid lizards. Current Science (Bangalore) 95(7): 838-840.

Mahendra, B.C. 1984. Handbook of the snakes of India, Ceylon, Burma, Bangladesh and Pakistan. Annals of Zoology (Agra) 22(B): 1-412.

Manamendra-Arachchi, K.; Batuwita, S.; Pethiyagoda, R. 2007. A taxonomic revision of the Sri Lankan day-geckos (Reptilia: Gekkonidae: Cnemaspis), with description of new species from Sri Lanka and southern India. Zeylanica 7(1): 9-122.

Manamendra-Arachchi, K.; Liyanage, S. 1994. Conservation and distribution of the agamid lizards of Sri Lanka with illustrations of the extant species. Journal of South Asian Natural History 1(1): 77-96.

Manthey, S.; Manthey, U. 1980. Für den Terrarianer eine Reise wert: Sri Lanka. 2. Das Hochland von Nuwara Eliya. Aquarien Magazin 14(8): 404-411.

Manthey, S.; Manthey, U. 1981. Für den Terrarianer eine Reise wert: Sri Lanka. 4. Im Land der Weddas - die Trockengebiete. Aquarien Magazin 15(5): 298-301.

Manthey, U. 1981. Die Echsen des Ceylonesischen Regenwaldes und seiner Randgebiete. Sauria (Berlin) 3(2): 25-35.

Mehely, L.V. von 1897. Zur Herpetologie von Ceylon. Termes. Fuzetek 20: 55-70.

Mendis, A.S.; Fernando, C.H. 1962. A guide to the freshwater fauna of Ceylon. Fish. Res. St. Dept. Fish. Ceylon 12: 1-160.

Mills, S. 1994. Turning turtles. BBC Wildlife 12(8): 26-28.

Munsch, W. 1987. Warane in Sri Lanka (Ceylon) erlebt. Aquarium (Bornheim) 214: 206-210.

Murthy, T.S.N. 1992. A distributional analysis of the uropeltid snakes of India and Sri Lanka. Records of the Zoological Survey of India 91(3-4): 375-387.

Nanayakkara, G.L.A. 1991. Conservation of the snakes of Sri Lanka. Hamadryad 16(1-2): 46-47.

Nathanael, S.; Pradeep, D.M.N.; Samarakoon, S.M.M.; Silva, A. de 2004. Herpetofaunal diversity with special reference to endemic and nationally threatened species at Polgolla, an upland area in the mid country wet zone of Sri Lanka. Lyriocephalus 5(1-2): 86-95. (Special issue).

Nicholls, L. 1929. The identification of the land snakes of Ceylon. Ceylon Journal of Science 2: 91-142.

Nicholls, L. 1932. Notes on Ceylon snakes. Ceylon Journal of Science (B) 17: 39-40.

Norris, C.E. 1956. A new wild life reserve in Ceylon. Oryx 3: 240-243.

Pawlowski, S. 2011. Zur Situation von Meeresschildkröten in Sri Lanka. Marginata 29: 36-41.

Pawlowski, S. 2014. [The status of turtles in Sri Lanka]. (In German). Marginata 42: 46-53.

Pawlowski, S.; Krämer, C. 2008. Besuch zweier Aufzuchtstationen von Meeresschildkröten im Südwesten Sri Lankas. Sauria (Berlin) 30(2): 33-40.

Peabotuwage, I.; Nuwan, B.I.; Samarasinghe, D.; Perera, N.; Madawala, M.; Amarasinghe, D.; Kandambi, H.K.D.; Karunarathna, D.M.S.S. 2011. Range extension for Duttaphrynus kotagamai (Amphibia: Bufonidae) and a preliminary checklist of herpetofauna from the Uda Maliboda Trail in Samanala Nature Reserve, Sri Lanka. Amphibian and Reptile Conservation 5(2): 52-64.

Pearless, S.H. 1909. Snakes of Badulla. Spolia Zeylanica 6: 54-55.

Phillips, W.W.A. 1940. A list of the more important animals, birds and reptiles of Ceylon, with the Sinhalese and Tamil names of each. Loris 2: 140-144.

Phillips, W.W.A. 1941. Misnomers. [Trivial names of Ceylon animals]. Loris 2: 220-223.

Ponnambalam, C. 1960. Ceylon snakes. Only five of 64 species can deal death. Loris 8: 378-381.

Porej, D. 2001. Herpetofauna of the Muthurajawela Marsh, Sri Lanka, with notes on natural history. Herpetological Natural History 8(1): 27-35.

Pradeep, D.M.N.; Wijegunasekera, H.N.P. 2004. Predatory patterns of paddy field pests by amphibians and reptiles of Sri Lanka. Lyriocephalus 5(1-2): 96-102. (Special issue).

Punchihewa, G.S.G. 2000. Animals far and near. Navrang, New Delhi. 167 pp.

Pyron, R.A.; Kandambi, H.K.D.; Hendry, C.R.; Pushpamal, V.; Burbrink, F.T.; Somaweera, R. 2013. Genus-level phylogeny of snakes reveals the origins of species richness in Sri Lanka. Molecular Phylogenetics and Evolution 66(3): 969-978.

Rajakaruna, R.S.; Dissanayake, D.M.N.J.; Ekanayake, E.M.L.; Ranawana, K.B. 2009. Sea Turtle conservation in Sri Lanka: assessment of knowledge, attitude and prevalence of consumptive use of turtle products among coastal communities. Indian Ocean Turtle Newsletter 10: 1-13.

Rajapaksha, D.R.N.S.; Samarawickrama, V.A.P.; Ranawana, K.B. 2006. Herpetofaunal diversity in Dothalugala Man and Biosphere Reserve, of the Knuckles Forest Range, Sri Lanka. Russian Journal of Herpetology 13(2): 120-134.

Ranasinghe, L.; Uragoda, C.G. (eds.) 1983. Symposium: medically important snakes and snake-bite in Sri Lanka. Ceylon Medical Journal 28(3): 107-201.

Rathnayake, N.D. 2001. Some notes on Sri Lankan Cat Snakes. Loris 22(5): 51-52.

Ratnapala, R. 1983. Key to the medically important venomous snakes of Sri Lanka. Ceylon Medical Journal 28(3): 130.

Ratnapala, R.; Aloysius, D.J.; Ranasinghe, L. 1983. Deadly venomous land snakes of Sri Lanka. Ceylon Medical Journal 28(3): 112-117.

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Richardson, P. 1994. The turtle conservation project (TCP) Sri Lanka: how much is a turtle worth?. Testudo 4(1): 17-21.

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