Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus Recent updates Related bibliographies: Reptiles Snakes Colubridae North America Taxonomy of the genus Nerodia Biodiversity of the family Colubridae Bibliography of the genus Nerodia (North American Water Snakes) (Reptilia: Serpentes: Colubridae) Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted. Nerodia in general Allen, B.A.; Burghardt, G.M.; York, D.S. 1984. Species and sex differences in substrate preference and tongue flick rate in three sympatric species of water snakes (Nerodia). Journal of Comparative Psychology 98(4): 358-367. Allen, E.R.; Neill, W.T. 1957. The water snake. Florida Wild Life 1952 [1957]: unpaginated. Balfour, P.S.; Stitt, E.W. 2008. Transcontinental introductions of watersnakes (Nerodia) into California. Proceedings of the Vertebrate Pest Conference 23: 301-303. Bartlett, R.D. 2001. Notes on the American Water Snakes: genus Nerodia. Reptile & Amphibian Hobbyist 6(5): 46-54. Blaney, R.M.; Blaney, P.K. 1979. The Nerodia sipedon complex of water snakes in Mississippi and southeastern Louisiana. Herpetologica 35(4): 350-359. Chiasson, R.B.; Lowe, C.H. 1989. Ultrastructural Scale Patterns in Nerodia and Thamnophis. Journal of Herpetology 23(2): 109-118. Clay, W.M. 1938. A synopsis of the North American water snakes of the genus Natrix. Copeia 1938: 173-182. Cliburn, J.W. 1975. Diel activity of Natrix in Bear Creek, Tishomingo County, Mississippi. Journal of the Mississippi Academy of Sciences 19: 160-163. Cochran, P.A. 1986. Opportunistic scavenging by a garter snake in Colorado. Bulletin of the Chicago Herpetological Society 21(3-4): 94-95. Conant, R. 1961. Comments on the distribution of Natrix and Thamnophis eques in Mexico. Journal Ohio Herpetological Society 3(2): 27. Conant, R. 1969. A review of the water snakes of the genus Natrix in Mexico. Bulletin of the American Museum of Natural History 142: 1-140. Densmore, L.D.; Rose, F.L.; Kain, S.J. 1992. Mitochondrial DNA evolution and speciation in water snakes (genus Nerodia) with special reference to Nerodia harteri. Herpetologica 48(1): 60-68. Dwyer, C.M.; Kaiser, H. 1997. Relationship between skull form and prey selection in the thamnophiine snake genera Nerodia and Regina. Journal of Herpetology 31(4): 463-475. Eberle, W.G. 1972. Comparative chromosomal morphology of the New World natricine snake genera Natrix and Regina. Herpetologica 28(2): 98-105. Ferrier, W. 2000. Nerodia, New World Water Snakes. Reptilia (GB) 13: 56-64. Fontenot, L.W.; Platt, S.G.; Dwyer, C.M. 1993. Observations on crayfish predation by water snakes, Nerodia (Reptilia: Colubridae). Brimleyana 19: 95-99. Franklin, L. 1975. Care and feeding of native water snakes (genus Natrix). Bulletin Chicago Herpetological Society 10(1-2): 2-9. Gibbons, J.W.; Dorcas, M.E. 2004. North American watersnakes. A natural history. University of Oklahoma, Norman. 496 pp. Gregory, P.T. 2009. Variation in size of cover used by watersnakes (Nerodia): do smaller snakes use smaller rocks? Acta Oecologica 35(2): 182-187. Hampton, P.M. 2011. Comparison of cranial form and function in association with diet in natricine snakes. Journal of Morphology 272(12): 1435-1443. Hanebrink, E.; Byrd, W. 1986. Species composition and diversity of water snake (Nerodia sp.) populations in northeastern Arkansas. Bulletin of the Chicago Herpetological Society 21(3-4): 72-78. Hebrard, J.J.; Mushinsky, H.R. 1978. Habitat use by five sympatric water snakes in a Louisiana swamp. Herpetologica 34(3): 306-311. Hoser, R.T. 2012. Two new genera of water snake from North America. The subdivision of the genera Regina Baird and Girard, 1853 and Nerodia Baird and Girard, 1853 (Serpentes: Colubridae: Natricinae). Australasian Journal of Herpetology 11: 28-31. Keck, M.B. 1998. Habitat use by semi-aquatic snakes at ponds on a reclaimed strip mine. Southwestern Naturalist 43(1): 13-19. Kochman, H.I. 1977. Differentiation and hybridization in the Natrix fasciata complex (Reptilia: Serpentes): a non-morphological approach. Herpetological Review 8(3)(Suppl.): 11. Kofron, C.P. 1978. Foods and habitats of aquatic snakes (Reptilia, Serpentes) in a Louisiana swamp. Journal of Herpetology 12(4): 543-554. Lawson, R. 1987. Molecular studies of thamnophiine snakes: 1. The phylogeny of the genus Nerodia. Journal of Herpetology 21(2): 140-157. Lawson, R.; Meier, A.J.; Frank, P.G.; Moler, P.E. 1991. Allozyme variation and systematics of Nerodia fasciata - Nerodia clarkii complex of water snakes (Serpentes: Colubridae). Copeia 1991(3): 638-659. Malnate, E.V. 1960. Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae. Proceedings of the Academy of Natural Sciences of Philadelphia 112: 41-71. Mara, W.P. 1995. Water Snakes of North America. T.F.H. Publications, Neptune City, New Jersey. 64 pp. McVay, J.D.; Flores-Villela, O.; Carstens, B. 2015. Diversification of North American natricine snakes. Biological Journal of the Linnean Society 116(1): 1-12. Michot, T.C. 1982. Thermal and spatial ecology of three species of water snakes (Nerodia) in a Louisiana swamp. Dissertation Abstracts International B Sciences and Engineering 42(11): 4292. Mushinsky, H.R.; Hebrard, J.J. 1977. Food partitioning by five species of water snakes in Louisiana. Herpetologica 33(2): 162-166. Mushinsky, H.R.; Hebrard, J.J.; Vodopich, D.S. 1982. Ontogeny of water snake foraging ecology. Ecology 63(6): 1624-1629. Pewtress, R. 1984. Keeping North American water snakes in captivity (Nerodia). Herptile 9(3): 110-113. Rossi, J.V.; Rossi, R. 2000. Captive care of North American colubrid snakes (king snakes, rat snakes, indigos, hognoses, garters, waters and others). Journal of Herpetological Medicine and Surgery 10(3-4): 31-33. Rossi, J.V.; Rossi, R. 2000. Husbandry of North American colubrid snakes. Journal of Herpetological Medicine and Surgery 10(3-4): 24-30. Rossman, D.A. 1995. A second external character for distinguishing garter snakes (Thamnophis) from water snakes (Nerodia). Herpetological Review 26(4): 182-183. Rossman, D.A.; Eberle, W.G. 1977. Partition of the genus Natrix, with preliminary observations on evolutionary trends in natricine snakes. Herpetologica 33(1): 34-43. Sanderson, W.E. 1983. Systematics of the water snakes of the Nerodia cyclopion complex. M.S. thesis, Louisiana State University, Baton Rouge. 103 pp. Scudder, L.M. 1984. Food consumption and growth in three species of water snakes of the genus Nerodia (Colubridae). Dissertation Abstracts International B Sciences and Engineering 45(1): 92. Strathemann, U. 1995. Freilandhaltung nordamerikanischer Wassernattern der Gattungen Thamnophis und Nerodia Teil 2: Erfahrungen mit Thamnophis und Nerodia. Sauria (Berlin) 17(2): 15-23. Strathemann, U. 1995. Freilandhaltung nordamerikanischer Wassernattern der Gattungen Thamnophis und Nerodia. Teil 1: Planung, Anlage und Bau der Freilandanlage. Sauria (Berlin) 17(1): 31-34. Viosca, P. 1924. A contribution to our knowledge of the water snakes. Copeia 1924(126): 3-13. Warren, M.L.; Haag, W.R.; Commens, A.M. 2004. Observations of watersnake (Nerodia, Colubridae) predation on darters (Percidae). Southeastern Fishes Council Proceedings 47: 1-3. Nerodia clarkii Ackley, J.W.; Meylan, P.A. 2010. Watersnake eden: use of stormwater retention ponds by Mangrove Salt Marsh Snakes (Nerodia clarkii compressicauda) in urban Florida. Herpetological Conservation and Biology 5(1): 17-22. Barbour, T.; Noble, G.K. 1915. Notes on the watersnake, Natrix compressicauda. Proceedings of the Academy of Natural Sciences of Philadelphia 67: 29-35. Bishop, M.B. 1940. Notes concerning two broods of young of Natrix compressicauda. Copeia 1940: 128. Carr, A.F.; Goin, C.J. 1942. Rehabilitation of Natrix sipedon taeniata Cope. Proceedings of the New England Zoological Club 21: 47-54. Chamberlain, E.B. 1937. Clark's watersnake on the Mid-Texas coast. Copeia 1937: 140. Cliburn, J.W. 1957. Some southern races of the common water snake, Natrix sipedon. Herpetologica 13: 193-202. Cope, E.D. 1895. On some new North American snakes. American Naturalist 29: 676-680. Dunson, W.A. 1979. Occurrence of partially striped forms of the mangrove snake Nerodia fasciata compressicauda Kennicott and comments on the status of N. f. taeniata Cope. Florida Scientist 42(2): 102-112. Hansknecht, K.A. 2008. Lingual luring by mangrove saltmarsh snakes (Nerodia clarkii compressicauda). Journal of Herpetology 42(1): 9-15. Hansknecht, K.A.; Burghardt, G.M. 2010. Stimulus control of lingual predatory luring and related foraging tactics of Mangrove Saltmarsh Snakes (Nerodia clarkii compressicauda). Journal of Comparative Psychology 124(2): 159-165. Holbrook, J.D.; Chesnes, T.; Boss, H.; Berman, A.; Chiyka, J.; Bell, C.; Bell, C. 2016. Geographic distribution: Nerodia clarkii compressicauda (Mangrove Saltmarsh Watersnake). Herpetological Review 47(1): 84. Jansen, K.P.; Mushinsky, H.R.; Karl, S.A. 2008. Population genetics of the mangrove salt marsh snake, Nerodia clarkii compressicauda, in a linear, fragmented habitat. Conservation Genetics 9(2): 401-410. Jones, M.P. 1977. Rulemaking actions May-June 6, 1977. Proposed rulings. Atlantic salt marsh snake. Endangered Species Technical Bulletin 2(6): 5-6. Lawson, R.; Meier, A.J.; Frank, P.G.; Moler, P.E. 1991. Allozyme variation and systematics of Nerodia fasciata - Nerodia clarkii complex of water snakes (Serpentes: Colubridae). Copeia 1991(3): 638-659. List, J.C.; Smith, P.W. 1954. A rare snake anomaly. American Midland Naturalist 51(1): 312-313. Ljustina, O.; Erdmann, J.A.; Doody, J.S. 2017. Natural history notes: Nerodia clarkii (Saltmarsh Watersnake). Retreat site / habitat use. Herpetological Review 48(3): 678-679. Marion, K.R. 1986. Gulf salt marsh snake. pp. 47-48. In: Mount, R.H. (ed.). Vertebrate animals of Alabama in need of special attention. Alabama Agricultural Experiment Station, Auburn, Alabama. 124 pp. McCardle, L.D.; Haase, A.; Fontenot, C.L. 2014. Geographic distribution: Nerodia clarkii (Gulf Salt Marsh Snake). Herpetological Review 45(1): 95. Miller, D.E. 1985. Rain water drinking by the mangrove water snake, Nerodia fasciata compressicauda. Herpetological Review 16(3): 71. Miller, D.E.; Mushinsky, H.R. 1990. Foraging ecology and prey size in the mangrove water snake, Nerodia fasciata compressicauda. Copeia 1990(4): 1099-1106. Mohrman, T.J.; Ennen, J.R.; Qualls, C.; Kreiser, B. 2008. Natural history notes: Nerodia clarkii (Saltmarsh Watersnake). Predation. Herpetological Review 39(3): 355-356. Mullin, S.J.; Mushinsky, H.R. 1995. Foraging ecology of the mangrove salt marsh snake, Nerodia clarkii compressicauda: effects of vegetational density. Amphibia-Reptilia 16(2): 167-175. Mullin, S.J.; Mushinsky, H.R. 1997. Use of experimental enclosures to examine foraging success in water snakes: a case study. Journal of Herpetology 31(4): 565-569. Neill, W.T. 1965. Notes on aquatic snakes, Natrix and Tretanorhinus, in Cuba. Herpetologica 21(1): 62-67. Phillips, C. 1940. The flat-tailed water snake. Proceedings Florida Acad. Sci. 4: 210-211. Rightmire, J.E. 2001. Some threatened U.S. snakes: five species on the red list. Reptile & Amphibian Hobbyist 6(10): 27-32. Shahrudin, S. 2013. Natural history notes: Nerodia clarkii clarkii (Gulf Saltmarsh Watersnalce). Predation. Herpetological Review 44(1): 156-157. Swanson, V.L. 1948. Natrix sipedon compressicauda at Key West, Florida. Herpetologica 4(3): 105-106. Nerodia cyclopion Cooper, W.E. 1977. Geographic distribution: Natrix cyclopion cyclopion x floridana. USA: Alabama: Baldwin Co. Herpetological Review 8(1): 13-14. Dartez, S.F.; Hampton, P.M.; Haertle, N.E.; Monteiro, C.S. 2011. Natural history notes: Lampropeltis getula holbrooki (Speckled Kingsnake). Diet. Herpetological Review 42(2): 292. Garton, J.S. 1970. Descriptive and ecological notes on Natrix cyclopion in Illinois. Herpetologica 26: 454-461. Karlström, A.; Steen, D.A. 2016. Geographic distribution: Nerodia cyclopion (Mississippi Green Watersnake). Herpetological Review 47(4): 631. Keiser, E.D. 1958. The green water snake in Illinois. Herpetologica 13: 260. Kofron, C.P. 1979. Reproduction of aquatic snakes in south central Louisiana. Herpetologica 35(1): 44-50. Lodrigue, K.J. 2007. Geographic distribution: Nerodia cyclopion cyclopion (Green Watersnake). Herpetological Review 38(4): 488. McAllister, C.T.; Robison, H.W.; Trauth, S.E.; Tumlison, R. 2014. Geographic distribution: Nerodia cyclopion (Mississippi Green Watersnake). Herpetological Review 45(4): 665. Mccardle, L.D.; Fontenot, C.L. 2016. The influence of thermal biology on road mortality risk in snakes. Journal of Thermal Biology 56: 39-49. McDonald, H.S. 1960. Nasal obstruction during the premolt period in Natrix. Herpetologica 16: 115-119. Mushinsky, H.R.; Hebrard, J.J.; Walley, M.G. 1980. The role of temperature on the behavioral and ecological associations of sympatric water snakes. Copeia 1980(4): 744-754. Sanderson, W.E. 1983. Systematics of the water snakes of the Nerodia cyclopion complex. M.S. thesis, Louisiana State University, Baton Rouge. 103 pp. Sanderson, W.E. 1993. Additional evidence for the specific status of Nerodia cyclopion and Nerodia floridana (Reptilia: Colubridae). Brimleyana 19: 83-94. Scudder Davis, R.M.; Burghardt, G.M. 1996. Ontogenetic changes in growth efficiency in laboratory-reared water snakes of the genus Nerodia. The Snake 27(2): 75-84. Scudder, R.M.; Burghardt, G.M. 1983. A comparative study of defensive behaviour in three sympatric species of water snakes (Nerodia). Zeitschrift für Tierpsychologie 63(1): 17-26. Trauth, S.E. 1990. Flooding as a factor in the decimation of a population of green water snakes (Nerodia cyclopion cyclopion) from Arkansas. Bulletin of the Chicago Herpetological Society 25(1): 1-3. Trauth, S.E.; Neasbitt, W.H. 1987. Geographic distribution: Nerodia cyclopion cyclopion (Green Water Snake). Herpetological Review 18(1): 20. Werler, J.E. 1948. Natrix cyclopion cyclopion in Texas. Herpetologica 4: 148. Nerodia erythrogaster Adler, K.K. 1963. An ecological study of the snake Natrix erythrogaster, with remarks on its postglacial dispersal. Journal Ohio Herpetological Society 4(1/2): 61-62. Arbaugh, R.; Arbaugh, T.; Steen, D.A. 2015. Natural history notes: Nerodia erythrogaster (Plain-bellied Watersnake). Predation. Herpetological Review 46(4): 647-648. Attum, O.; Lee, Y.M.; Kingsbury, B.A. 2009. The status of the northern population of the Copper-bellied Watersnake, Nerodia erythrogaster neglecta. Northeastern Naturalist 16(3): 317-320. Attum, O.; Lee, Y.M.; Roe, J.H.; Kingsbury, B.A. 2008. Wetland complexes and upland-wetland linkages: landscape effects on the distribution of rare and common wetland reptiles. Journal of Zoology (London) 275(3): 245-251. Beane, J.C.; Thorp, T.J. 2008. Geographic distribution: Nerodia erythrogaster erythrogaster (Red-bellied Watersnake). Herpetological Review 39(2): 242. Belford, T.J. 2015. Geographic distribution: Nerodia erythrogaster (Plain-bellied Watersnake). Herpetological Review 46(3): 386. Boundy, J.; David, P. 2015. The taxonomic status of the snake name Tropidonotus roulei Chabannaud, 1917. Herpetological Review 46(2): 295-297. Bousefield, S. 1982. Observations on the yellow-bellied water-snake (Nerodia erythrogaster flavigaster). Herptile 7(4): 23-24. Browning, M.J.S. 1967. Natrix erythrogaster erythrogaster in the Piedmont region of South Carolina. Herpetologica 23: 59. Brumwell, M.J. 1951. An ecological survey of the Fort Leavenworth Military Reservation. American Midland Naturalist 45(1): 187-231. Bufalino, A.P.; Scott, A.F. 2003. The distribution of Nerodia erythrogaster in the Lower Cumberland River Basin of Kentucky and Tennessee. Herpetological Review 34(1): 77-78. Bufalino, A.P.; Scott, A.F. 2008. Statistical analyses of pattern variation in the copper-bellied (Nerodia erythrogaster neglecta) and yellow-bellied (N. e. flavigaster) watersnake subspecies. American Midland Naturalist 159(1): 30-41. Bustard, R. 1959. Hardy snakes. Aquarist and Pondkeeper 24: 49-50. Byrd, W.; Hanebrink, E.; Meshaka, W. 1988. Food, feeding behaviour, sex ratios and measurements of three species of water snakes (Nerodia spp.) collected from northeastern Arkansas. Bulletin of the Chicago Herpetological Society 23(4): 55-57. Camper, J.D. 2009. Core terrestrial habitat around wetlands: contributions from the spatial ecology of the redbelly watersnake (Nerodia erythrogaster erythrogaster). Copeia 2009(3): 556-562. Cantu-Martínez, J.M.; Quintero, E.; Zaidan, F.; Harrison, C.R. 2011. Geographic distribution: Nerodia erythrogaster transversa (Blotched Watersnake). Herpetological Review 42(3): 395. Carpenter, C.C. 1958. Reproduction, young, eggs and food of Oklahoma snakes. Herpetologica 14: 113-115. Chabanaud, P. 1917. Descriptions de trois espèces nouvelles de reptiles de l'Afrique. Bulletin du Muséum National d'Histoire Naturelle (Paris) 23(1): 219-225. Christiansen, J.L.; LeClere, J.B. 2002. Nerodia erythrogaster neglecta in Iowa with comments on ontogenesis of color and pattern. Herpetological Review 33(2): 152-154. Christman, B.L.; Wallace, J.E.; Bartel, S.C. 2000. Geographic distribution: Nerodia erythrogaster transversa (Blotched Water Snake). USA: Texas: Presidio Co. Herpetological Review 31(1): 56. Clay, W.M. 1934. Rediscovery of the red-bellied watersnake, Natrix sipedon erythrogaster (Forster) in Michigan. Copeia 1934(2): 95-96. Colvin, R.; Dennison, J. 2012. Geographic distribution: Nerodia erythrogaster flavigaster (Yellow-bellied Watersnake). Herpetological Review 43(1): 106. Conant, R. 1934. The red-bellied water snake, Natrix sipedon erythrogaster (Forster) in Ohio. Ohio Journal of Science 34(1): 21-28. Conant, R. 1943. Natrix erythrogaster erythrogaster in the northeastern part of its range. Herpetologica 11(2): 83-86. Conant, R. 1949. Two new races of Natrix erythrogaster. Copeia 1949: 1-15. Conant, R. 1953. Three new water snakes of the genus Natrix from Mexico. Natural History Miscellanea 126: 1-9. Conant, R. 1955. Notes on Natrix erythrogaster from the eastern and western extremes of its range. Natural History Miscellanea 147: 1-3. Conant, R. 1955. Two new Ohio localities for the red-bellied water snake. Ohio Journal of Science 55(1): 61-62. Conant, R. 1960. Water snakes in the desert. America's First Zoo 12: 31-35. Conant, R. 1963. Another new water snake of the genus Natrix from the Mexican plateau. Proceedings of the Biological Society of Washington 76: 169-172. Conant, R. 1965. Notes on reproduction in two natricine snakes from Mexico. Herpetologica 21(2): 140-144. Connior, M.B. 2010. Geographic distribution: Nerodia erythrogaster flavigaster (Yellow-bellied Watersnake). Herpetological Review 41(4): 517. Cooper, J.E. 1969. A red-bellied watersnake from Maryland's western coastal plain. Journal of Herpetology 3: 185-186. Daniel, R.E. 2006. New state size records for Missouri reptiles. Missouri Herpetological Association Newsletter 19: 14-15. Diener, R.A. 1957. An ecological study of the plain-bellied water snake. Herpetologica 13: 203-211. Dowling, H.G. 1956. Geographic relations of Ozarkian amphibians and reptiles. Southwestern Naturalist 1(4): 174-189. Duke, S.T. 2010. Minimum habitat requirements of the Copper-bellied Watersnake (Nerodia erythrogaster neglecta). Bulletin of the Chicago Herpetological Society 45(2): 29-31. Dunn, E.R.; Allen, R. 1935. The red-bellied watersnake in Pennsylvania. Copeia 1935: 180-181. Eubanks, J.T.; Pecor, K.W. 2003. Natural history notes: Nerodia erythrogaster flavigaster, N. rhombifer, and N. sipedon pleuralis (Yellow-bellied, Diamond-backed, and Midland Watersnakes). Foraging behavior. Herpetological Review 34(1): 70. Freed, P.S. 1983. Life history notes: Nerodia erythrogaster flavigaster (Yellowbelly Water Snake). Coloration. Herpetological Review 14(3): 75. Freeman, H.W. 1955. Part V. The amphibia and reptilia of the Savannah river project area. 2. Chelonia. 3. Crocodilia, Sauria and Serpentes. U.S. Cur. Publ. Biol. 1(4): 239-244, 275-291. Gillingham, J.C.; Rush, T. 1974. Notes on the fishing behaviour of water snakes. Journal of Herpetology 8(4): 384-385. Greding, E.J. 1964. Food of matrix in Hunt County, Texas. Southwestern Naturalist 9: 206. Grogan, W.L. 1985. New distribution records for Maryland reptiles and amphibians. Bulletin of the Maryland Herpetological Society 21(2): 74-75. Hampton, P.M. 2007. Natural history notes: Nerodia erythrogaster flavigaster (Yellow-bellied Watersnake). Anti-predatory behavior. Herpetological Review 38(1): 91. Hampton, P.M. 2007. Natural history notes: Nerodia erythrogaster flavigaster (Yellow-bellied Watersnake). Diet. Herpetological Review 38(1): 91. Hampton, P.M.; Ford, N.B. 2007. Effects of flood suppression on natricine snake diet and prey overlap. Canadian Journal of Zoology 85(7): 809-814. Hill, R.L. 2012. Geographic distribution: Nerodia erythrogaster erythrogaster (Red-bellied Watersnake). USA: Georgia: Gwinnett Co. Herpetological Review 43(4): 621. Hill, R.L.; Lock, B.; Brothers, D.; Camdelyn, R. 2012. Geographic distribution: Nerodia erythrogaster erythrogaster (Red-bellied Watersnake). USA: Georgia: Dekalb Co. Herpetological Review 43(4): 621. Hofmann, E.P. 2018. Natural history notes: Nerodia erythrogaster (Plain-bellied Watersnake). Defensive behavior. Herpetological Review 49(1): 135. Irwin, K.J. 2002. Geographic distribution: Nerodia erythrogaster (Plain-bellied Watersnake). Herpetological Review 33(4): 325. Kingsbury, B.A.; Grimes, R.A. 2001. A description and assessment of the copperbelly Water Snake conservation agreement. Transactions of the North American Wildlife and Natural Resources Conference 66: 538-554. Kofron, C.P. 1979. Reproduction of aquatic snakes in south central Louisiana. Herpetologica 35(1): 44-50. Lacki, M.J.; Hummer, J.W.; Fitzgerald, J.L. 1994. Application of line transects for estimating population density of the endangered Copperbelly Water Snake in southern Indiana. Journal of Herpetology 28(2): 241-245. Lacki, M.J.; Hummer, J.W.; Fitzgerald, J.L. 2005. Population patterns of copperbelly Water Snakes (Nerodia erythrogaster neglecta) in a riparian corridor impacted by mining and reclamation. American Midland Naturalist 153(2): 357-369. Lamer, J.T.; Jenkins, S.E.; Tucker, J.K.; Dolan, C.R. 2008. Geographic distribution: Nerodia erythrogaster (Plain-bellied Watersnake). Herpetological Review 39(2): 242. Laposha, N.A.; Parmerlee, J.S.; Powell, R.; Smith, D.D. 1985. Life history notes: Nerodia erythrogaster transversa (Blotched Water Snake). Reproduction. Herpetological Review 16(3): 81. Laughlin, H.E. 1959. Stomach contents of some aquatic snakes from Lake McAlester, Pittsburgh County, Oklahoma. Texas Journal of Science 11(1): 83-85. Laurent, E.J.; Kingsbury, B.A. 2003. Habitat separation among three species of Water Snakes in northwestern Kentucky. Journal of Herpetology 37(2): 229-235. Lutterschmidt, W.I.; Schaefer, J.F. 1996. Mist netting: adapting a technique from ornithology for sampling semi-aquatic snake populations. Herpetological Review 27(3): 131-132. Makowsky, R.; Marshall, J.C.; McVay, J.; Chippindale, P.T.; Rissler, L.J. 2010. Phylogeographic analysis and environmental niche modeling of the Plain-bellied Watersnake (Nerodia erythrogaster) reveals low levels of genetic and ecological differentiation. Molecular Phylogenetics and Evolution 55(3): 985-995. Marshall, J.C.; Kingsbury, B.A.; Minchella, D.J. 2009. Microsatellite variation, population structure, and bottlenecks in the threatened copperbelly Water Snake. Conservation Genetics 10(2): 465-476. McAllister, C.T. 1980. Geographic distribution: Nerodia erythrogaster transversa. Herpetological Review 11(1): 15. McCoid, M.J. 1999. 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