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Taxonomy of the genus Regina Biodiversity of the family Colubridae

Bibliography of the genus
Regina (Queen Snakes)

(Reptilia: Serpentes: Colubridae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Regina in general

Alfaro, M.E.; Arnold, S.J. 2001. Molecular systematics and evolution of Regina and the thamnophiine snakes. Molecular Phylogenetics and Evolution 21(3): 408-423.

Dwyer, C.M.; Kaiser, H. 1997. Relationship between skull form and prey selection in the thamnophiine snake genera Nerodia and Regina. Journal of Herpetology 31(4): 463-475.

Eberle, W.G. 1972. Comparative chromosomal morphology of the New World natricine snake genera Natrix and Regina. Herpetologica 28(2): 98-105.

Ernst, C.H.; Whitfield-Gibbons, J.; Dorcas, M.E. 2002. Regina. Catalogue of American Amphibians and Reptiles 756: 756.1-756.4.

Gibbons, J.W.; Dorcas, M.E. 2004. North American watersnakes. A natural history. University of Oklahoma, Norman. 496 pp.

Godley, J.S.; McDiarmid, R.W.; Rojas, N.N. 1984. Estimating prey size and number in crayfish-eating snakes, genus Regina. Herpetologica 40(1): 82-88.

Hoser, R.T. 2012. Two new genera of water snake from North America. The subdivision of the genera Regina Baird and Girard, 1853 and Nerodia Baird and Girard, 1853 (Serpentes: Colubridae: Natricinae). Australasian Journal of Herpetology 11: 28-31.

International Commission on Zoological Nomenclature 1962. Opinion 632. Regina Baird & Girard, 1853 (Reptilia); designation of a type-species under the plenary powers. Bulletin of Zoological Nomenclature 19: 146-147.

Malnate, E.V. 1960. Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae. Proceedings of the Academy of Natural Sciences of Philadelphia 112: 41-71.

Mara, W.P. 1995. Water Snakes of North America. T.F.H. Publications, Neptune City, New Jersey. 64 pp.

McVay, J.D.; Carstens, B. 2013. Testing monophyly without well-supported gene trees: evidence from multi-locus nuclear data conflicts with existing taxonomy in the snake tribe Thamnophiini. Molecular Phylogenetics and Evolution 68(3): 425-431.

McVay, J.D.; Flores-Villela, O.; Carstens, B. 2015. Diversification of North American natricine snakes. Biological Journal of the Linnean Society 116(1): 1-12.

Price, R. 1983. Microdermatoglyphics: the Liodytes-Regina problem. Journal of Herpetology 17(3): 292-294.

Rossman, D.A. 1963. Relationships and taxonomic status of the North American natricine snake genera Liodytes, Regina, and Clonophis. Occasional Papers Museum of Zoology Louisiana State University 29: 1-29.

Ruthven, A.G.; Thompson, C. 1913. The variations in the number of vertebrae and ventral scutes in two snakes of the genus Regina. American Naturalist 47: 625-632.

Smith, H.M.; Huheey, J.E. 1960. The watersnake genus Regina. Transactions of the Kansas Academy of Sciences 63: 156-164.

Regina grahamii

Baker, F.C.; Woodruff, F.M. 1897. Note on Natrix grahamii, B. & G. Science 2(2)(5): 447.

Baker, S.J.; Kirk, D.; Schmidt, C.E.; Dreslik, M.J. 2007. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 38(1): 105-106.

Byrd, W.W.; Hanebrink, E.L.; Files, C. 1989. Geographic distribution: Regina grahami (Graham's Crayfish Snake). Herpetological Review 20(1): 13.

Byrd, W.W.; Hanebrink, E.L.; Files, C. 1990. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 21(2): 41.

Cleveland, E.D. 1986. County record for Graham's crawfish snake (Regina grahami). Transactions of the Kansas Academy of Science 89(1-2): 9.

Coleman, J.L.; Pruett, J.A. 2008. New county records of reptiles and amphibians from East Texas. Herpetological Review 39(2): 246-247.

Collins, J.T.; Collins, S.L. 2009. Regina grahamii (Graham's Crayfish Snake). Kansas. Journal of Kansas Herpetology 30: 11.

Cox, C.L.; Farrar, E.S.; Hey, J.D. 2005. Geographic distribution: Regina grahami (Graham's Crayfish Snake). Herpetological Review 36(3): 340.

Dowling, H.G. 1956. Geographic relations of Ozarkian amphibians and reptiles. Southwestern Naturalist 1(4): 174-189.

Edmond, B.S. 1993. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 24(2): 68.

Goodman, J.D. 1948. A report on the reptiles collected by J.M. Shaffer from the Keokuk Area, 1863-1895. Proceedings of the Iowa Academy of Science 55: 365-366.

Guthrie, J.E. 1930. A color mutation in Graham's watersnake. Journal Hered. Washington 21: 237-240.

Guthrie, J.E. 1930. Color dimorphism in Graham's watersnake, Natrix grahami (Baird & Girard). Copeia 1930: 39.

Hall, R.J. 1969. Ecological observations on Graham's water-snake (Regina grahami Baird and Girard). American Midland Naturalist 81: 156-163.

Hibbitts, T.J.; Adams, C.S. 2015. Geographic distribution: Regina grahamae (Graham's Crayfish Snake). Herpetological Review 46(2): 221.

Jones, M.C.; Reilly, S.; Forstner, M.R.J.; Dixon, J.R. 2005. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 36(4): 470.

Killebrew, F.C.; Rhea, M. 1980. Geographic distribution: Regina grahami. Herpetological Review 11(2): 39.

Kofron, C.P. 1979. Reproduction of aquatic snakes in south central Louisiana. Herpetologica 35(1): 44-50.

Kofron, C.P.; Dixon, J.R. 1980. Observations on aquatic colubrid snakes in Texas. Southwestern Naturalist 25(1): 107-109.

Landwer, A.J.; Lee, T.E. 2001. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 32(2): 124.

Laughlin, H.E. 1959. Stomach contents of some aquatic snakes from Lake McAlester, Pittsburgh County, Oklahoma. Texas Journal of Science 11(1): 83-85.

Liner, E.A. 1982. Life history notes: Regina grahami (Graham's Crayfish Snake). Coloration. Herpetological Review 13(2): 48.

Manjarrez, J.; Macias García, C.; Drummond, H. 2017. Morphological convergence in a Mexican garter snake associated with the ingestion of a novel prey. Ecology and Evolution 7(18): 7178-7186

McAllister, C.T.; Bibb, D.K. 1991. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 22(2): 68.

Miner, R.N. 1959. A size record for Graham's water snake. Herpetologica 15: 234.

Muelleman, P.J.; Montgomery, C.E. 2011. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 42(2): 244.

Mushinsky, H.R.; Hebrard, J.J. 1977. Food partitioning by five species of water snakes in Louisiana. Herpetologica 33(2): 162-166.

Mushinsky, H.R.; Hebrard, J.J. 1977. The use of time by sympatric water snakes. Canadian Journal of Zoology 55(9): 1545-1550.

Mushinsky, H.R.; Hebrard, J.J.; Walley, M.G. 1980. The role of temperature on the behavioral and ecological associations of sympatric water snakes. Copeia 1980(4): 744-754.

Nakamure, E.L.; Smith, H. 1960. A comparative study of selected characters in certain American species of watersnakes. Transactions of the Kansas Academy of Sciences 63: 102-113.

Outline, J.E. 1930. Freak color inheritance in Graham's water snake. Proceedings of the Iowa Academy of Science 32: 375-377.

Ramsey, L.W. 1954. A possible range extension for Natrix grahami. Herpetologica 10: 188.

Ramsey, Z.D.; McAllister, C.T. 2004. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 35(3): 294.

Schmidt, C. 2003. Regina grahamii (Graham's Crayfish Snake). Journal of Kansas Herpetology 6: 8.

Seigel, R.A. 1992. Ecology of a specialized predator: Regina grahami in Missouri. Journal of Herpetology 26(1): 32-37.

Thornton, O.W.; Smith, J.R. 1995. Natural history notes: Regina grahami (Graham's Crayfish Snake). Reproduction. Herpetological Review 26(2): 102-103.

Walley, H.D. 1977. Geographic distribution: Regina grahami. USA: Illinois: Carroll Co. Herpetological Review 8(2): 39.

Walston, L.J.; Walston, L.J. 2008. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 39(3): 373.

Waters, R.M.; Burghardt, G.M. 2005. The interaction of food motivation and experience in the ontogeny of chemoreception in crayfish snakes. Animal Behaviour 69(2): 363-374.

Williams, A.A. 1997. Geographic distribution: Regina grahamii (Graham's Crayfish Snake). Herpetological Review 28(2): 98.

Wilson, A.K. 1999. New Illinois amphibian and reptile distribution records from the Kaskaskia River drainage. Herpetological Review 30(2): 118-120.

Young, R.A. 1973. Notes on the Graham's water snake, Regina grahami Baird and Girard in DuPage County, Illinois. Bulletin Chicago Herpetological Society 8(1-2): 4-5.

Regina septemvittata

Adler, K.K. 1962. Life history studies of the queen snake, Natrix septemvittata (Say), in central Ohio. Journal Ohio Herpetological Society 3(4): 60.

Adler, K.K.; Tilley, S.G. 1960. A fish and a snail in the diet of Natrix septemvittata (Say). Journal Ohio Herpetological Society 2(4): 28-29.

Allen, K. 2003. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 34(2): 170.

Anton, T.G.; Redmer, M. 1991. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 22(1): 26.

Arndt, R.G. 1985. Distributional notes on some Delaware snakes, with two first records for the state. Bulletin of the Maryland Herpetological Society 21(2): 67-73.

Arndt, R.G. 1994. Geographic distribution: Regina septemvittata (Queen Snake). USA: South Carolina: Barnwell Co. Herpetological Review 25(4): 168.

Barton, L.J.; Morningstar, L.L.; Hall, J.M. 2015. Geographic distribution: Regina septemvittata (Queensnake). Herpetological Review 46(4): 576.

Bekker, K.; Timar, J. 2004. Natural history notes: Regina septemvittata (Queen Snake). Diet. Herpetological Review 35(1): 72.

Bradford, J.W.; Butterfield, B.P. 2016. Geographic distribution: Regina septemvittata (Queensnake). Herpetological Review 47(3): 428-429.

Branson, B.A.; Baker, E.C. 1973. Sexual dimorphism in Kentucky queen snakes (Regina septemvittata based on scute counts. Transactions of the Kentucky Academy of Science 34(3-4): 57-58.

Branson, B.A.; Baker, E.C. 1974. An ecological study of the queen snake, Regina septemvittata (Say) in Kentucky. Tulane Studies in Zoology and Botany 18(4): 153-171.

Butler, J.A. 1977. The physiological and behavioural mechanisms that relate to feeding strategies of Regina septemvittata during environmental temperature extremes. Herpetological Review 8(3): 3.

Christensen, T.; Christensen, M. 2006. Regina septemvittata (Queen Snake). Catesbeiana 26(2): 78.

Cliburn, J.W. 1975. Natrix septemvittata in Tishomingo County, Mississippi. Journal of the Mississippi Academy of Sciences 19: 159-160.

Conant, R. 1960. The queen snake, Natrix septemvittata, in the interior highlands of Arkansas and Missouri, with comments upon similar disjunct distributions. Proceedings of the Academy of Natural Sciences of Philadelphia 112: 25-40.

Davis, J.G. 2009. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 40(4): 457.

Dowling, H.G. 1956. Geographic relations of Ozarkian amphibians and reptiles. Southwestern Naturalist 1(4): 174-189.

Dundee, H.A. 2003. Natural history notes: Regina septemvittata (Queen Snake). Aberrant pattern. Herpetological Review 34(1): 72.

Ernst, C.H. 2002. Regina septemvittata. Catalogue of American Amphibians and Reptiles 757: 757.1-757.5.

Ernst, C.H. 2003. Natural history of the Queen Snake, Regina septemvittata, in southeastern Pennsylvania, U.S.A. Herpetological Bulletin 85: 2-11.

Finkler, M.S.; Claussen, D.L. 1999. Influence of temperature, body size, and inter-individual variation on forced and voluntary swimming and crawling speeds in Nerodia sipedon and Regina septemvittata. Journal of Herpetology 33(1): 62-71.

Fontenot, L.W.; Platt, S.G. 1996. Natural history notes: Regina septemvittata (Queen Snake). Reproduction. Herpetological Review 27(4): 205.

Ford, N.B. 1982. Courtship behavior of the Queen Snake, Regina septemvittata. Herpetological Review 13(3): 72.

Gerald, G.W. 2006. Natural history notes: Regina septemvittata (Queen Snake). Predation. Herpetological Review 37(4): 480.

Hayslett, M.; Scranton, J. 1995. Regina septemvitta [septemvittata] (Queen Snake). Catesbeiana 15(2): 48.

Herman, D.W.; Johnson, N. 1986. Life history notes: Regina septemvittata (Queen Snake). Dicephalism. Herpetological Review 17(1): 22.

Hromada, S.J. 2017. Geographic distribution: Regina septemvittata (Queensnake). Herpetological Review 48(2): 392.

Jackrel, S.L.; Reinert, H.K. 2011. Behavioral responses of a dietary specialist, the Queen Snake (Regina septemvittata) to potential chemoattractants released by its prey. Journal of Herpetology 45(3): 272-276.

Johnston, B.L. 2004. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 35(1): 84.

Judd, W.W. 1956. Observations on the habitat and food of the queen snake, Natrix septemvittata, at London, Ontario. Canadian Field Naturalist 69(4): 167-168.

Kapfer, J.M.; Mueller, W.P.; Porzky, M.M.; Casper, G.S. 2009. Geographic distribution: Regina septemvittata (Queen Snake). USA: Wisconsin. Herpetological Review 40(1): 116.

Klauber, L.M. 1948. Some misapplications of the Linnaean names applied to American snakes. Copeia 1948: 1-14.

Lacy, G.B. 1996. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 27(3): 154-155.

Layne, J.R.; Ford, N.B. 1984. Flight distance of the Queen Snake, Regina septemvittata. Journal of Herpetology 18(4): 496-498.

Lee, D.S. 1974. Another record for the queen snake in southern Maryland. Bulletin of the Maryland Herpetological Society 9(4): 107.

Lipps, G.; Davis, J.G. 2014. Geographic distribution: Regina septemvittata (Queensnake). Herpetological Review 45(1): 96.

Manjarrez, J.; Macias García, C.; Drummond, H. 2017. Morphological convergence in a Mexican garter snake associated with the ingestion of a novel prey. Ecology and Evolution 7(18): 7178-7186

McBride, B. 1967. Notes on three rare Ontario snakes. Bull. Can. Amph. Rept. Conserv. Soc. 5(6): 1-3.

Merkle, D. 1987. Field notes. New county records for the Queen Snake Regina septemvittata in the central piedmont of Virginia. Catesbeiana 7(2): 19-20.

Mitchell, J.C. 2005. Regina septemvittata (Queen Snake). Catesbeiana 25(1): 34.

Mitchell, J.C. 2005. Status of the Queen Snake (Regina septemvittata) on the University of Richmond campus. Catesbeiana 25(1): 24-25.

Mitchell, J.C.; Rouse, G. 2005. Regina septemvittata (Queen Snake). Catesbeiana 25(1): 34-35.

Nakamure, E.L.; Smith, H. 1960. A comparative study of selected characters in certain American species of watersnakes. Transactions of the Kansas Academy of Sciences 63: 102-113.

Neill, W.T. 1941. A dicephalic queen snake. Copeia 1941: 266.

Neill, W.T. 1963. A new subspecies of the queen snake Natrix septemvittata from southern Alabama. Herpetologica 19: unpaginated.

Newcomer, R.T.; Taylor, D.H.; Guttman, S.I. 1974. Celestial orientation in two species of water snakes (Natrix sipedon and Regina septemvittata). Herpetologica 30(2): 194-200.

Oldham, C.R.; Price, S.J.; Boys, W.A.; Fleckenstein, L.J. 2015. Natural history notes: Regina septemvittata (Queensnake). Defensive behavior / death-feigning. Herpetological Review 46(2): 276-277.

Petzing, J.E.; Phillips, C.A. 1998. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 29(2): 115.

Pierson, T.W. 2012. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 43(2): 308.

Pinder, M.J. 1996. Regina septemvittata (Queen Snake). Catesbeiana 16(1): 9.

Plummer, M.V. 1980. Geographic distribution: Regina septemvittata. Herpetological Review 11(3): 81.

Plummer, M.V. 2016. Geographic distribution: Regina septemvittata (Queensnake). Herpetological Review 47(2): 264.

Plummer, M.V.; Goy, J.M. 2016. Geographic distribution: Regina septemvittata (Queensnake). Herpetological Review 47(2): 264.

Raney, E.C.; Roecker, R.M. 1947. Food and growth of two species of watersnakes from western New York. Copeia 1947: 171-174.

Ray, W.J. le 1928. The queen snake Natrix septemvittata in Ontario. Canadian Field Naturalist 42: 42.

Reid, S.M.; Nocera, J.J. 2015. Composition of native crayfish assemblages in southern Ontario rivers affected by Rusty Crayfish (Orconectes rusticus Girard, 1852) invasions - implications for endangered Queensnake recovery. Aquatic Invasions 10(2): 189-198.

Reinert, H.K. 1975. Another winter record of a snake (Natrix septemvittata). Bulletin Philadelphia Herpetological Society 23: 7.

Roble, S.M. 1995. Regina septemvittata (Queen Snake). Catesbeiana 15(1): 24-25.

Scott, A.F.; Zirkle, E. 1992. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 23(1): 27.

Spangler, J.A.; Mount, R.H. 1969. The taxonomic status of the natricine snake Regina septemvittata mabila (Neill). Herpetologica 25: 113-119.

Spinner, L. 2017. Long live the Queen! Lebensweise und Haltung der Königinnennatter. Terraria-Elaphe 2017(63): 38-42.

Spurr, R.H.; Smith, D.C. 1980. Sights of Queen Snakes along the Ausable River. Bulletin Canadian Amphibian and Reptile Conservation Society 17(3): 1-4.

Stanley, J.W.; Trauth, S.E. 2007. Distribution of the Queen Snake (Regina septemvittata) in Arkansas. Journal of the Arkansas Academy of Science 61: 99-103.

Thesing, B.J.; Clause, A.G. 2018. Geographic distribution: Filling gaps with specimens: new herpetofaunal county records from Georgia, USA. Herpetological Review 49(1): 80-84.

Trauth, S.E. 1988. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 19(1): 20.

Trauth, S.E. 1991. Distribution, scutellation, and reproduction in the Queen Snake, Regina septemvittata (Serpentes: Colubridae), from Arkansas. Arkansas Academy of Science Proceedings 45: 103-106.

Triplehorn, C.A. 1946. A large specimen and a high embryo count for the queen snake. Copeia 1946(1): 76.

Tuck, R.G. 1974. A craniodichotomous queensnake, Regina septemvittata (Say) (Serpentes: Natricidae), from Frederick County, Maryland. Bulletin of the Maryland Herpetological Society 9(2): 22-23.

Waters, R.M.; Burghardt, G.M. 2005. The interaction of food motivation and experience in the ontogeny of chemoreception in crayfish snakes. Animal Behaviour 69(2): 363-374.

Williams, M.; Foster, N. 2012. Geographic distribution: Regina septemvittata (Queensnake). USA: Tennessee: Maury Co. Herpetological Review 43(3): 450.

Williamson, S. 2001. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 32(4): 279.

Wood, J.T. 1944. Fall aggregation of the queen snake. Copeia 1944: 253.

Wood, J.T. 1949. Observations on Natrix septemvitata (Say) in south western Ohio. American Midland Naturalist 42(3): 744-750.

Wood, J.T.; Duellman, W.E. 1950. Size and scutellation in Natrix septemvittata (Say) in southwestern Ohio. American Midland Naturalist 43(1): 173-178.

Wright, R.; Wagamon, W. 2002. Regina septemvittata (Queen Snake). Catesbeiana 22(2): 55.

Wusterbarth, T.L.; King, R.B.; Duvall, M.R.; Grayburn, W.S.; Burghardt, G.M. 2010. Phylogenetically widespread multiple paternity in New World natricine snakes. Herpetological Conservation and Biology 5(1): 86-93.

Young, R.A.; Vandeventer, T.L.; Jones, R. 1988. Geographic distribution: Regina septemvittata (Queen Snake). Herpetological Review 19(3): 60.