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Taxonomy of the genus Uma Biodiversity of the family Phrynosomatidae

Bibliography of the genus Uma (North American Fringe-toed Lizards)

(Reptilia: Sauria: Phrynosomatidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Uma in general

Adest, G.A. 1977. Genetic relationships in the genus Uma (Iguanidae). Copeia 1977(1): 47-52.

Adest, G.A. 1978. The relations of the sand lizards Uma, Callisaurus and Holbrookia (Sauria: Iguanidae): an electrophoretic study. Ph.D. dissertation, University of California, Los Angeles.

Carpenter, C.C. 1963. Patterns of behaviour in three farms of the fringe-toed lizards (Uma-Iguanidae). Copeia 1963: 406-412.

Carpenter, C.C. 1967. Display patterns of the Mexican iguanid lizards of the genus Uma. Herpetologica 23: 285-293.

Cope, E.D. 1894. On the iguanian genus Uma, Baird. American Naturalist 28: 434-435.

Cope, E.D. 1895. On the species of Uma and Xantusia. American Naturalist 29: 938-939.

Cornett, J. 1983. A masterpiece of adaptation. Uma, the fringe toed sand lizard. Pacific Discovery 36(2): 2-10.

Cox, D.C.; Tanner, W.W. 1977. Osteology and myology of the head and neck regions of Callisaurus, Cophosaurus, Holbrookia and Uma (Reptilia: Iguanidae). Great Basin Naturalist 37(1): 35-56.

Earle, A.M. 1962. The middle ear of the genus Uma compared to those of the other sand lizards. Copeia 1962: 185-188.

Heifetz, W. 1941. A review of the lizards of the genus Uma. Copeia 1941(2): 99-111.

Lannom, J.R. 1962. A different method of catching the desert lizards, Callisaurus and Uma. Copeia 1962: 437-438.

Mayhew, W.W. 1961. Photoperiodic response of female fringe-toed lizards. Science 134: 2104-2105.

Mayhew, W.W. 1964. Taxonomic status of California populations of the lizard genus Uma. Herpetologica 20: 170-183.

Mayhew, W.W. 1967. Comparative reproduction in three species of the genus Uma. pp. 45-61. In: Milstead, W.W. (ed.). Lizard ecology - a symposium. University of Missouri Press, Columbia, Missouri. 300 pp.

Morafka, D.J.; Adest, G.A.; Reyes, L.M.; Aguirre-Leon, G.; Lieberman, S.S. 1992. Differentiation of North American deserts: a phylogenetic evaluation of a vicariance model. Tulane Studies in Zoology and Botany Supplementary Publication 1: 195-226.

Norris, K.S. 1958. The evolution and systematics of the iguanid genus Uma and its relation to the evolution of other North American desert reptiles. Bulletin of the American Museum of Natural History 114: 247-326.

Pough, F.H. 1969. Physiological aspects of the burrowing of sand lizards (Uma, Iguanidae) and other lizards. Comparative Biochemistry and Physiology 31: 869-884.

Pough, F.H. 1969. The biology of burrowing in the sand lizard, Uma. Dissertation Abstracts International (B) 29: 4446-4447.

Shaw, C.E. 1950. Lizards in the diet of captive Uma. Herpetologica 6(2): 36-37.

Stebbins, R.C. 1943. Adaptations in the nasal passages for sand burrowing in the saurian genus Uma. American Naturalist 77(768): 38-52.

Stebbins, R.C. 1944. Some aspects of the ecology of the iguanid genus Uma. Ecological Monographs (Durham) 14(3): 311-313.

Switak, K.H. 1979. Leben in der Wüste. Die Fransenzehen-Leguane der Gattung Uma Baird. Aquarium (Bornheim) 13(123): 409-411.

Wilgenbusch, J.; Queiroz, K. de 2000. Phylogenetic relationships among the phrynosomatid Sand Lizards inferred from mitochondrial DNA sequences generated by heterogeneous evolutionary processes. Systematic Biology 49(3): 592-612.

Zalusky, S.B.; Gaudin, A.J.; Swanson, J.R. 1980. A comparative study of cranial osteology in the North American sand lizards, genus Uma (Reptilia: Iguanidae). Copeia 1980(2): 296-310.

Uma cowlesi

Shaw, C.E. 1952. Notes on the eggs and young of some United States and Mexican lizards. 1. Herpetologica 8(3): 71-79.

Smith, P.W.; Hensley, M.M. 1959. Notes on a small collection of amphibians and reptiles from the vicinity of the Pinacate lava cap in north-western Sonora, Mexico. Transactions of the Kansas Academy of Sciences 61: 64-76.

Uma exsul

Ballesteros-Barrera, C.; Martinez-Meyer, E.; Gadsden, H. 2007. Effects of land-cover transformation and climate change on the distribution of two microendemic lizards, genus Uma, of northern Mexico. Journal of Herpetology 41(4): 733-740.

Commins, M.L.; Savitzky, A.H. 1973. Fiels observations on a population of the sand lizard Uma exsul. Journal of Herpetology 7(1): 51-53.

Cooper, W.E.; Castaneda, G.; Pena, C.G. de la 2006. Phylogenetic constraints do not block food chemical discrimination in the omnivorous phrynosomatid lizard Uma exsul. Journal of Herpetology 40(3): 329-335.

Cooper, W.E.; Castañeda-Gaytan, J.G.; García-de la Peña, C.; Wilson, B.S.; Caldwell, J.P.; Vitt, L.J. 2013. Foraging modes of some Jamaican, Costa Rican, and Mexican lizards. Herpetology Notes 6: 591-597.

Estrada-Rodriguez, J.L.; Gadsden, H.; Pacheco, S.V.L.; Lopez-Corrujedo, H. 2004. A new capture technique for the Coahuila Fringe-toed Lizard (Uma exsul) and other desert lizards. Herpetological Review 35(3): 244-245.

Estrada-Rodriguez, J.L.; Leyva-Pacheco, S.V. 2007. Natural history notes: Uma exsul (Coahuila Fringe-toed Lizard). Swimming. Herpetological Review 38(1): 84-85.

Gadsden, H.; Carrazco, M.L.D.; Gil Martinez, R. 2006. Reproduction in the arenicolous Mexican lizard Uma exsul. Journal of Herpetology 40(1): 117-122.

Gadsden, H.; Palacios-Orona, L.E.; Cruz-Soto, G.A. 2001. Diet of the Mexican fringe-toed lizard (Uma exsul). Journal of Herpetology 35(3): 493-496.

Gadsden-Esparza, H.; López-Corrujedo, H.; Estrada-Rodríguez, J.L.; Romero-Méndez, U. 2001. Biología poblacional de la lagartija de arena de Coahuila Uma exsul (Sauria: Phrynosomatidae): implicaciones para su conservación. Boletin de la Sociedad Herpetologica Mexicana 9(2): 51-66.

Garcia-de la Pena, C.; Castaneda, G.; Gadsden, H.; Contreras-Balderas, A.J. 2007. Niche segregation within a dune lizard community in Coahuila, Mexico. Southwestern Naturalist 52(2): 251-257.

Garcia-de la Pena, C.; Castaneda-Gaytan, G.; Lazcano, D.; Rosales, A.J. 2003. Natural history notes: Uma exsul (Fringe-toed Sand Lizard). Predation. Herpetological Review 34(4): 370.

Garcia-de la Pena, C.; Gadsden, H.; Contreras-Balderas, A.J.; Castaneda, G. 2007. Ciclos de actividad diaria y estacional de un gremio de saurios en las dunas de arena de Viesca, Coahuila, Mexico. Revista Mexicana de Biodiversidad 78(1): 141-147.

Garcia-de la Pena, C.; Gadsden, H.; Lopez-Corrujedo, H.; Lazcano, D. 2005. Natural history notes: Uma exsul (Coahuila Fringe-toed Sand Lizard). Body temperature. Herpetological Review 36(1): 66-67.

Lemos-Espinal, J.A.; Auth, D.L.; Chiszar, D.; Smith, H.M. 2002. Observations on the Chihuahua fringe-toed lizard, Uma paraphygas. Bulletin of the Chicago Herpetological Society 37(1): 4-7.

Leyva-Pacheco, S.V.; Gadsden-Esparza, H.; Estrada-Rodriguez, J.L.; Romero Mendez, U. 2004. Natural history notes: Uma exsul (Coahuila Fringe-toed Lizard). Burrows. Herpetological Review 35(2): 174.

Pena, C.G. de la; Gaytan, G.C. 2006. Uma exsul. The Fringe-toed Sand Lizard. Reptilia (GB) 44: 74-78.

Pough, F.H.; Morafka, D.J.; Hillman, P.E. 1978. The ecology and burrowing behaviour of the Chihuahuan fringe-footed lizard, Uma exsul. Copeia 1978(1): 81-86.

Schmidt, K.P.; Bogert, C.M. 1947. A new fringe-footed sand lizard from Coahuila, Mexico. American Museum Novitates 1339: 1-9.

Schulte, J.J.; Smith, G.R.; Lemos-Espinal, J.A. 2017. Diet of juvenile lizards, Uma exsul, from Coahuila, Mexico. Southwestern Naturalist 62(1): 69–71.

Uma inornata

Anonymous. 1981. California lizard added to threatened list. Field Museum of Natural History Bulletin 52(1): 31.

Anonymous. 1984. Lizard bites California development prospects. Chicago Herpetological Society Newsletter 1984(February): 1 p. (Unpaginated. Reprinted from The Daily Sentinel (Grand Junction Co.), December 27, 1983).

Anonymous. 1984. Lizard with big teeth. Chicago Herpetological Society Newsletter 1984(March): 1 p. (Unpaginated. Reprinted from Buffalo News, February 19, 1984).

Anonymous. 1989. The sand's fine for diving in lizard's new reserve. Animal Kingdom 92: 17.

Barrows, C.; Muth, A.; Fisher, M.; Lovich, J. 1995. Coachella Valley fringe-toed lizards. pp. 137-138. In: LaRoe, E.T.; Farris, G.S.; Puckett, C.E.; Doran, P.D. & Mac, M.J. (eds.). Our living resources: a report to the nation on the distribution, abundance, and health of U.S. plants, animals and ecosystems. U.S. Department of the Interior, National Biological Service, Washington, D.C. 530 pp.

Barrows, C.W. 1996. An ecological model for the protection of a dune ecosystem. Conservation Biology 10(3): 888-891.

Barrows, C.W. 1997. Habitat relationships of the Coachella Valley fringe-toed lizard (Uma inornata). Southwestern Naturalist 42(2): 218-223.

Barrows, C.W. 2006. Population dynamics of a threatened Sand Dune Lizard. Southwestern Naturalist 51(4): 514-523.

Barrows, C.W.; Allen, M.F. 2007. Persistence and local extinctions of endangered lizard Uma inornata on isolated habitat patches. Endangered Species Research 3(1): 61-68.

Barrows, C.W.; Preston, K.L.; Rotenberry, J.T.; Allen, M.F. 2008. Using occurrence records to model historic distributions and estimate habitat losses for two psammophilic lizards. Biological Conservation 141(7): 1885-1893.

Barrows, C.W.; Rotenberry, J.T.; Allen, M.F. 2010. Assessing sensitivity to climate change and drought variability of a sand dune endemic lizard. Biological Conservation 143(3): 731-736.

Beatley, T. 1994. Reconciling urban growth and endangered species: the Coachella Valley habitat conservation plan. pp. 231-250. In: Platt, R.H.; Rowntree, R.A. & Muick, P.C. The ecological city: preserving and restoring urban biodiversity. University of Massachusetts Press, Amherst. 291 pp.

Bender, M. 1980. Coachella Valley fringe-toed lizard Threatened. Endangered Species Technical Bulletin 5(10): 1, 3.

Bolster, B.C.; Cornett, J.W.; Hansen, R.W.; Muth, A.; Rorabaugh, J. 1994. Coachella Valley fringe-toed lizard. pp. 268-270. In: Thelander, C.G. & Crabtree, M. (eds.): Life on the edge: a guide to California's endangered natural resources: wildlife. Biosystems Books, Santa Cruz, California. 550 pp.

Chen, X.W.; Barrows, C.W.; Li, B.A. 2006. Is the Coachella Valley fringe-toed lizard (Uma inornata) on the edge of extinction at Thousand Palms Preserve in California? Southwestern Naturalist 51(1): 28-34.

Chen, X.W.; Barrows, C.W.; Li, B.A. 2006. Phase coupling and spatial synchrony of subpopulations of an endangered dune lizard. Landscape Ecology 21(8): 1185-1193.

Cornett, J.W. 1982. Interbreeding between Uma inornata and Uma notata. Southwestern Naturalist 27(2): 223.

Durtsche, R.D. 1992. Feeding time strategies of the fringe-toed lizard, Uma inornata, during breeding and non-breeding seasons. Oecologia (Heidelberg) 89(1): 85-89.

Durtsche, R.D. 1995. Foraging ecology of the fringe-toed lizard, Uma inornata, during periods of high and low food abundance. Copeia 1995(4): 915-926.

England, A.S. 1983. The Coachella Valley, an endangered ecosystem progress report on conservation and management efforts. Cal-Neva Wildlife 1983: 148-156.

Finnley, D. (ed.) 1980. Critical Habitat reproposed for Coachella Valley lizard. Endangered Species Technical Bulletin 5(6): 1, 13.

Hedtke, S.M.; Zamudio, K.R.; Phillips, C.A.; Losos, J.; Brylski, P. 2007. Conservation genetics of the endangered Coachella Valley fringe-toed lizard (Uma inornata). Herpetologica 63(4): 411-420.

Holing, D. 1987. Lizard and the links. Audubon 89(6): 38-42,44,46,48-49.

Kinzel, J. 1968. Sand lizards. Herpetology 2(2): 7.

Král, B. 1969. A karyotype study of the Fringe-toed lizard Uma inornata Cope, 1895. Zool. Listy 18: 185-194.

Mayhew, W.W. 1965. Reproduction in the sand-dwelling lizard Uma inornata. Herpetologica 21: 39-55.

Mayhew, W.W. 1983. Conflicts between the fringe-toed lizard and development in the Coachella Valley. Cal-Neva Wildlife 1983: 144-147.

Neset, T.; Lisle, H.F. de 1977. Proposed endangered species. Herpetology 9(4): 7-8.

Pough, F.H. 1973. Uma inornata. Catalogue of American Amphibians and Reptiles 126: 1-2.

Trepanier, T.L.; Murphy, R.W. 2001. The Coachella Valley fringe-toed lizard (Uma inornata): Genetic diversity and phylogenetic relationships of an endangered species. Molecular Phylogenetics and Evolution 18(3): 327-334.

Turner, F.B.; Weaver, D.C.; Rorabaugh, J.C. 1984. Effects of reduction in windblown sand on the abundance of the fringe toed lizard (Uma inornata) in the Coachella Valley, California. Copeia 1984(2): 370-378.

Vandergast, A.G.; Wood, D.A.; Thompson, A.R.; Fisher, M.; Barrows, C.W.; Grant, T.J. 2016. Drifting to oblivion? Rapid genetic differentiation in an endangered lizard following habitat fragmentation and drought. Diversity and Distributions 22(3): 344-357.

Wells, K. 1984. How a little lizard gets Palm Springs to do its bidding. Developers were destroying its sandy desert habitat; it could shut them down. Chicago Herpetological Society Newsletter 1984(August): 2 pp. (Unpaginated. Reprinted from the Wall Street Journal, July 9, 1984).

Uma notata

Adler, K. 1986. Orientation mechanisms in a desert lizard, Uma notata. pp. 583-587. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European Herpetological Meeting (3rd Ordinary General Meeting of the Societas Europaea Herpetologica) Prague 1985. Charles University, Prague. 754 pp.

Adler, K.; Phillips, J.B. 1985. Orientation in a desert lizard (Uma notata): time-compensated compass movement and polarotaxis. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 156(4): 547-552.

Bart, C.E. 1931. On the occurrence of a throat-fan in the sand lizard, Uma notata Baird, with notes on the adaptive specializations of the form. Copeia 1931: 15-16.

Cornett, J.W. 1982. Interbreeding between Uma inornata and Uma notata. Southwestern Naturalist 27(2): 223.

Gottscho, A.D.; Wood, D.A.; Vandergast, A.G.; Lemos-Espinal, J.A.; Gatesy, J.; Reeder, T.W. 2017. Lineage diversification of fringe-toed lizards (Phrynosomatidae: Uma notata complex) in the Colorado Desert: delimiting species in the presence of gene flow. Molecular Phylogenetics and Evolution 106: 103-117.

Kleinbergs-Krisans, S.; Catlett, R.H. 1975. Radiosensitivity of the arenicolous lizard, Uma notata. Radiation Res. 61(1): 110-117.

Mayhew, W.W. 1966. Reproduction in the arenicolous lizard Uma notata. Ecology 47: 9-18.

Meyers, J.J.; Herrel, A. 2005. Prey capture kinematics of ant-eating lizards. Journal of Experimental Biology 208(1): 113-127.

Norris, K.S. 1951. The lizard that swims in the sand. Natural History (New York) 60: 404-407.

Norris, K.S. 1963. Environment and a lizard. Desert history is clue to adaptations. Natural History (New York) 72(1): 54-59.

Ortega, A.; Gonzalez-Romero, A.; Barbault, R. 1986. Rythmes journaliers d'activite et partage des ressources dans une communaute de lézards du desert de sonora, Mexique. Revue d'Ecologie la Terre et la Vie 41(4): 355-360.

Pough, F.H. 1970. The burrowing ecology of the sand lizard, Uma notata. Copeia 1970: 145-157.

Pough, F.H. 1977. Uma notata. Catalogue of American Amphibians and Reptiles 197: 1-2.

Punzo, F. 1982. Tail autotomy and running speed in the lizards Cophosaurus texanus and Uma notata. Journal of Herpetology 16(3): 329-331.

Shaw, C.E. 1952. Notes on the eggs and young of some United States and Mexican lizards. 1. Herpetologica 8(3): 71-79.

Sugerman, R.A.; Applegarth, J.S. 1980. An instance of natural cannibalism by Uma n. notata Baird. Herpetological Review 11(4): 90.

Uma paraphygas

Ballesteros-Barrera, C.; Martinez-Meyer, E.; Gadsden, H. 2007. Effects of land-cover transformation and climate change on the distribution of two microendemic lizards, genus Uma, of northern Mexico. Journal of Herpetology 41(4): 733-740.

Castañeda, G.; García-de la Peña, C.; Gadsden-Esparza, H.; Contreras-Balderas, A.J.; Cooper, W.E. 2011. Tail loss incidence in the Chihuahuan fringe toed-lizard Uma paraphygas (Squamata: Phrynosomatidae). Revista Mexicana de Biodiversidad 82(1): 193-198.

Castaneda-Gaytan, G.; Gadsden, H.; Lopez-Corrujedo, H.; Estrada-Rodriguez, J.L. 2003. Historia de vida de Uma paraphygas (Sauria: Phrynosomatidae) en la Reserva de la Biosfera de Mapimi, Durango. Acta Zoologica Mexicana Nueva Serie 89: 169-184.

Gadsden, E.H.; Palacios Orona, L.E. 1997. Seasonal dietary patterns of the Mexican fringe-toed lizard (Uma paraphygas). Journal of Herpetology 31(1): 1-9.

Gadsden-Esparza, H.; Méndez-de la Cruz, F.R.; Gil-Martínez, R.; Casas-Andreu, G. 1993. Patrón reproductivo de una lagartija (Uma paraphygas) en peligro de extinctión. Boletin de la Sociedad Herpetologica Mexicana 5(2): 52-50.

Garcia-de la Pena, C.; Gadsden, H.; Castaneda-Gaytan, G.; Lopez-Corrujedo, H. 2005. Body-ambient temperature relationships of the endemic Chihuahua fringe-toe lizard, Uma paraphygas. Bulletin of the Chicago Herpetological Society 40(9): 165-168.

Lemos-Espinal, J.A.; Auth, D.L.; Chiszar, D.; Smith, H.M. 2002. Observations on the Chihuahua fringe-toed lizard, Uma paraphygas. Bulletin of the Chicago Herpetological Society 37(1): 4-7.

Williams, K.L.; Chrapliwy, P.S.; Smith, H.M. 1960. A new fringe-footed lizard (Uma) from Mexico. Transactions of the Kansas Academy of Sciences 62 [1959]: 166-172.

Uma rufopunctata

Foster, C.D.; Tietgen, M.; Baumer, M. 2015. Yuman Fringe-toed Lizard (Uma rufopunctata) care and breeding at the Phoenix Zoo. Herpetological Review 46(1): 46-49.

Goldberg, S.R. 2012. Reproduction in the Yuman Fringe-toed Lizard, Uma rufopunctata (Squamata: Phrynosomatidae). Southwestern Naturalist 57(4): 456-457.

Goodman, C.M.; Schraft, H.A.; Clark, R.W. 2017. Natural history notes: Crotalus cerastes (Sidewinder). Diet / Scavenging. Herpetological Review 48(3): 670.

Tietgen, M.; Foster, C.D. 2014. Uma rufopunctata (Yuman Fringe-toed Lizard) behavior. Herpetological Review 45(4): 636-637.

Uma scoparia

Brown, T.W.; Lillywhite, H.B. 1992. Autecology of the Mojave Desert Sidewinder, Crotalus cerastes cerastes, at Kelso Dunes, Mojave Desert, California, USA. pp. 279-308. In: Campbell, J.A. & Brodie, E.D. (eds.). Biology of the pitvipers. Selva, Tyler, Texas. 467 pp.

Carothers, J.H. 1986. An experimental confirmation of morphological adaptation: toe fringes in the sand-dwelling lizard Uma scoparia. Evolution 40(4): 871-874.

Elvin, D.W. 1960. Range extension for Uma scoparia Cope. Herpetologica 16: 31.

Funk, R.S. 1974. Geographic distribution: Uma scoparia. USA: Arizona: Yuma Co. Herpetological Review 5(1): 20.

Gottscho, A.D.; Marks, S.B.; Jennings, W.B. 2014. Speciation, population structure, and demographic history of the Mojave Fringe-toed Lizard (Uma scoparia), a species of conservation concern. Ecology and Evolution 4(12): 2546-2562.

Gracie, A.E.; Murphy, R.W. 1986. Life history notes: Gambelia wislizenii (Leopard Lizard). Food. Herpetological Review 17(2): 47.

Irschick, D.J.; Jayne, B.C. 1998. Effects of incline on speed, acceleration, body posture and hindlimb kinematics in two species of lizard Callisaurus draconoides and Uma scoparia. Journal of Experimental Biology 201(2): 273-287.

Irschick, D.J.; Jayne, B.C. 1999. Comparative three-dimensional kinematics of the hindlimb for high-speed bipedal and quadrupedal locomotion of lizards. Journal of Experimental Biology 202(9): 1047-1065.

Jayne, B.C.; Daggy, M.W. 2000. The effects of temperature on the burial performance and axial motor pattern of the sand-swimming of the Mojave fringe-toed lizard Uma scoparia. Journal of Experimental Biology 203(7): 1241-1252.

Jayne, B.C.; Ellis, R.V. 1998. How inclines affect the escape behaviour of a dune-dwelling lizard, Uma scoparia. Animal Behaviour 55(5): 1115-1130.

Jayne, B.D.; Irschick, D.J. 2000. A field study of incline use and preferred speeds for the locomotion of lizards. Ecology (Washington, D.C.) 81(11): 2969-2983.

Jones, J.L.; Stocking, S.E. 2017. Geographic distribution: Uma scoparia (Mojave Fringe-toed Lizard). Herpetological Review 48(2): 390-391.

Korff, W.L.; McHenry, M.J. 2011. Environmental differences in substrate mechanics do not affect sprinting performance in sand lizards (Uma scoparia and Callisaurus draconoides). Journal of Experimental Biology 214(1): 122-130.

Leavitt, D.J.; Crumbo, W.L.; Myrand, J.M.; Masters, E.H. 2017. The geographic range of Uma scoparia Cope, 1894 (Squamata: Phrynosomatidae) in Arizona. Check List 13(2)(Article 2090): 1-4.

Mayhew, W.W. 1966. Reproduction in the psammophilous lizard Uma scoparia. Copeia 1966: 14-122.

Murphy, R.W.; Trepanier, T.L.; Morafka, D.J. 2006. Conservation genetics, evolution and distinct population segments of the Mojave fringe-toed lizard, Uma scoparia. Journal of Arid Environments 67 (Supplement): 226-247.

Pough, F.H. 1974. Uma scoparia. Catalogue of American Amphibians and Reptiles 155: 1-2.

Schlüter, U. 2001. Das Leguanportrait: Uma scoparia Cope, 1894. Iguana Rundschreiben 14(1): 4-6.