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Taxonomy of the genus Carlia Biodiversity of the family Scincidae

Bibliography of the genus
Carlia (Australian-Pacific Rainbow Skinks)

(Reptilia: Sauria: Scincidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Carlia in general

Bedford, G. 1995. Natural history notes: Carlia (Scincidae). Tail waving and prey piracy. Herpetological Review 26(2): 99-100.

Couper, P.J.; Worthington-Wilmer, J.; Roberts, L.; Amey, A.P.; Zug, G.R. 2005. Skinks currently assigned to Carlia aerata (Scincidae: Lygosominae) of north-eastern Queensland: a preliminary study of cryptic diversity and two new species. Australian Journal of Zoology 53(1): 35-49.

Coventry, A.J. 1971. The discovery of the lizard genus Carlia (Scincidae: Lygosominae) in Victoria. Victorian Naturalist 88(1): 20-22.

Dolman, G.; Hugall, A.F. 2008. Combined mitochondrial and nuclear data enhance resolution of a rapid radiation of Australian rainbow skinks (Scincidae: Carlia). Molecular Phylogenetics and Evolution 49: 782-794.

Dolman, G.; Moritz, C. 2006. A multilocus perspective on refugial isolation and divergence in rainforest skinks (Carlia). Evolution 60(3): 573-582.

Donnellan, S.C.; Couper, P.J.; Saint, K.M.; Wheaton, L. 2009. Systematics of the Carlia 'fusca' complex (Reptilia: Scincidae) from northern Australia. Zootaxa 2227: 1-31.

Fritts, T.H.; Rodda, G.H. 1998. The role of introduced species in the degradation of island ecosystems: a case history of Guam. Annual Review of Ecology and Systematics 29: 113-140.

Greer, A.E. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology (Supplementary Series) 31: 1-67.

Ingram, G.J.; Covacevich, J. 1989. Revision of the genus Carlia (Reptilia, Scincidae) in Australia with comments on Carlia bicarinata of New Guinea. Memoirs of the Queensland Museum 27(2): 443-490.

Kraus, F. 2007. Taxonomic partitioning within Papuan members of the Carlia novaeguineae complex (Squamata: Scincidae). Journal of Herpetology 41(3): 410-423.

McCoid, M.J. 1995. Natural history notes: Carlia cf. fusca (NCN). Behavior. Herpetological Review 26(1): 35.

Mitchell, P.J. 1953. A brief revision of the four-fingered members of the genus Leiolopisma (Lacertilia). Records of the Australian Museum 11(1): 75-90.

Shah, B. 2001. Mite infestations in Australian skinks: seasonal, geographical and ecological variation. Herpetofauna (Sydney) 31(2): 102-107.

Storr, G.M. 1974. The genus Carlia (Lacertilia, Scincidae) in Western Australia and Northern Territory. Records of the Western Australian Museum 3(2): 151-165.

Stuart-Fox, D.M.; Hugall, A.F.; Moritz, C. 2002. A molecular phylogeny of rainbow skinks (Scincidae: Carlia): taxonomic and biogeographic implications. Australian Journal of Zoology 50(1): 39-51.

Wells, R. 1975. Notes on an unidentified skink of the genus Carlia from Black Mountain, N.E. Queensland. Herpetofauna (Sydney) 7(2): 11.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Zug, G.R.; Allison, A. 2006. New Carlia fusca complex lizards (Reptilia: Squamata: Scincidae) from New Guinea, Papua-Indonesia. Zootaxa 1237: 27-44.

Carlia aenigma

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia ailanpalai

Austin, C.C.; Rittmeyer, E.N.; Oliver, L.A.; Andermann, J.O.; Zug, G.R.; Rodda, G.H.; Jackson, N.D. 2011. The bioinvasion of Guam: inferring geographic origin, pace, pattern and process of an invasive lizard (Carlia) in the Pacific using multi-locus genomic data. Biological Invasions 13(9): 1951-1967.

Boyarski, V.L.; Savidge, J.A.; Rodda, G.H. 2008. Brown treesnake (Boiga irregularis) trappability: attributes of the snake, environment and trap. Applied Herpetology 5(1): 47-61.

Buden, D.W. 2009. Carlia ailanpalai (Reptilia: Scincidae): an invasive species of lizard in the federated states of Micronesia. Pacific Science 63(2): 243-251.

Campbell, E.W.; Yackel-Adams, A.A.; Converse, S.J.; Fritts, T.H.; Rodda, G.H. 2012. Do predators control prey species abundance? An experimental test with Brown Treesnakes on Guam. Ecology (Washington D C) 93(2): 1194-1203.

Goldberg, S.R. 2011. Natural history notes: Carlia ailanpalai (NCN). Hermaphroditism. Herpetological Review 42(1): 92-93.

Goldberg, S.R.; Kraus, F. 2012. Reproduction in the invasive lizard, Carlia ailanpalai (Squamata: Scincidae) from Oceania. Russian Journal of Herpetology 19(3): 199-202.

McCoid, M.J.; Henke, S.E.; Hensley, R.A. 2005. Husbandry and captive reproduction in Carlia ailanpalai (Scincidae). Herpetological Review 36(3): 292-293.

Tadevosyan, T.L.; Lardner, B. 2016. Natural history notes: Carlia ailanpalai (Curious Skink). Diet. Herpetological Review 47(1): 134-135.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia amax

Potter, S.; Bragg, J.G.; Peter, B.M.; Bi, K.; Moritz, C. 2016. Phylogenomics at the tips: inferring lineages and their demographic history in a tropical lizard, Carlia amax. Molecular Ecology 25(6): 1367-1380.

Turner, G. 1995. Notes on Carlia amax (Scincidae). Herpetofauna (Sydney) 25(2): 54-56.

Carlia aramia

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia babarensis

Kopstein, F. 1926. Reptilien von den Molukken und den Benachbarten Inseln. Zoologische Mededelingen (Leiden) 9: 71-112.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia beccarii

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia bicarinata

O'Shea, M. 1993. Life history notes: Carlia bicarinata (Bicarinate Four-fingered Skink). Partial albinism. Herpetological Review 24(2): 59.

Zug, G.R.; Barber, M.M.; Dudley, J.C. 1982. Gonadal histology and reproduction in Carlia bicarinata (Scincidae, Sauria, Reptilia) of the Port Moresby area, Papua New Guinea. Herpetologica 38(3): 418-425.

Carlia bomberai

Zug, G.R.; Allison, A. 2006. New Carlia fusca complex lizards (Reptilia: Squamata: Scincidae) from New Guinea, Papua-Indonesia. Zootaxa 1237: 27-44.

Carlia caesius

Goldberg, S.R. 2018. Natural history notes: Carlia caesius. Reproduction. Herpetological Review 49(2): 326.

Zug, G.R.; Allison, A. 2006. New Carlia fusca complex lizards (Reptilia: Squamata: Scincidae) from New Guinea, Papua-Indonesia. Zootaxa 1237: 27-44.

Carlia crypta

Singhal, S.; Hoskin, C.J.; Couper, P.; Potter, S.; Moritz, C. 2018. A framework for resolving cryptic species: a case study from the lizards of the Australian Wet Tropics. Systematic Biology 67(6):1061-1075.

Carlia decora

Hoskin, C.J.; Couper, P.J. 2012. Description of two new Carlia species (Reptilia: Scincidae) from north-east Australia, elevation of Carlia pectoralis inconnexa Ingram & Covacevich 1989 to full species status, and redescription of Carlia pectoralis (de Vis 1884). Zootaxa 3546: 1-28.

Carlia diguliensis

Kopstein, F. 1926. Reptilien von den Molukken und den Benachbarten Inseln. Zoologische Mededelingen (Leiden) 9: 71-112.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia dogare

Covacevich, J.A.; Ingram, G.J. 1978. Three new species of rainbow skinks of the genus Carlia from northern Queensland. Victorian Naturalist 92(1-2): 19-22.

Vickers, M.; Schwarzkopf, L. 2016. A simple method to predict body temperature of small reptiles from environmental temperature. Ecology and Evolution 6(10): 3059-3066.

Carlia eothen

Goldberg, S.R.; Kraus, F. 2008. Notes on reproduction of Carlia eothern and Lamprolepis smaragdina (Squamata: Scincidae) from Papua New Guinea. Herpetofauna (Sydney) 38(2): 110-115.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia fusca

Donnellan, S.C.; Couper, P.J.; Saint, K.M.; Wheaton, L. 2009. Systematics of the Carlia 'fusca' complex (Reptilia: Scincidae) from northern Australia. Zootaxa 2227: 1-31.

O'Shea, M. 1993. Life history notes: Carlia fusca (Brown Four-fingered Skink). Display behavior. Herpetological Review 24(3): 105.

Ota, H.; Hikida, T.; Matsui, M.; Hasegawa, M. 1988. Karyotype of a scincid lizard, Carlia fusca, from Guam, the Mariana Islands. Zoological Science (Tokyo) 5(4): 901-903.

Wilhoft, D.C. 1961. Temperature responses in two tropical Australian skinks. Herpetologica 17: 109-113.

Wilhoft, D.C.; Reiter, E.O. 1965. Sexual cycle of the lizard, Leiolopisma fuscum, a tropical Australian skink. Journal of Morphology 116: 379-387.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia gracilis

Doody, J.S.; Taylor, R.V. 1997. A note on the foraging behaviour of the snake Dendrelaphis punctulata in the Northern Territory. Herpetofauna (Sydney) 27(2): 55-56.

Nickay, J.L. 2007. Natural history notes: Carlia gracilis (Slender Rainbow Skink) and Glaphyromorphus darwiniensis (Darwin Skink). Feeding aggregation. Herpetological Review 38(4): 450-451.

Carlia inconnexa

Hoskin, C.J.; Couper, P.J. 2012. Description of two new Carlia species (Reptilia: Scincidae) from north-east Australia, elevation of Carlia pectoralis inconnexa Ingram & Covacevich 1989 to full species status, and redescription of Carlia pectoralis (de Vis 1884). Zootaxa 3546: 1-28.

Carlia insularis

Afonso Silva, A.C.; Santos, N.; Ogilvie, H.A.; Moritz, C. 2017. Validation and description of two new north-western Australian Rainbow skinks with multispecies coalescent methods and morphology. PeerJ 5: e3724.

Carlia isostriacantha

Afonso Silva, A.C.; Santos, N.; Ogilvie, H.A.; Moritz, C. 2017. Validation and description of two new north-western Australian Rainbow skinks with multispecies coalescent methods and morphology. PeerJ 5: e3724.

Carlia jarnoldae

Covacevich, J.A.; Ingram, G.J. 1978. Three new species of rainbow skinks of the genus Carlia from northern Queensland. Victorian Naturalist 92(1-2): 19-22.

Langkilde, T.; Alford, R.A.; Schwarzkopf, L. 2005. No behavioural compensation for fitness costs of autotomy in a lizard. Austral Ecology 30(7): 713-718.

Langkilde, T.; Schwarzkopf, L. 2003. Observations of mating behavior and reproduction in the scincid Carlia jarnoldae. Herpetological Review 34(4): 325-327.

Langkilde, T.; Schwarzkopf, L.; Alford, R.A. 2003. An ethogram for adult male rainbow skinks, Carlia jarnoldae. Herpetological Journal 13(3): 141-148.

Langkilde, T.; Schwarzkopf, L.; Alford, R.A. 2005. The function of tail displays in male rainbow skinks (Carlia jarnoldae). Journal of Herpetology 39(2): 325-328.

Vickers, M.; Schwarzkopf, L. 2016. A random walk in the park: an individual-based null model for behavioral thermoregulation. American Naturalist 187(4): 481-490.

Vickers, M.; Schwarzkopf, L. 2016. A simple method to predict body temperature of small reptiles from environmental temperature. Ecology and Evolution 6(10): 3059-3066.

Carlia johnstonei

Afonso Silva, A.C.; Bragg, J.G.; Potter, S.; Fernandes, C.; Coelho, M.M.; Moritz, C. 2017. Tropical specialist vs. climate generalist: Diversification and demographic history of sister species of Carlia skinks from northwestern Australia. Molecular Ecology 26(15): 4045-4058

Carlia leucotaenia

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia longipes

Couper, P.J.; Amey, A.P. 2010. Tips for separating Carlia rostralis from Carlia longipes - resolving the inadequacies of existing keys. Herpetofauna (Sydney) 40(1): 13-15.

Donnellan, S.C.; Couper, P.J.; Saint, K.M.; Wheaton, L. 2009. Systematics of the Carlia 'fusca' complex (Reptilia: Scincidae) from northern Australia. Zootaxa 2227: 1-31.

Goodman, B.A.; Schwarzkopf, L.; Krockenberger, A.K. 2013. Phenotypic integration in response to incubation environment adaptively influences habitat choice in a tropical lizard. American Naturalist 182(5): 666-673.

O'Brien, M. 1994. Predation on lizards by Carlia longipes. Herpetofauna (Sydney) 24(2): 43.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia luctuosa

Tanner, V.M. 1952. Pacific islands herpetology. 6. Tahiti and Marquesas Islands, New Guinea and Australia. Great Basin Naturalist 12: 1-12.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia munda

Bedford, G. 1995. Natural history notes: Carlia (Scincidae). Tail waving and prey piracy. Herpetological Review 26(2): 99-100.

Greer, A.E. 1975. Notes on the systematics of the genus Carlia (Lacertilia: Scincidae). 1. Carlia melanopogon Gray 1845. Herpetologica 31(1): 70-75.

Metcalfe, D.C.; Peterson, M. 2005. Natural history notes: Carlia munda (NCN). Predation; fire response. Herpetological Review 36(4): 445.

Valentine, L.E.; Roberts, B.; Schwarzkopf, L. 2007. Mechanisms driving avoidance of non-native plants by lizards. Journal of Applied Ecology 44(1): 228-237.

Vickers, M.; Schwarzkopf, L. 2016. A simple method to predict body temperature of small reptiles from environmental temperature. Ecology and Evolution 6(10): 3059-3066.

Carlia mysi

O'Shea, M.; Paiva, O. 2014. Natural history notes: Carlia curta (Papuan Litter Skink) and Carlia mysi (Mys' Rainbow Skink). Mass avian predation. Herpetological Review 45(3): 493-494.

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia nigrauris

Zug, G.R. 2010. An outlying Carlia population from Java and comments on species groups within the genus Carlia (Reptilia: Squamata: Scincidae). Proceedings of the California Academy of Sciences 61(7-18): 389-408.

Carlia pectoralis

Blomberg, S.P.; Owens, I.P.F.; Stuart-Fox, D.M. 2001. Ultraviolet reflectance in the small skink Carlia pectoralis. Herpetological Review 32(1): 16-17.

Hoskin, C.J.; Couper, P.J. 2012. Description of two new Carlia species (Reptilia: Scincidae) from north-east Australia, elevation of Carlia pectoralis inconnexa Ingram & Covacevich 1989 to full species status, and redescription of Carlia pectoralis (de Vis 1884). Zootaxa 3546: 1-28.

Jamieson, B.G.M.; Scheltinga, D.M. 1994. The ultrastructure of spermatozoa of the Australian skinks, Ctenotus taeniolatus, Carlia pectoralis and Tiliqua scincoides scincoides (Scincidae, Reptilia). Memoirs of the Queensland Museum 37(1): 181-193.

Valentine, L.E.; Roberts, B.; Schwarzkopf, L. 2007. Mechanisms driving avoidance of non-native plants by lizards. Journal of Applied Ecology 44(1): 228-237.

Carlia peronii

Greer, A.E. 1976. Notes on the systematics of the genus Carlia (Lacertilia: Scincidae). 2. Carlia peroni (Duméril and Bibron 1839). Herpetologica 32(4): 371-377.

Carlia pulla

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia quinquecarinatus

Donnellan, S.C.; Couper, P.J.; Saint, K.M.; Wheaton, L. 2009. Systematics of the Carlia 'fusca' complex (Reptilia: Scincidae) from northern Australia. Zootaxa 2227: 1-31.

Carlia rhomboidalis

Dolman, G. 2008. Evidence for differential assortative female preference in association with refugial isolation of rainbow skinks in Australia's tropical rainforests. PLoS ONE 3(10): e3499, 1-9.

Smith, T.B.; Schneider, C.J.; Holder, K. 2001. Refugial isolation versus ecological gradients: Testing alternative mechanisms of evolutionary divergence in four rainforest vertebrates. Genetica (Dordrecht) 112-113: 383-398.

Stuart-Fox, D.M.; Schneider, C.J.; Moritz, C.; Couper, P.J. 2001. Comparative phylogeography of three rainforest-restricted lizards from mid-east Queensland. Australian Journal of Zoology 49(2): 119-127.

Wilhoft, D.C. 1961. Temperature responses in two tropical Australian skinks. Herpetologica 17: 109-113.

Wilhoft, D.C. 1963. Reproduction in the tropical Australian skink, Leiolopisma rhomboidalis. American Midland Naturalist 70: 442-461.

Carlia rostralis

Couper, P.J.; Amey, A.P. 2010. Tips for separating Carlia rostralis from Carlia longipes - resolving the inadequacies of existing keys. Herpetofauna (Sydney) 40(1): 13-15.

Lloyd, R.; Alford, R.A.; Schwarzkopf, L. 2009. Chemical discrimination among predators by lizards: responses of three skink species to the odours of highand low-threat varanid predators. Austral Ecology 34(1): 50-54.

Manicom, C.; Alford, R.; Schoener, T.W.; Schwarzkopf, L. 2014. Mechanisms causing variation in sexual size dimorphism in three sympatric, congeneric lizards. Ecology (Washington D C) 95(6): 1531-1544.

Manicom, C.; Schwarzkopf, L. 2011. Diet and prey selection of sympatric tropical skinks. Austral Ecology 36(5): 485-496.

Vickers, M.; Manicom, C.; Schwarzkopf, L. 2011. Extending the cost-benefit model of thermoregulation: high-temperature environments. American Naturalist 177(4): 452-461.

Whittier, J. 1993. Behavioural repertoire of Carlia rostralis (Scincidae) in the wet tropics of Queensland, Australia. pp. 305-310. In: Lunney, D. & Ayers, D. (eds.). Herpetology in Australia: a diverse discipline. Surrey Beatty & Sons, Chipping Norton, New South Wales. 414 pp.

Whittier, J. 1993. Ecological notes on Carlia rostralis in rainforest and associated habitat in the southern wet tropics. Memoirs of the Queensland Museum 34(1): 125-129.

Whittier, J.; Martin, J. 1992. Aspects of social behaviour and dominance in male rainbow skinks, Carlia rostralis. Australian Journal of Zoology 40(1): 73-79.

Carlia rubigo

Hoskin, C.J.; Couper, P.J. 2012. Description of two new Carlia species (Reptilia: Scincidae) from north-east Australia, elevation of Carlia pectoralis inconnexa Ingram & Covacevich 1989 to full species status, and redescription of Carlia pectoralis (de Vis 1884). Zootaxa 3546: 1-28.

Carlia rubrigularis

Andersson, M.; Krockenberger, A.; Schwarzkopf, L. 2010. Experimental manipulation reveals the importance of refuge habitat temperature selected by lizards. Austral Ecology 35(3): 294-299.

Dolman, G. 2008. Evidence for differential assortative female preference in association with refugial isolation of rainbow skinks in Australia's tropical rainforests. PLoS ONE 3(10): e3499, 1-9.

Goodman, B.A. 2006. Costs of reproduction in a tropical invariant-clutch producing lizard (Carlia rubrigularis). Journal of Zoology (London) 270(2): 236-243.

Goodman, B.A. 2006. The effects of maternal size on clutch traits in a tropical invariant-clutch lizard, Carlia rubrigularis (Scincidae). Amphibia-Reptilia 27(4): 505-511.

Lane, A.M. 2006. Observations of courtship, copulation, and gestation in the wet tropical endemic skink Carlia rubrigularis. Herpetological Review 37(1): 46-47.

Lloyd, R.; Alford, R.A.; Schwarzkopf, L. 2009. Chemical discrimination among predators by lizards: responses of three skink species to the odours of highand low-threat varanid predators. Austral Ecology 34(1): 50-54.

Manicom, C.; Alford, R.; Schoener, T.W.; Schwarzkopf, L. 2014. Mechanisms causing variation in sexual size dimorphism in three sympatric, congeneric lizards. Ecology (Washington D C) 95(6): 1531-1544.

Manicom, C.; Schwarzkopf, L. 2011. Diet and prey selection of sympatric tropical skinks. Austral Ecology 36(5): 485-496.

Moritz, C.; Langham, G.; Kearney, M.; Krockenberger, A.; VanDerWal, J.; Williams, S. 2012. Integrating phylogeography and physiology reveals divergence of thermal traits between central and peripheral lineages of tropical rainforest lizards. Royal Society Philosophical Transactions Biological Sciences 367(1596): 1680-1687.

Phillips, B.L.; Baird, S.J.E.; Moritz, C. 2004. When vicars meet: a narrow contact zone between morphologically cryptic phylogeographic lineages of the rainforest skink, Carlia rubrigularis. Evolution 58(7): 1536-1548.

Pintor, A.F.V.; Schwarzkopf, L.; Krockenberger, A.K. 2016. Hydroregulation in a tropical dry-skinned ectotherm. Oecologia (Berlin) 182(4): 925-931.

Schneider, C.; Smith, T.B.; Larison, B.; Moritz, C. 1999. A test of alternative models of diversification in tropical rainforests: ecological gradients vs. rainforest refugia. Proceedings of the National Academy of Sciences of the United States of America 96(24): 13869-13873.

Shoo, L.P.; Wilson, R.; Williams, Y.M.; Catterall, C.P. 2014. Putting it back: woody debris in young restoration plantings to stimulate return of reptiles. Ecological Management & Restoration 15(1): 84-87.

Singhal, S.; Bi, K. 2017. History cleans up messes: The impact of time in driving divergence and introgression in a tropical suture zone. Evolution 71(7): 1888-1899

Singhal, S.; Moritz, C. 2013. Reproductive isolation between phylogeographic lineages scales with divergence. Proceedings of the Royal Society Biological Sciences Series B 280(1772): 20132246, 1-8.

Smith, T.B.; Schneider, C.J.; Holder, K. 2001. Refugial isolation versus ecological gradients: Testing alternative mechanisms of evolutionary divergence in four rainforest vertebrates. Genetica (Dordrecht) 112-113: 383-398.

Torr, G.A. 1994. Combat behaviour in the rainforest skink Carlia rubrigularis. Herpetofauna (Sydney) 24(2): 40.

Turner, G. 2010. Some predator-prey interactions involving reptiles and frogs from the wet tropics region, north Queensland. Queensland Naturalist 48(4-6): 78-84.

Turner, G.S. 2018. Natural history notes: Cryptophis nigrescens (Eastern Small-eyed Snake). Diet. Herpetological Review 49(2): 339.

Vickers, M.; Manicom, C.; Schwarzkopf, L. 2011. Extending the cost-benefit model of thermoregulation: high-temperature environments. American Naturalist 177(4): 452-461.

Carlia schmeltzii

Covacevich, J.A.; Ingram, G.J. 1978. Three new species of rainbow skinks of the genus Carlia from northern Queensland. Victorian Naturalist 92(1-2): 19-22.

Doody, J.S.; Schembri, B. 2014. Natural history notes: Carlia schmeltzii (Robust Rainbow Skink). Environmentally-cued hatching. Herpetological Review 45(3): 494.

Hacking, J.; Rickard, A.; Schwarzkopf, L. 2014. Why do lizards avoid weeds? Biological Invasions 16(4): 935-947.

Carlia sexdentatus

Donnellan, S.C.; Couper, P.J.; Saint, K.M.; Wheaton, L. 2009. Systematics of the Carlia 'fusca' complex (Reptilia: Scincidae) from northern Australia. Zootaxa 2227: 1-31.

Vickers, M.; Schwarzkopf, L. 2016. A random walk in the park: an individual-based null model for behavioral thermoregulation. American Naturalist 187(4): 481-490.

Carlia storri

Lloyd, R.; Alford, R.A.; Schwarzkopf, L. 2009. Chemical discrimination among predators by lizards: responses of three skink species to the odours of highand low-threat varanid predators. Austral Ecology 34(1): 50-54.

Manicom, C.; Alford, R.; Schoener, T.W.; Schwarzkopf, L. 2014. Mechanisms causing variation in sexual size dimorphism in three sympatric, congeneric lizards. Ecology (Washington D C) 95(6): 1531-1544.

Manicom, C.; Schwarzkopf, L. 2011. Diet and prey selection of sympatric tropical skinks. Austral Ecology 36(5): 485-496.

Vickers, M.; Manicom, C.; Schwarzkopf, L. 2011. Extending the cost-benefit model of thermoregulation: high-temperature environments. American Naturalist 177(4): 452-461.

Carlia sukur

Zug, G.R.; Kaiser, H. 2014. A new species of four-toed skink (Squamata: Scincidae: Carlia peronii species group) from Pulau Sukur, Indonesia, and biogeographic notes on the herpetofauna of Flores and Komodo. Proceedings of the Biological Society of Washington 126(4): 379-392.

Carlia tetradactyla

Fischer, J.; Lindenmayer, D.; Cowling, A. 2003. Habitat models for the four-fingered skink (Carlia tetradactyla) at the microhabitat and landscape scale. Wildlife Research 30(5): 495-504.

Fischer, J.; Lindenmayer, D.B.; Cowling, A. 2004. The challenge of managing species at multiple scales: Reptiles in an Australian grazing landscape. Journal of Applied Ecology 41(1): 32-44.

Letnic, M.I.; Fox, B.J. 1997. The impact of industrial fluoride fallout on faunal succession following sand mining of dry sclerophyll forest at Tomago, NSW. - 1. Lizard recolonisation. Biological Conservation 80(1): 63-81.

Michael, D.R.; Cunningham, R.B.; Lindenmayer, D.B. 2010. Microhabitat relationships among five lizard species associated with granite outcrops in fragmented agricultural landscapes of south-eastern Australia. Austral Ecology 35(2): 215-226.

Porter, R. 1993. A record of communal egg-laying in the skink Carlia tetradactyla. Memoirs of the Queensland Museum 33(1): 60.

Carlia triacantha

Afonso Silva, A.C.; Bragg, J.G.; Potter, S.; Fernandes, C.; Coelho, M.M.; Moritz, C. 2017. Tropical specialist vs. climate generalist: Diversification and demographic history of sister species of Carlia skinks from northwestern Australia. Molecular Ecology 26(15): 4045-4058

Carlia tutela

Zug, G.R. 2004. Systematics of the Carlia "fusca" lizards (Squamata: Scincidae) of New Guinea and nearby islands. Bishop Museum Bulletins in Zoology 5: 1-83.

Carlia vivax

Goodman, B.A. 2006. Clutch traits in the skink Carlia vivax. Herpetofauna (Sydney) 36(1): 27-30.

Singh, S.; Smyth, A.K.; Blomberg, S.P. 2002. Effect of a control burn on lizards and their structural environment in a eucalypt open-forest. Wildlife Research 29(5): 447-454.

Singh, S.; Smyth, A.K.; Blomberg, S.P. 2002. Thermal ecology and structural habitat use of two sympatric lizards (Carlia vivax and Lygisaurus foliorum) in subtropical Australia. Austral Ecology 27(6): 616-623.

Carlia wundalthini

Hoskin, C.J. 2014. A new skink (Scincidae: Carlia) from the rainforest uplands of Cape Melville, north-east Australia. Zootaxa 3869(3): 224–236.