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© Rune Midtgaard

Taxonomy of the family Agamidae
Bibliography of the genus Harpesaurus
Biodiversity of the family Agamidae


Sunda Horned Agamas, Nose-horned Agamas, Hook-nosed Agamas



1851 Arpephorus Dumeril [not Arpephorus Stephens 1829 (Insecta: Coleoptera)] (type species: Arpephorus tricinctus Dumeril 1851; syn. Wermuth 1967)
1885 Harpesaurus Boulenger [substitute name for Arpephorus Dumeril 1851]
1924 Hylagama Mertens (type species: Hylagama borneensis Mertens 1924; syn. Das & Yaakob 2007; Manthey 2010)
Contents: 5 species, all of which are endemic.
Remarks: Previously included Thaumatorhynchus. One species (borneensis) was formerly placed in a separate genus, Hylagama.
Distribution: Indonesia (Java, Kalimantan, Nias, Sumatra), Malaysia (Sarawak).

Harpesaurus beccarii

West Sumatran Horned Agama*, (Beccari's Horned Mountain Lizard)

Beccaris Hornagame

Vestsumatransk Hornagam*

1888 Harpesaurus beccarii Doria

Distribution: Indonesia (Sumatra).

Harpesaurus borneensis

Bornean Horned Agama, Wood Agama



1924 Hylagama borneensis Mertens
Harpesaurus borneensis Manthey & Grossmann 1997
1978 Harpesaurus thescelorhinos King (Moody 1980; Manthey & Grossmann 1997)

Distribution: Indonesia (Kalimantan), Malaysia (Sarawak).

Harpesaurus ensicauda

Nias Horned Agama, (Werner's Horned Agama)



1913 Harpesaurus ensicauda Werner

Distribution: Indonesia (Nias).

Harpesaurus modigliani

East Sumatran Horned Agama*, (Modigliani's Horned Agama)

Modiglianis Hornagame

Østsumatransk Hornagam*

1933 Harpesaurus modigliani Vinciguerra

Distribution: Indonesia (Sumatra).

Harpesaurus tricinctus

Three-banded Horned Agama


Trebåndet Hornagam*

1851 Arpephorus tricinctus Dumeril
Harpesaurus tricinctus Boulenger 1885

Distribution: Indonesia (Java).