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Taxonomy of the family Colubridae
Bibliography of the genus Helicops
Biodiversity of the family Colubridae








Genus
Helicops

South American Keelbacks, South American Water Snakes, Neotropical Water Snakes

Scheelaugennattern

Sydamerikanske Vandsnoge

1828 Helicops Wagler (type species: Coluber carinicaudus Wied 1824)
1843 Tachynectes Fitzinger (type species: Homalopsis leopardina Schlegel 1837)
1843 Uranops Fitzinger (type species: Coluber angulatus Linnaeus 1758)
1849 Uranops Gray [not Uranops Fitzinger 1843 (Serpentes)] (type species: Coluber angulatus Linnaeus 1758)
Contents: 19 species, of which 11 (57.9%) are endemic.
Distribution: South America.
Reported from: Argentina, Bolivia, Brazil (Acre, Alagoas, Amapa, Amazonas, Bahia, Ceara, Distrito Federal, Espirito Santo, Goias, Maranhao, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Para [incl. Marajo Island], Paraiba, Parana, Pernambuco, Piaui, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Rondonia, Roraima, Santa Catarina, Sao Paulo, Sergipe, Tocantins), Colombia, Ecuador, French Guiana, Guyana, Paraguay, Peru, Surinam, Trinidad & Tobago (Trinidad), Uruguay, Venezuela.

Helicops angulatus

Brown-banded Keelback, Banded South American Water Snake

Braunbändige Scheelaugennatter

Brunbĺndet Vandsnog

1758 Coluber angulatus Linnaeus
Helicops angulatus Wagler 1830
1802 Coluber Surinamensis Shaw (Peters & Orejas-Miranda 1970)
1824 Natrix aspera Wagler (Peters & Orejas-Miranda 1970)
1869 Helicops cyclops Cope (Peters & Orejas-Miranda 1970)
1869 Helicops fumigatus Cope (Peters & Orejas-Miranda 1970)

Distribution: Bolivia, Brazil (Acre, Alagoas, Amapa, Amazonas, Bahia, Ceara, Distrito Federal, Goias, Maranhao, Mato Grosso, Mato Grosso do Sul, Para [incl. Marajo Island], Paraiba, Pernambuco, Piaui, Rio Grande do Norte, Rondonia, Roraima, Sergipe, Tocantins), Colombia, Ecuador, French Guiana, Guyana, Peru, Surinam, Trinidad & Tobago (Trinidad), Venezuela.


Helicops apiaka

Apiaká Keelback

Apiaká-Scheelaugennatter

Apiaká-vandsnog

2013 Helicops apiaka Kawashita-Ribeiro, Ávila & Morais

Distribution: Brazil (Mato Grosso).


Helicops boitata

Poconé Keelback

Poconé-Scheelaugennatter

Poconé-vandsnog

2019 Helicops boitata Moraes-da-Silva, Amaro, Sales Nunes, Strüssmann, Teixeira, Andrade, Sudré, Recoder, Rodrigues & Curcio

Distribution: Brazil (Mato Grosso).





Helicops carinicaudus

Keel-tailed Keelback

Kielschwanznatter

Keel-tailed Keelback

1824 Coluber carinicaudus Wied
Helicops carinicaudus Wagler 1830

Remarks: Previously included infrataeniatus. Records from Colombia (Pérez-Santos & Moreno 1988) refer to danieli (Yuki 1994). Correct publication year of the original description is 1824, not 1825 (Vanzolini & Myers 2015; Costa & Bérnils 2015). Records from Mato Grosso do Sul are erroneous (Costa & Bérnils 2018).
Distribution: Brazil (Bahia, Espirito Santo, Parana, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, Sao Paulo).


Helicops danieli

Colombian Keelback

Kolumbianische Scheelaugennatter

Colombiansk Vandsnog

1938 Helicops danieli Amaral

Distribution: Colombia.


Helicops gomesi

Săo Paulo Keelback

Săo-Paulo-Scheelaugennatter

Săo Paulo-vandsnog

1922 Helicops gomesi Amaral

Remarks: Correct publication year of the original description is 1922, not 1921 (Vanzolini 1977; Costa & Bérnils 2015).
Distribution: Brazil (Goias, Sao Paulo).


Helicops hagmanni

Amazonian Keelback

Amazonas-Scheelaugennatter

Amazonas-vandsnog

1910 Helicops hagmanni Roux

Distribution: Brazil (Acre, Amapa, Amazonas, Para [incl. Marajo Island], Rondonia), Colombia, Venezuela.


Helicops hogei

Venezuelan Keelback

Venezuelanische Scheelaugennatter

Venezuelansk Vandsnog

1964 Helicops hogei Lancini

Remarks: Wallach, Williams & Boundy (2014) listed hogei as a synonym of scalaris, but without discussion.
Distribution: Venezuela.


Helicops infrataeniatus

Surinam Keelback, Pampean Water Snake

Surinam-Scheelaugennatter

Surinam-vandsnog

1865 Helicops infrataeniatus Jan
Helicops carinicauda infrataeniata Griffin 1916
1865 Helicops carinicaudus gastrosticta Jan (Peters & Orejas-Miranda 1970)
1878 Helicops trivittatus Cope [not Myron trivittatus Gray 1849] (Peters & Orejas-Miranda 1970)
1885 Helicops baliogaster Cope (Peters & Orejas-Miranda 1970)
1897 Helicops pictiventris Werner (Cei 1993)

Remarks: Previously regarded as a subspecies of carinicaudus. Revalidated as a separate species by Deiques & Cechin (1991).
Distribution: Argentina, Brazil (Mato Grosso do Sul, Parana, Rio Grande do Sul, Santa Catarina, Sao Paulo), Paraguay, Uruguay.


Helicops leopardinus

Leopard Keelback, Spotted Keelback

Leopard-Kielschwanznatter

Leopard-vandsnog

1837 Homalopsis leopardina Schlegel
Helicops leopardinus Jan 1865
1854 Helicops leprieurii Duméril, Bibron & Duméril (Peters & Orejas-Miranda 1970)

Distribution: Argentina, Bolivia, Brazil (Alagoas, Amapa, Amazonas, Bahia, Ceara, Distrito Federal, Maranhao, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Para, Marajo Island, Paraiba, Pernambuco, Piaui, Rio Grande do Norte, Rondonia, Sergipe, Tocantins), Colombia, Ecuador, French Guiana, Paraguay, Peru, Surinam.


Helicops modestus

Brazilian Olive Keelback

Brasilianische Braune Scheelaugennatter

Brasiliansk Brun Vandsnog

1861 Helicops modestus Günther
1866 Helicops assimilis Reinhardt (Peters & Orejas-Miranda 1970)

Distribution: Brazil (Bahia, Distrito Federal, Goias, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Para, Parana, Sao Paulo).


Helicops nentur

Minas Gerais Keelback

Minas-Gerais-Scheelaugennatter

Minas Gerais-vandsnog

2016 Helicops nentur Costa, Santana, Leal, Koroiva & Garcia

Distribution: Brazil (Minas Gerais).


Helicops pastazae

Pastaza River Keelback

Pastaza-Scheelaugennatter

Pastaza-vandsnog

1934 Helicops pastazae Shreve

Distribution: Colombia, Ecuador, Peru, Venezuela.


Helicops petersi

Napo Keelback

Napo-Scheelaugennatter

Napo-vandsnog

1976 Helicops petersi Rosmann

Distribution: Ecuador.


Helicops polylepis

South American Many-scaled Keelback

Vielschuppige Scheelaugennatter

Mangeskćllet Vandsnog

1861 Helicops polylepis Günther
1862 Tachynectes chrysostictus Cope (Peters & Orejas-Miranda 1970)
1865 Helicops spixii Jan (Peters & Orejas-Miranda 1970)

Distribution: Bolivia, Brazil (Acre, Amapa, Amazonas, Goias, Mato Grosso, Mato Grosso do Sul, Para [incl. Marajo Island], Rondonia, Tocantins), Colombia, Peru.


Helicops scalaris

Ladder Keelback

Treppen-Scheelaugennatter

Trappe-vandsnog

1865 Helicops scalaris Jan

Distribution: Colombia, Venezuela.


Helicops tapajonicus

Tapajós River Keelback

Tapajós-Scheelaugennatter

Tapajós-vandsnog

2005 Helicops tapajonicus Da Frota

Distribution: Brazil (Para).


Helicops trivittatus

Three-striped Keelback

Dreistreifige Scheelaugennatter

Trestribet Vandsnog

1849 Myron trivittatus Gray
Helicops trivittatus Boulenger 1863

Distribution: Brazil (Amapa, Goias, Maranhao, Mato Grosso, Para [incl. Marajo Island], Tocantins).


Helicops yacu

Loreto Keelback

Loreto-Scheelaugennatter

Loreto-vandsnog

1975 Helicops yacu Rossman & Dixon
Hydrops yacu [lapsus?] Henle & Ehrl 1991

Remarks: Rossman & Abe (1979) suggested that yacu might be as subspecies of pastazae. Presence in Brazil reported by Rossman & Abe (1979), but apparently overlooked by subsequent workers (e.g., Costa & Bernils 2015).
Distribution: Brazil (Acre), Peru.