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Taxonomy of the family Teiidae
Bibliography of the genus Holcosus
Biodiversity of the family Teiidae








Genus
Holcosus

Northwestern Ameivas*

Nordwestliche Ameiven*

Nordvestlige Ameivaer*

1862 Holcosus Cope (type species: Ameiva septemlineata Dumeril 1851)
Contents: 19 species, of which 10 (52.6%) are endemic.
Remarks: Previously regarded as a synonym of Ameiva (e.g., Peters & Donoso-Barros 1970). Revalidated by Harvey, Ugueto & Gutberlet (2012). This generic taxonomic arrangement was considered unwarranted by Giugliano, Nogueira, Valdujo, Collevatti & Colli (2013), but the validity of Holcosus was supported by Tucker, Colli, Giugliano, Hedges, Hendry, Lemmon, Lemmon, Sites & Pyron (2016).
Introduced to: Nicaragua (Islas del Maiz: Isla Pequeńa del Maiz).
Distribution: Mexico, Central America, NW. South America.
Reported from: Belize, Colombia (incl. Gorgona Island), Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico (Campeche, Chiapas, Colima, Guerrero, Hidalgo, Jalisco, Michoacan, Nayarit, Oaxaca, Puebla, Queretaro, Quintana Roo, San Luis Potosi, Tabasco, Tamaulipas, Veracruz, Yucatan), Nicaragua (incl. Islas del Maiz: Isla Grande del Maiz), Panama.

Holcosus amphigrammus

Eastern Mexican Ameiva*

Ostmexicanische Ameive*

Řstmexicansk Ameiva*

1945 Ameiva undulata amphigramma Smith & Laufe
Holcosus undulatus amphigrammus Harvey, Ugueto & Gutberlet 2012*
Holcosus amphigrammus Meza-Lázaro & Nieto-Montes de Oca 2015
1946 Ameiva undulata podarga Smith & Laufe (Meza-Lázaro & Nieto-Montes de Oca 2015)
Holcosus undulatus podargus Harvey, Ugueto & Gutberlet 2012*

Remarks: Current presence in Nuevo Leon needs confirmation, since the species has not been found in the state since 1934 (Lemos-Espinal, Wylie & Smith 2017).
Distribution: Mexico (Hidalgo, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tabasco, Tamaulipas, Veracruz).


Holcosus anomalus

Cauca Ameiva*, (Echternacht's Ameiva)

Cauca-Ameive*

Cauca-ameiva*

1977 Ameiva anomala Echternacht
Holcosus anomalus Harvey, Ugueto & Gutberlet 2012

Distribution: Colombia.


Holcosus bridgesii

Colombian Ameiva*

Kolumbianische Ameive*

Colombiansk Ameiva*

1869 Holcosus bridgesii Cope
Ameiva bridgesii Barbour & Noble 1915

Distribution: Colombia (incl. Gorgona Island), Ecuador.


Holcosus chaitzami

Alta Verapaz Ameiva*, (Chaitzam's Ameiva)

Alta-Verapaz-Ameive*

Alta Verapaz-ameiva*

1924 Ameiva chaitzami Stuart
Holcosus chaitzami Harvey, Ugueto & Gutberlet 2012

Remarks: Previously included thomasi.
Distribution: Guatemala, Mexico (Chiapas).


Holcosus festivus

Central American Ameiva, Central American Whiptail, Middle American Ameiva, Tiger Ameiva, Festive Ameiva

Zentralamerikanische Ameive

Centralamerikansk Ameiva*

1856 Cnemidophorus festivus Lichtenstein
Ameiva festivus Bocourt 1874
Holcosus festivus Harvey, Ugueto & Gutberlet 2012
1862 Ameiva eutropia Cope (Peters & Donoso-Barros 1970; Echternacht 1974)
1873 Ameiva edwardsii Bocourt (Peters & Donoso-Barros 1970)
Ameiva festiva edwardsii Stuart 1943
Holcosus festivus edwardsii Harvey, Ugueto & Gutberlet 2012*
1956 Ameiva festiva occidentalis Taylor (Peters & Donoso-Barros 1970)
Holcosus festivus occidentalis Harvey, Ugueto & Gutberlet 2012*

Other common names:
edwardsii: Edwards' Middle American Ameiva
Distribution: Belize, Colombia, Costa Rica, Guatemala, Honduras, Mexico (Chiapas, Tabasco), Nicaragua, Panama.


Holcosus gaigeae

Yucatan Ameiva*

Yucatan-Ameive*

Yucatan-ameiva*

1946 Ameiva undulata gaigeae Smith & Laufe
Holcosus undulatus gaigeae Harvey, Ugueto & Gutberlet 2012*
Holcosus gaigeae Meza-Lázaro & Nieto-Montes de Oca 2015

Remarks: Not mentioned for Yucatan by González-Sánchez, Johnson, García-Padilla, Mata-Silva, DeSantis & Wilson (2017), although reported in several works (e.g., Smith & Taylor 1950; Duellman 1965; Dundee, White & Rico-Gray 1986; Meza-Lázaro & Nieto-Montes de Oca 2015).
Distribution: Belize, Guatemala, Mexico (Campeche, Quintana Roo, Yucatan).


Holcosus hartwegi

Northern Chiapan Ameiva*

Nördlicher Chiapas-Ameive

Nordlig Chiapas-ameiva*

1946 Ameiva undulata hartwegi Smith
Holcosus undulatus hartwegi Harvey, Ugueto & Gutberlet 2012*
Holcosus hartwegi Meza-Lázaro & Nieto-Montes de Oca 2015

Distribution: Guatemala, Honduras, Mexico (Chiapas, Campeche, Quintana Roo, Yucatan).


Holcosus leptophrys

Delicate Ameiva

Costa-Rica-Ameive*

Costa Rica-ameiva*

1893 Amiva leptophrys Cope
Holcosus leptophrys Harvey, Ugueto & Gutberlet 2012
1915 Ameiva ruthveni Barbour & Noble (Peters & Donoso-Barros 1970)

Distribution: Colombia, Costa Rica, Panama.


Holcosus miadis

Islas del Maiz Ameiva*

Islas-del-Maiz-Ameive*

Islas del Maiz-ameiva*

1929 Ameiva festiva miadis Barbour & Loveridge
Ameiva undulata miadis Peters & Donoso-Barros 1970
Holcosus undulatus miadis Harvey, Ugueto & Gutberlet 2012*
Holcosus miadis Meza-Lázaro & Nieto-Montes de Oca 2015*

Introduced to: Nicaragua (Islas del Maiz: Isla Pequeńa del Maiz).
Distribution: Nicaragua (Islas del Maiz: Isla Grande del Maiz).


Holcosus niceforoi

Sasaima Ameiva*

Sasaima-Ameive*

Sasaima-ameiva*

1943 Ameiva festiva niceforoi Dunn
Ameiva niceforoi Echternacht 1970
Holcosus niceforoi Harvey, Ugueto & Gutberlet 2012

Distribution: Colombia.


Holcosus orcesi

Azuay Ameiva*

Azuay-Ameive*

Azuay-ameiva*

1964 Ameiva orcesi Peters
Holcosus orcesi Harvey, Ugueto & Gutberlet 2012

Distribution: Ecuador.


Holcosus parvus

Guatemalan Rainbow Ameiva*

Guatemalischer Regenbogen-Ameive*

Guatemalansk Regnbue-ameiva*

1915 Ameiva undulata parva Barbour & Noble
Holcosus undulatus parvus Harvey, Ugueto & Gutberlet 2012*
Holcosus parvus Meza-Lázaro & Nieto-Montes de Oca 2015

Distribution: Costa Rica, El Salvador, Guatemala, Mexico (Chiapas, Oaxaca).


Holcosus pulcher

Nicaraguan Rainbow Ameiva*

Nicaraguanischer Regenbogen-Ameive*

Nicaraguansk Regnbue-ameiva*

1861 Ameiva pulchra Hallowell
Ameiva undulata pulchra Stuart 1942
Holcosus undulatus pulcher Harvey, Ugueto & Gutberlet 2012*
Holcosus pulcher Meza-Lázaro & Nieto-Montes de Oca 2015*

Remarks: Status uncertain, but tentatively regarded as a valid species herein, following Meza-Lázaro & Nieto-Montes de Oca (2015), who concluded that pulcher is unlikely to be conspecific with undulatus.
Distribution: Nicaragua.


Holcosus quadrilineatus

Four-lined Ameiva

Vierstreifen-Ameive*

Firstribet Ameiva*

1861 Cnemidophorus quadrilineatus Hallowell
Ameiva quadrilineata Cope 1862
Holcosus quadrilineatus Harvey, Ugueto & Gutberlet 2012
1876 Amiva gabbiana Cope (Peters & Donoso-Barros 1970)

Distribution: Costa Rica, Nicaragua, Panama.


Holcosus septemlineatus

Seven-lined Ameiva

Siebenstreifige Ameive*

Syvstribet Ameiva*

1851 Ameiva septemlineata Dumeril
Holcosus septemlineatus Harvey, Ugueto & Gutberlet 2012
1859 Ameiva sex-scutata Günther (Peters & Donoso-Barros 1970)

Distribution: Colombia, Ecuador.


Holcosus sinister

Western Mexican Ameiva*

Westmexicanische Ameive*

Vestmexicansk Ameiva*

1946 Ameiva undulata sinistra Smith & Laufe
Holcosus undulatus sinistrus Harvey, Ugueto & Gutberlet 2012*
Holcosus sinister Meza-Lázaro & Nieto-Montes de Oca 2015

Remarks: Includes older, northwestern records of dextra (see undulatus), but these records may represent a distinct species (Meza-Lázaro & Nieto-Montes de Oca 2015). Records from Morelos (e.g., Smith & Laufe 1946; Smith & Taylor 1950; Davis & Smith 1953), are erroneous (Echternacht 1971; Castro-Franco & Bustos-Zagal 1994).
Distribution: Mexico (Colima, Guerrero, Jalisco, Michoacan, Nayarit, Puebla).


Holcosus stuarti

Palenque Ameiva*

Palenque-Ameive*

Palenque-ameiva*

1940 Ameiva undulata stuarti Smith
Holcosus undulatus stuarti Harvey, Ugueto & Gutberlet 2012*
Holcosus stuarti Meza-Lázaro & Nieto-Montes de Oca 2015

Distribution: Mexico (Campeche, Chiapas, Tabasco).


Holcosus thomasi

Rio Grijalva Ameiva*

Rio-Grijalva-Ameive*

Rio Grijalva-ameiva*

1946 Ameiva undulata thomasi Smith & Laufe
Holcosus chaitzami thomasi Harvey, Ugueto & Gutberlet 2012*
Holcosus thomasi Meza-Lázaro & Nieto-Montes de Oca 2015

Remarks: Previously regarded as a subspecies of undulata (e.g., Peters & Donoso-Barros 1970) or a synonym of chaitzami (e.g., Echternacht 1970). Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Distribution: Guatemala, Mexico (Chiapas).


Holcosus undulatus

Mexican Rainbow Ameiva*, Metallic Ameiva

Mexicanische Regenbogen-Ameive*

Mexicansk Regnbue-ameiva*

1834 Cnemidophorus undulatus Wiegmann
Ameiva undulata Gray 1845
Holcosus undulatus Harvey, Ugueto & Gutberlet 2012
1946 Ameiva undulata dextra Smith & Laufe (Meza-Lázaro & Nieto-Montes de Oca 2015, see remarks)
Holcosus undulatus dextrus Harvey, Ugueto & Gutberlet 2012*

Remarks: Previously included amphigrammus, gaigeae, hartwegi, miadis, parvus, pulcher, sinister, stuarti, and thomasi. Northwestern records of dextra refer to sinister, but may represent a distinct species (Meza-Lázaro & Nieto-Montes de Oca 2015).
Distribution: Mexico (Guerrero, Oaxaca).