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Taxonomy of the genus Naja Biodiversity of the family Elapidae [terrestrial species]

Bibliography of the genus
Naja (Typical Cobras)

(Reptilia: Serpentes: Elapidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Naja in general

Anonymous. 1925. Tiger killed by a cobra. Journal of the Bombay Natural History Society 30: 705-706.

Anonymous. 1967. Male cobra looks for its dead mate. Loris 11: 124.

Ashe, J. 1984. Cobras. Swara 7(3): 15-18.

Avadhani, R. 2011. Kobras in der hinduistischen Kultur. Reptilia (D) 16(89): 29-34.

Bannerman, W.B. 1905. On the distribution of the varieties of cobra (Naia tripudians) in India. Journal of the Bombay Natural History Society 16: 638-643.

Bannerman, W.B. 1907. A further note on the distribution of the varieties of cobra in India. Journal of the Bombay Natural History Society 17: 1031-1032.

Bannerman, W.B.; Pocha, J.P. 1912. Cobra breeding at Paul. Journal of the Bombay Natural History Society 21: 1337-1339.

Barnard, H.A. 1978. Cobra reminiscences. Loris 14(6): 368-370.

Bethencourt-Ferreira, J. 1901. Sôbre a distribução das cobras do gênero 'Naia' em Africa. Journal Sci. Lisb. (2) 6: 129-137.

Bharatan, E. 1971. Cobra and monitor lizard. Journal of the Bombay Natural History Society 68(3): 838-839.

Bogert, C.M. 1954. Snakes that spit their venom. Animal Kingdom 57: 66-70.

Bouillon, A. 1965. Serpentes danger! Zooleo 2: 1-76.

Brien, E.O. 1923. Cobra going down a hole tail foremost. Journal of the Bombay Natural History Society 29: 303.

Broadley, D.G. 1966. Notes on the taxonomy and distribution of the African cobras of the genus Naja. Journal of the Herpetological Association of Africa 2: 21-29.

Broadley, D.G. 1968. A review of the African cobras of the genus Naja (Serpentes: Elapinae). Arnoldia (Rhodesia) 3(29): 1-14.

Broadley, D.G.; Wüster, W. 2004. A review of the southern African 'non-spitting' cobras (Serpentes: Elapidae: Naja). African Journal of Herpetology 53(2): 101-122.

Brongersma, L.D. 1947. Zoological notes from Port Dickson. 1. Amphibians and reptiles. Zoologische Mededelingen (Leiden) 27: 300-308.

Cotton, T. 2014. Comments on Spracklandus Hoser, 2009 (Reptilia, Serpentes, ELAPIDAE): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published (Case 3601). Bulletin of Zoological Nomenclature 71(3): 181-182.

Deraniyagala, P.E.P. 1932. Herpetological Notes. Ceylon Journal of Science (B) 17: 44-55.

Deraniyagala, P.E.P. 1960. The taxonomy of the cobras of southeastern Asia. Spolia Zeylanica 29: 41-63.

Deraniyagala, P.E.P. 1961. The taxonomy of the cobras of south eastern Asia. Part 2. Spolia Zeylanica 29: 205-232.

Duckett, J.E. 1987. The common cobra (Naja naja) and its hood markings in Malaysia. Malayan Naturalist 40(3-4): 46-47.

Elliot, R.H. 1933. The mongoose and the cobra. Blackwood's 1933: 662.

Ferreira, J.B. 1930. Some notes on the habits and peculiarities of the African snakes of the genus Naja, their taxonomic status and geographical distribution. Bulletin of the Antivenin Institute of America 4: 54-57.

Fukuyama, T.; Sawai, Y.; Okonogi, T. 1974. Comparative study of necrosis induced by Naja naja naja, Naja n. atra, Naja n. kaouthia and Naja n. philippinensis. The Snake 6(2): 83-85.

George, C.P. 1904. A cobra feeding on eggs. Journal of the Bombay Natural History Society 16: 174.

Ghosh, S.K. 1946. Short note about the black cobra. Journal Bengal Nat. Hist. Soc. 20(3): 105-108.

Heurn, W.C. van 1931. Herpetologische aanteekeningen. II, IV. De Tropische Natuur 20: 61-63, 205-209.

Horton, C.W. 1948. On the mechanics of spitting in the African spitting Cobras. Copeia 1948: 23-26.

Hoser, R.T. 2009. A reclassification of the true cobras; species formerly referred to the genera Naja, Boulengerina and Paranaja. Australasian Journal of Herpetology 7: 1-15.

Hoser, R.T. 2012. Exposing a fraud! Afronaja Wallach, Wüster and Broadley 2009, is a junior synonym of Spracklandus Hoser 2009. Australasian Journal of Herpetology 2012(9): 1-64.

Hoser, R.T. 2013. Case 3601. Spracklandus Hoser, 2009 (Reptilia, Serpentes, Elapidae): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published. Bulletin of Zoological Nomenclature 70(4): 234-237.

Hoser, R.T. 2013. Two new species of true cobra in the genus Boulengeria Dollo, 1886 from West Africa and South Africa (Serpentes: Elapidae). Australasian Journal of Herpetology 20: 3-7.

Hoser, R.T. 2015. Comment on Spracklandus Hoser, 2009 (Reptilia, Serpentes, Elapidae): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published (Case 3601). Bulletin of Zoological Nomenclature 72(1): 61-64.

Kaiser, H. 2014. Comments on Spracklandus Hoser, 2009 (Reptilia, Serpentes, Elapidae): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published. (Case 3601). Bulletin of Zoological Nomenclature 71(1): 30-35.

Khan, M.S. 1984. A cobra with an unusual hood pattern. The Snake 16(2): 131-134.

Lauraine, P. 1931. The snake charming sisters of Holy Popa: women as hereditary handlers of cobras. Ill. London News Oct. 3: 528-529.

Laurent, R.F. 1973. Le concept de sous-espèce et la variation géographique chez Naja melanoleuca et Naja nigricollis (Reptilia, Ophidia). Revue de Zoologie et Botanique Africaine (Bruxelles) 87(1): 198-204.

Leviton, A.E. 1964. Contribution to a review of Philippine snakes. 7. The snakes of the genera Naja and Ophiophagus. Philippine Journal of Science 93: 531-550.

Loveridge, A. 1944. Further revisions of African snake genera. Bulletin of the Museum of Comparative Zoology at Harvard 95: 121-247.

Nutphand, W. 1986. Cobra. Thai Zoological Center, Bangkok. Booklet no. 1(12). 15 pp.

Perret, J.L. 1960. Une nouvelle et remarkable espèce d'Atractaspis (Viperidae) et quelques autres serpents d'Afrique. Revue Suisse de Zoologie 67(5): 129-139.

Rhodin, A.G.J.; Kaiser, H.; Dijk, P.P. van; Wüster, W.; O'Shea, M.; Auliya, M. & al. 2015. Comment on Spracklandus Hoser, 2009 (Reptilia, Serpentes, Elapidae): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published (Case 3601). Bulletin of Zoological Nomenclature 72(1): 65-78.

Roman, B. 1973. Une collection de serpents peu commune. Notes et Documents Voltaiques 6(2): 36-58.

Shine, R.; Branch, W.R.; Webb, J.K.; Harlow, P.S.; Shine, T.; Keogh, J.S. 2007. Ecology of cobras from southern Africa. Journal of Zoology (London) 272(2): 183-193.

Smith, M.A. 1946. Cobras and king cobras. Zoo Life 1: 39-42.

Stejneger, L.H. 1936. Types of the amphibian and reptilian genera proposed by Laurenti in 1768. Copeia 1936(3): 133-141.

Stumpel, A.H.P. 1981. Water snake (Xenochrophis piscator) imitates cobra. (Reptilia: Serpentes: Colubridae). Salamandra 17(3-4): 203-204.

Szyndlar, Z. 1992. Osteology of African cobras: preliminary observations. pp. 425-428. In: Korsos, Z. & Kiss, I. (eds.): Proceedings of the 6th Ordinary General Meeting of the Societas Europaea Herpetologica, 19-23 August 1991, Budapest, Hungary. Hungarian Natural History Museum, Budapest. 531 pp.

Szyndlar, Z.; Rage, J.C. 1990. West Palearctic cobras of the genus Naja (Serpentes: Elapidae): interrelationships among extinct and extant species. Amphibia-Reptilia 11(4): 385-400.

Thomson, S. 2014. Comment on Spracklandus Hoser, 2009 (Reptilia, Serpentes, Elapidae): request for confirmation of the availability of the generic name and for the nomenclatural validation of the journal in which it was published. (Case 3601). Bulletin of Zoological Nomenclature 71(2): 133-135.

Tscherbakoff, S.G. 1935. Feeding of cobras in captivity. Journal of the Bombay Natural History Society 38: 321-329.

Vijayaraghavan, B. 2005. Are there spitting cobras in India? Cobra (Chennai) 61: 1-5.

Vozenilek, P. 1976. [Spitting cobras]. (In Czech). Ziva 24(1): 28-29.

Wallach, V.; Wüster, W.; Broadley, D.G. 2009. In praise of subgenera: taxonomic status of cobras of the genus Naja Laurenti (Serpentes: Elapidae). Zootaxa 2236: 26-36.

Werner, Y.L. 1999. Does the cobra obey the snake charmer's flute? Hamadryad 24(1): 23-28.

Whitaker, R. 1969. Defanged cobras, how they survive. Journal of the Bombay Natural History Society 66: 632-634.

Whitaker, R.; Whitaker, S. 2012. Venom, antivenom production and the medically important snakes of India. Current Science (Bangalore) 103(6): 635-643.

Wilks, A. 2017. Pick your poison. Bulletin of the Chicago Herpetological Society 52(7): 118-121

Woods, D.H. 1958. Cobra enters hole backwards. African Wild Life 12: 77.

Wüster, W. 1992. Cobras and other herps in Southeast Asia. British Herpetological Society Bulletin 39: 19-25.

Wüster, W. 1996. Taxonomic changes and toxinology: systematic revisions of the asiatic cobras (Naja naja species complex). Toxicon 34(4): 399-406.

Wüster, W. 2002. The cobras of the genus Naja in India. pp. 98-133. In: Sharma, B.D. (ed.). Indian poisonous snakes: an ecological and clinical study. Anmol Publications, New Delhi. 341 pp.

Wüster, W.; Chirio, L.; Trape, J.F.; Ineich, I.; Jackson, K.; Greenbaum, E.; Barron, C.; Kusamba, C.; Nagy, Z.T.; Storey, R.; Hall, C.; Wüster, C.E.; Barlow, A.; Broadley, D.G. 2018. Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455(1): 68–98.

Wüster, W.; Crookes, S.; Ineich, I.; Mané, Y.; Pook, C.E.; Trape, J.F.; Broadley, D.G. 2007. The phylogeny of cobras inferred from mitochondrial DNA sequences: Evolution of venom spitting and the phylogeography of the African spitting cobras (Serpentes: Elapidae: Naja nigricollis complex). Molecular Phylogenetics and Evolution 45(2): 437-453.

Wüster, W.; Thorpe, R.S. 1987. Geographical variation in cobras of the genus Naja in southeast Asia: a multivariate analysis. pp. 449-452. In: van Gelder, J.J., Strijbosch, H. & Bergers, P.J.M. (eds.). Proceedings of the 4th Ordinary General Meeting of the Societas Europaea Herpetologica. Societas Europaea Herpetologica, Faculty of Sciences, Nijmegen. 473 pp.

Wüster, W.; Thorpe, R.S. 1989. Population affinities of the asiatic cobra (Naja naja) species complex in south-east Asia: reliability and random resampling. Biological Journal of the Linnean Society 36: 391-409.

Wüster, W.; Thorpe, R.S. 1991. Asiatic cobras: systematics and snakebite. Experientia (Basel) 47(2): 205-209.

Wüster, W.; Thorpe, R.S. 1992. Asiatic cobras: population systematics of the Naja naja species complex (Serpentes: Elapidae) in India and Central Asia. Herpetologica 48(1): 69-85.

Wüster, W.; Thorpe, R.S. 1992. Dentitional phenomena in cobras revisited: spitting and fang structure in the Asiatic species of Naja (Serpentes: Elapidae). Herpetologica 48(4): 424-434.

Wüster, W.; Thorpe, R.S.; Cox, M.J.; Jintakune, P.; Nabhitabhata, J. 1995. Population systematics of the snake genus Naja (Reptilia: Serpentes: Elapidae) in Indochina: multivariate morphometrics and comparative mitochondrial DNA sequencing (cytochrome oxidase 1). Journal of Evolutionary Biology 8(4): 493-510.

Young, B.A.; Dunlap, K.; Koenig, K.; Singer, M. 2004. The buccal buckle: the functional morphology of venom spitting in cobras. Journal of Experimental Biology 207(20): 3483-3494.

Zug, G.R. 2014. Comment on the proposed confirmation of the availability of Spracklandus Hoser, 2009 (Reptilia, Squamata, ELAPIDAE). (Case 3601). Bulletin of Zoological Nomenclature 71(4): 252-253.

Naja anchietae

Broadley, D.G. 1961. Three new snakes added to the Southern Rhodesia list. Journal of the Herpetological Association of Rhodesia 15(1): 10-12.

Broadley, D.G.; Wüster, W. 2004. A review of the southern African 'non-spitting' cobras (Serpentes: Elapidae: Naja). African Journal of Herpetology 53(2): 101-122.

Werner, F. 1903. Über Reptilien und Batrachier aus Guatemala und China in der zoologischen Staats-Sammlung in München, nebst einem Anhang über seltene Formen aus anderen Gebieten. Abhandlungen der Bayerischen Akademie der Wissenschaften 22(2): 343-384.

Naja annulata

Bouillon, A. 1965. Serpentes danger! Zooleo 2: 1-76.

Boulenger, G.A. 1895. On the type specimen of Boulengerina stormsi, an elapoid snake from Central Africa. Proceedings of the Zoological Society of London 1895: 865-866.

Branch, W.R.; Conradie, W. 2013. Geographical distribution: Naja annulata annulata (Bucholtz & Peters, 1876). Angola. African Herp News 59: 50-53.

Cochran, D.M. 1944. Dangerous reptiles (popular). Rep. Smithson. Instn. 1943: 275-325.

Dainty, M. 2011. Die Gebänderte Wasserkobra, Boulengerina annulata. Reptilia (D) 16(89): 24-28.

Ditmars, R.L. 1935. Snakes and surprises. Bulletin of the New York Zoological Society 38: 101-105.

Ionides, C.J.P.; Pitman, C.R.S. 1965. Notes on three East African venomous snake populations. Puku 3: 87-95.

Kauffeld, C.F. 1947. The Hooden Clan. Fauna (Philadelphia) 9: 82-87.

Kratzer, H. 1965. Über die Tanganyika-Wasserkobra (Boulengerina annulata stormsi). Salamandra 1: 61-67.

Kudryavtsev, S.V.; Mamet, S.V.; Ganina, L.V. 2008. [Some successful results in husbandry and breeding of the rare and hard-to-keep venomous snakes at the Moscow zoo (part YIII: 2007-2008 years)]. (In Russian). Nauchnye Issledovaniya v Zoologicheskikh Parkakh 24: 9-18.

Laurent, R.F. 1952. Batraciens et reptiles recemment acquis par le Musee du Congo Belge. Revue de Zoologie et Botanique Africaine (Bruxelles) 45: 198-203.

Loveridge, A. 1931. On two amphibious snakes of the Central African Lake Region. Bulletin of the Antivenin Institute of America 5(1): 7-12.

Loveridge, A. 1960. The water-cobra of Lake Tanganyika. Journal of the East Africa Natural History Society 23(5): 189.

Newby, J.E. 1961. Snake catching at Lake Tanganyika. Nat. Soc. Cent. Afr. J. 2: 11-12.

O'Shea, M.; McIntyre, P.B. 2005. Natural history notes: Boulengerina annulata stormsi (Storm's Water Cobra). Attempted predation. Herpetological Review 36(2): 189.

Pauwels, O.S.G.; Albert, J.L.; Arrowood, H.; Mvele, C.; Casanova, M.; Dodane, J.B.; Morgan, J.; Primault, L.; Thepenier, L.; Fenner, J.N. 2017. Miscellanea Herpetologica Gabonica X. Bulletin of the Chicago Herpetological Society 52(8): 133-138.

Pauwels, O.S.G.; Lavoue, S. 2004. Geographic distribution: Boulengerina annulata annulata (Banded Water Cobra). Herpetological Review 35(3): 290.

Perret, J.L. 1961. Études herpétologiques Africaines. III. Bulletin Soc. Neuchâtel. Sci. Nat. (3) 84: 133-138.

Pickford, N. 1970. Storm's water cobra. Africana 4(1): 16-17.

Shaw, C.E. 1963. Notes on the eggs, incubation and young of some African reptiles. British Journal of Herpetology 3: 63-71.

Vesey-Fitzgerald, D. 1959. The water cobras of Mpulungu. Journal of the Herpetological Association of Rhodesia 8: 5-6.

Naja annulifera

Broadley, D.G. 1995. The snouted cobra, Naja annulifera, a valid species in southern Africa. Journal of the Herpetological Association of Africa 44(2): 26-32.

Broadley, D.G.; Wüster, W. 2004. A review of the southern African 'non-spitting' cobras (Serpentes: Elapidae: Naja). African Journal of Herpetology 53(2): 101-122.

Phillips, M.A.; Waterman, J.M. 2014. Anti-snake behaviour in a facultative cooperative breeder, the Cape Ground Squirrel. Behaviour 151(12-13): 1735-1758.

Sauer, F. 1959. Naja haje annulifera Peters in Südwest-Afrika. Zoologischer Anzeiger 162: 113-116.

Naja arabica

Al Asmari, A.; Rajamohammed, A.M.; Khan, H.A. 2014. Identification and phylogeny of Arabian snakes: Comparison of venom chromatographic profiles versus 16S rRNA gene sequences. Saudi Journal of Biological Sciences 21(5): 436-441.

Els, J. 2011. First record on the production of the Arabian cobra (Naja arabica) (Scortecci, 1932) in captivity. African Herp News 54: 4-6.

Haas, G. 1957. Some amphibians and reptiles from Arabia. Proceedings of the California Academy of Sciences (Ser. 4) 29(4): 47-86.

Kowalski, T.; Grossmann, W. 2016. Das Porträt: Naja arabica Scortecci, 1932. Sauria (Berlin) 38(4): 1-2.

Trape, J.F.; Chirio, L.; Broadley, D.G.; Wüster, W. 2009. Phylogeography and systematic revision of the Egyptian cobra (Serpentes: Elapidae: Naja haje) species complex, with the description of a new species from West Africa. Zootaxa 2236: 1-25.

Naja ashei

Branch, W.R.; Menegon, M.; Beraducci, J. 2010. Geographical distribution: Naja ashei Wüster & Broadley, 2007. Ashe's Spitting Cobra. Tanzania. African Herp News 52: 21-23.

Wüster, W.; Broadley, D.G. 2007. Get an eyeful of this: a new species of giant spitting cobra from eastern and north-eastern Africa (Squamata: Serpentes: Elapidae: Naja). Zootaxa 1532: 51-68.

Naja atra

Anderson, L.A.; Dufton, M.J. 1997. The action of Taiwan cobra venom on methionine enkephalin: a useful assay for oligopeptidase content. Toxicon 35(7): 1113-1123.

Chiou, S.H.; Hung, C.C.; Huang, H.C.; Chen, S.T.; Wang, K.T.; Yang, C.C. 1995. Sequence comparison and computer modelling of cardiotoxins and cobrotoxin isolated from Taiwan cobra. Biochemical and Biophysical Research Communications 206(1): 22-32.

Fletcher, J.E.; Jiang, M.S. 1995. Presynaptically acting snake venom phospholipase A2 enzymes attack unique substrates. Toxicon 33(12): 1565-1576.

Herklots, G.A.C. 1938. Land snakes of Hongkong. IV. Naja. Hong Kong Naturalist 8(3-4): 189-200.

Hu, B.C.; Hwang, M.H.; Ho, S.S.; Chou, S.A.; Hsieh, T.T.; Tsai, B. 1966. A preliminary report on some ecological observations of Agkistrodon halys and Naja naja atra. Acta Zoologica Sinica 18: 187-194.

Ji, X.; Chen, H.L.; Du, W.G.; Zhu, B.Q. 2002. [Radiotelemetry of thermoregulation and thermal tolerance on Chinese cobras (Naja atra) overwintering in a laboratory enclosure]. (In Chinese, English summary). Acta Zoologica Sinica 48(5): 591-598.

Ji, X.; Du, W.G. 2001. Effects of thermal and hydric environments on incubating eggs and hatchling traits in the cobra, Naja naja atra. Journal of Herpetology 35(2): 186-194.

Ji, X.; Du, W.G.; Qu, Y.F.; Lin, L.H. 2009. Nonlinear continuum of egg size-number trade-offs in a snake: is egg-size variation fitness related? Oecologia (Berlin) 159(4): 689-696.

Ji, X.; Du, W.G.; Xu, W.Q. 1999. Experimental manipulation of egg size and hatchling size in the cobra, Naja naja atra (Elapidae). Netherlands Journal of Zoology 49(3): 167-175.

Ji, X.; Wang, Z.W. 2005. Geographic variation in reproductive traits and trade-offs between size and number of eggs of the Chinese cobra (Naja atra). Biological Journal of the Linnean Society 85(1): 27-40.

Lin, L.H.; Hua, L.; Qu, Y.F.; Gao, J.F.; Ji, X. 2014. The phylogeographical pattern and conservation of the Chinese Cobra (Naja atra) across its range Based on mitochondrial control region sequences. PLoS ONE 9(9): e106944.

Lin, L.H.; Ji, X.; Lu, H.L.; Luo, L.G. 2005. Variation in body temperature as a consequence of the response to heating lights under a light-dark cycle in the Chinese cobra Naja atra. Acta Zoologica Sinica 51(1): 38-45.

Lin, L.H.; Qu, Y.F.; Li, H.; Zhou, K.Y.; Ji, X. 2012. Genetic structure and demographic history should inform conservation: Chinese Cobras currently treated as homogenous show population divergence. PLoS ONE 7(4): e36334, 1-9.

Lin, L.H.; Zhao, Q.; Ji, X. 2008. Conservation genetics of the Chinese cobra (Naja atra) investigated with mitochondrial DNA sequences. Zoological Science (Tokyo) 25(9): 888-893.

Mao, S.H. 1963. Variation in colour pattern and hood marking of Taiwan cobras. Bull. Inst. Zool. Academia Sinica 2: 55-59.

Mao, S.H. 1993. Common terrestrial venomous snakes of Taiwan. National Museum of Natural Science Special Publications (Taichung) 5: 1-108.

Mavromichalis, J.; Bloem, S. 1995. A successful breeding of Naja naja sputatrix atra. Litteratura Serpentium (English Edition) 15(2): 26-30.

Moriguchi, H.; Toriba, M.; Miyata, H.; Sakai, A.; Sugano, H. 1982. Notes on reproductions of captive snakes observed in the Japan Snake Institute during the years 1980-1981. The Snake 14(1): 57-60.

Olexa, A.; Stettler, P.H. 1976. Zur Fortpflanzung der Indochinesischen Kobra Naja naja atra Cantor, 1842. Aquarium Aqua Terra 10(88): 460-464.

Romer, J.D. 1958. Partial albinism in a Chinese cobra. Copeia 1958: 334.

Sivaraman, T.; Kumar, T.K.S.; Yang, P.W.; Yu, C. 1997. Cardiotoxin-like basic protein (CLBP) from Naja naja atra is not a cardiotoxin. Toxicon 35(9): 1367-1371.

Wang, Z.; Lin, L.; Ji, X. 2014. Unhatched and hatched eggshells of the Chinese Cobra Naja atra. Asian Herpetological Research 5(4): 276-280.

Xiang, J.; Sun, P.Y.; Zhang, H.S.; Fu, S.Y. 1997. Incubation and utilization of energy and material during embryonic development in eggs of Naja naja atra. Journal of Herpetology 31(2): 302-306.

Naja christyi

Bouillon, A. 1965. Serpentes danger! Zooleo 2: 1-76.

Laurent, R.F. 1956. Notes herpétologiques africaines. Revue de Zoologie et Botanique Africaine (Bruxelles) 53(3-4): 229-256.

Naja crawshayi

Anonymous. 1958. The races of Naja nigricollis occurring in Rhodesia. Journal of the Herpetological Association of Rhodesia 2(1): 5-6.

Bourgeois, M. 1964. Les reptiles de la Kasapa (Elisábethville). Publ. Univ. Elisábethville 7: 65-89.

Naja guineensis

Auliya, M.; Wagner, P.; Böhme, W. 2012. The herpetofauna of the Bijagos archipelago, Guinea-Bissau (West Africa) and a first country-wide checklist. Bonn Zoological Bulletin 61(2): 255-281.

Wüster, W.; Chirio, L.; Trape, J.F.; Ineich, I.; Jackson, K.; Greenbaum, E.; Barron, C.; Kusamba, C.; Nagy, Z.T.; Storey, R.; Hall, C.; Wüster, C.E.; Barlow, A.; Broadley, D.G. 2018. Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455(1): 68–98.

Naja haje

Alves, R. 1960. Some case histories of snake bites in Southern Rhodesia. Journal of the Herpetological Association of Rhodesia 9(1): 15-17.

Behler, J.L.; Brazaitis, P.J. 1974. Breeding the Egyptian cobra Naja haje at the New York Zoological Park. International Zoo Yearbook 14: 83-84.

Broadley, D.G. 1958. Snakes of Southern Rhodesia. 6. The egyptian cobra (Naja haje haje). African Wild Life 12: 125-128.

Broadley, D.G. 1962. The herpetofauna of the Inyanga National Park. Journal of the Herpetological Association of Rhodesia 17/18: 8-10.

Cloudsley-Thompson, J.L. 1994. Cleopatra's asp. British Herpetological Society Bulletin 50: 18-19.

Cochran, D.M. 1944. Dangerous reptiles (popular). Rep. Smithson. Instn. 1943: 275-325.

Dickinson, W.H. 1948. Observations on Naja haje (Linnaeus) feigning death. Herpetologica 4: 147.

Egan, L.; Alexander, G.J. 1997. Fang structure of Hemachatus haemachatus (Serpentes: Elapidae) - a comparison to another spitter and non-spitter. pp. 157-158. In: Wyk, J.H. van (ed.). Proceedings of the FitzSimons Commemorative Symposium (South African lizards: 50 years of progress) and Third H.A.A. Symposium on African herpetology: held at the Transvaal Museum, Pretoria, South Africa, 11-15 October 1993. Herpetological Association of Africa, Matieland. 227 pp.

FitzSimons, V. 1956. Reptiles and Amphibians of the Wonderboom Nature Reserve. Fauna and Flora (Pretoria) 7: 89-106.

Foley, H.; Parrot, L. 1935. Sur la repartition du Naja haje L. (ophidiens proteoglyphes) en Algerie. Bull. Soc. Hist. nat. Alger 26(6-7): 106-168.

Hoge, A.R.; Nina, A.C. de M. 1964. Serpentes coietadas pelo Instituto Nacional de Pesquisas da Amazonia. Memorias do Instituto Butantan (Sao Paulo) 30 [1960-1962]: 71-96.

Hughes, D.F.; Behangana, M.; Wilber, L. 2017. Geographic distribution: Naja haje (Egyptian Cobra). Herpetological Review 48(2): 392.

Joleaud, L. 1934. Études de geographie zoologiques sur la Berberie. III. Le Naja. C. R. Ass. franc. Av. Sci. 58: 254-256.

Lambiris, A.J. 1967. Observations on the feeding behaviour of Naja haje and Naja nigricollis. Journal of the Herpetological Association of Africa 3: 36-37.

Newman, A.C. 1965. Further observations from Umtali snake park. Journal of the Herpetological Association of Rhodesia 23/24: 55-57.

Nkinin, S.W.; Chippaux, J.P.; Pietin, D.; Doljansky, Y.; Tremeau, O.; Menez, A. 1997. L'origine genetique de la variabilite des venins: impact sur la preparation des serums antivenimeux. Bulletin de la Societe de Pathologie Exotique 90(4): 277-281.

Poole, A. 1995. Ancient Egyptian snake mythology as seen at the British Museum. British Herpetological Society Bulletin 54: 37.

Scortecci, G. 1955. Viaggio di esplorazione biologica sulle montagne della Migiurtinia compiuto con il contribute del consiglio Nazionale delle Ricerche. Bollettino dei Musei e degli Istituti Biologici dell'Universita di Genova 25 [1953-1955]: 43-105.

Sosa, R.; Braga, L.; Schalk, C.M.; Pinto Ledezma, J.N. 2013. Natural history notes: Naja haje (Egyptian Cobra). Diet / ophiophagy. Herpetological Review 44(1): 155-156.

Stejneger, L.H. 1936. Types of the amphibian and reptilian genera proposed by Laurenti in 1768. Copeia 1936(3): 133-141.

Toschi, A. 1957. Sul comportamento in natura del Cobra (Naia haie (L.)). Ric. Zool. Appl. Caccia (Suppl.) 2(8): 1-8.

Trape, J.F.; Chirio, L.; Broadley, D.G.; Wüster, W. 2009. Phylogeography and systematic revision of the Egyptian cobra (Serpentes: Elapidae: Naja haje) species complex, with the description of a new species from West Africa. Zootaxa 2236: 1-25.

Vasil'ev, D.B. 1995. On the clutch size of the Egyptian cobra Naja haje haje (L.) of the Moscow Zoo. Russian Journal of Herpetology 2(2): 178.

Vernet, R. 2000. Observations preliminaires sur l'herpetofaune de la Republique de Djibouti. Bulletin de la Societe Herpetologique de France 93: 37-43.

Villiers, A. 1950. Contribution a l'étude de l'Air (Mission L. Chopard et A. Villiers). Reptiles. Ophidiens et cheloniens. Mem. Inst. Franc. Afr. Noire 10: 337-344.

Watkins-Pitchford, R. 1953. Cobras and puff-adders. Country Life 113(2924): 264-266.

Young, B.A.; Kardong, K.V. 2010. The functional morphology of hooding in cobras. Journal of Experimental Biology 213(9): 1521-1528.

Naja kaouthia

Ambu, S.; Sing, N.C.; Mokhtar, S. 1986. Growth rate of the copper-headed racer and the common cobra observed at the snake farm, Perlis, Malaysia. The Snake 18(1): 33-36.

Benke, Z. 1992. Clinical aspects and pathophysiology of a Naja naja kaouthia bite. pp. 79-82. In: Korsos, Z. & Kiss, I. (eds.): Proceedings of the 6th Ordinary General Meeting of the Societas Europaea Herpetologica, 19-23 August 1991, Budapest, Hungary. Hungarian Natural History Museum, Budapest. 531 pp.

Cox, M.J. 1989. Notes on the natural history (breeding) of Naja kaouthia and Python molurus bivittatus in Thailand. Bulletin of the Chicago Herpetological Society 24(7): 127-128.

Cox, M.J. 1995. Geographic distribution: Naja kaouthia (Monocled Cobra). Thailand: Loei Province. Herpetological Review 26(3): 156-157.

Das, S.; Biswas, A. 2017. Notes on the breeding ecology of monocled cobras (Naja kaouthia) from areas adjacent to Sundarban, India. Herpetological Bulletin 140: 33-34.

Dolezal, J.; Soukup, M. 1992. Predaceous behaviour of Indian cobra (Naja naja kaouthia): comparing of situation with and without visual information. Fauna Bohemiae Septentrionalis 17: 69-73.

Filippi, E.; Petretto, M. 2013. Natural history notes: Naja kaouthia (Monocellate Cobra). Reproduction / egg size. Herpetological Review 44(1): 156.

Fletcher, J.E.; Jiang, M.S. 1995. Presynaptically acting snake venom phospholipase A2 enzymes attack unique substrates. Toxicon 33(12): 1565-1576.

Gumprecht, A. 2009. Über das Vorkommen der Monokel-Kobra (Naja kouthia) in Nord-Thailand. Sauria (Berlin) 31(2): 60-62.

Hartmann, P.; Steiner, B. 1985. Naja naja kaouthia. Litteratura Serpentium 5(5): 203-204.

Kyi, S.W.; Zug, G.R. 2003. Unusual foraging behaviour of Naja kaouthia at the Moyingye Wetlands Bird Sanctuary, Myanmar. Hamadryad 27(2): 265-266.

Mitra, D. 1992. Snakes started mating early. Zoos' Print 7(3): 14.

Moriguchi, H.; Toriba, M.; Miyata, H.; Sakai, A.; Sugano, H. 1982. Notes on reproductions of captive snakes observed in the Japan Snake Institute during the years 1980-1981. The Snake 14(1): 57-60.

Pal, A.; Dey, S.; Roy, U.S. 2012. Seasonal diversity and abundance of herpetofauna in and around an industrial city of West Bengal, India. Journal of Applied Sciences in Environmental Sanitation 7(4): 281-286.

Panas, T.; Meesuk, W.; Chanhome, L.; Aowphol, A. 2012. Reproductive patterns of captive male and female Monocled Cobra, Naja kaouthia (Lesson, 1831). Zoological Studies 51(5): 692-700.

Prakash, S.; Mishra, A.K.; Raziuddin, M. 2012. A new record of cream coloured morph of Naja kaouthia Lesson, 1831 (Reptilia, Serpentes, Elapidae) from Hazaribag, Jharkhand, India. Biodiversity Journal 3(2): 153-155.

Santra, V.; Wüster, W. 2017. Natural history notes: Naja kaouthia (Monocled Cobra). Behavior / spitting. Herpetological Review 48(2): 455-456.

Schweizer, H. 1952. Über geschlüpfte Eigelege der Brillenschlange (Naja naja kaouthia) und der Hornviper (Aspis cerastes). DATZ (Die Aquarien- und Terrarien-Zeitschrift) 5: 19-21, 51-53.

Soderberg, P. 1970. The seven major poisonous snakes of Thailand. 1. The cobra (Naja naja kaouthia). Conservation News of South East Asia 6 [1965]: 19-21.

Strimple, P. 1992. Incubation and hatching of monacled cobra eggs. Litteratura Serpentium (English Edition) 12(4): 100.

Thomas, E. 1970. Kämpfe junger Brillenschlangen (Naja naja kaouthia). Salamandra 6: 52-54.

Trutnau, L. 2002. Beobachtungen an der Monokelkobra (Naja kaouthia Lesson, 1831) im Freiland und im Terrarium. Draco 3(4) (12): 45-49.

Wüster, W. 1998. The cobras of the genus Naja in India. Hamadryad 23(1): 15-32.

Young, B.A.; Kardong, K.V. 2010. The functional morphology of hooding in cobras. Journal of Experimental Biology 213(9): 1521-1528.

Naja katiensis

Rasmussen, J.B. 1996. Afrikanske slanger. 8. Naja katiensis. [African snakes. 8. Naja katiensis]. (In Danish, English summary). Nordisk Herpetologisk Forening 39(2): 41-44.

Trape, J.F.; Baldé, C. 2014. A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa 3900(3): 301-338.

Naja mandalayensis

Slowinski, J.B.; Wüster, W. 2000. A new cobra (Elapidae: Naja) from Myanmar (Burma). Herpetologica 56(2): 257-270.

Naja melanoleuca

Banks, C.B. 1984. Life history notes: Naja melanoleuca (Forest Cobra). Coprophagy. Herpetological Review 15(4): 113.

Haagner, G.V. 1990. The husbandry and captive propagation of forest cobras, Naja melanoleuca. Journal of the Herpetological Association of Africa 38: 57-60.

Haagner, G.V.; Carpenter, G. 1988. Notes on the reproduction of captive Forest Cobras, Naja melanoleuca (Serpentes: Elapidae). Journal of the Herpetological Association of Africa 34: 35-37.

Hinkel, H. 1987. Beuteerwerb bei einer Naja melanoleuca Hallowell, 1858 (Serpentes, Elapidae). Salamandra 23(4): 277.

Hoier, R. 1950. A travers plaines et volcans au Pare National Albert. Reptiles. pp. 119-127. Bruxelles. 174 pp.

Kauffeld, C.F. 1963. More at home with cobras. Bulletin of the Philadelphia Herpetological Society 11: 13-18.

Laurent, R.F. 1958. Notes herpetologiques africaines 2. Revue de Zoologie et Botanique Africaine (Bruxelles) 58: 115-128.

Laurent, R.F. 1973. Le concept de sous-espèce et la variation géographique chez Naja melanoleuca et Naja nigricollis (Reptilia, Ophidia). Revue de Zoologie et Botanique Africaine (Bruxelles) 87(1): 198-204.

Leloup, P. 1962. The egg-laying of Naja melanoleuca. Herpetologica 18: 71.

Lesage, F.M.C. 1956. Snake not quite charmed. Nigerian Field 21(3): 111-121.

Luiselli, L. 2002. Life-history correlates of suboptimal adaptation to rainforest biota by spitting cobras, Naja nigricollis, in southern Nigeria: comparative evidences with sympatric Forest Cobras, Naja melanoleuca. Revue d'Ecologie la Terre et la Vie 57(2): 123-133.

Luiselli, L.; Akani, G.C.; Corti, C.; Angelici, F.M. 2002. Is sexual size dimorphism in relative head size correlated with intersexual dietary divergence in West African Forest Cobras, Naja melanoleuca? Contributions to Zoology 71(4): 141-145.

Luiselli, L.; Angelici, F.M.; Akani, G.C. 2002. Comparative feeding strategies and dietary plasticity of the sympatric cobras Naja melanoleuca and Naja nigricollis in three diverging Afrotropical habitats. Canadian Journal of Zoology 80(1): 55-63.

Luiselli, L.; Bikikoro, L.; Chiyere Ude, L. 1997. Fish-eating by the cobras Naja melanoleuca and Naja nigricollis (Elapidae) in the Niger Delta (Port Harcourt, Rivers State, Nigeria). British Herpetological Society Bulletin 59: 13-14.

Menzies, J.I. 1963. West African snakes (Naja nigricollis and N. melanoleuca). Nigerian Field 28: 143.

Parker, R.H. 1962. The blacklipped forest cobra (Naja melanoleuca). African Wild Life 16: 35-38.

Pauwels, O.S.G.; Albert, J.L.; Arrowood, H.; Mvele, C.; Casanova, M.; Dodane, J.B.; Morgan, J.; Primault, L.; Thepenier, L.; Fenner, J.N. 2017. Miscellanea Herpetologica Gabonica X. Bulletin of the Chicago Herpetological Society 52(8): 133-138.

Pauwels, O.S.G.; Carlino, P.; Chirio, L.; Meunier, Q.; Okouyi Okouyi, J.V.; Orbell, C.; Rousseaux, D.; Testa, O. 2017. Miscellanea Herpetologica Gabonica IX. Bulletin of the Chicago Herpetological Society 52(6): 97-102.

Pauwels, O.S.G.; Essono, T.B.B.; Carlino, P.; Chirio, L.; Huijbregts, B.; Leuteritz, T.E.J.; Rousseaux, D.; Tobi, E.; Vigna, C.; Neer, W. van 2017. Miscellanea Herpetologica Gabonica VII. Bulletin of the Chicago Herpetological Society 52(1): 1-7.

Perret, J.L. 1961. Études herpétologiques Africaines. III. Bulletin Soc. Neuchâtel. Sci. Nat. (3) 84: 133-138.

Rahm, U. 1953. Einige Schlangen des westafrikanischen Urwaldes. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 6: 292-294.

Rasmussen, J.B. 1995. Afrikanske slanger. 7. Naja melanoleuca [African snakes. 7. Naja melanoleuca]. (In Danish, English summary). Nordisk Herpetologisk Forening 38(6): 89-96.

Shaw, C.E. 1956. Hail to the new champ! Zoonooz 29(11): 10-11.

Struhsaker, T.T.; McKey, D. 1975. Two cusimanse mongooses attack a black cobra. Journal of Mammalogy 56(3): 721-722.

Swan, M.; Banks, C. 2005. Natural history notes: Naja melanoleuca (Forest Cobra). Longevity. Herpetological Review 36(3): 325.

Trutnau, L. 1984. Erfahrungen bei der Pflege und Nachzucht von zwei afrikanischen Naja-Arten: der Schwarzweissen Hutschlange Naja melanoleuca und der Kapkobra Naja nivea. Herpetofauna (Weinstadt) 6(31): 15-26.

Tryon, B.W. 1979. Reproduction in captive forest cobras, Naja melanoleuca (Serpentes: Elapidae). Journal of Herpetology 13(4): 499-504.

Tryon, B.W.; Whitehead, J. 1986. Life history notes: Naja melanoleuca (African Forest Cobra). Caudal use. Herpetological Review 17(3): 61.

Warner, J.K. 2011. Natural history notes: Naja melanoleuca (Forest Cobra). Diet and foraging behavior. Herpetological Review 42(2): 295.

Watkins-Pitchford, R. 1953. Cobras and puff-adders. Country Life 113(2924): 264-266.

Wilson, D. 1959. The hatching and rearing of the cobra (Naja melanoleuca). British Journal of Herpetology 2: 159-162.

Wüster, W.; Chirio, L.; Trape, J.F.; Ineich, I.; Jackson, K.; Greenbaum, E.; Barron, C.; Kusamba, C.; Nagy, Z.T.; Storey, R.; Hall, C.; Wüster, C.E.; Barlow, A.; Broadley, D.G. 2018. Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455(1): 68–98.

Yeomans, L. 2011. Die Schwarzweiße Kobra, Naja melanoleuca. Teil 1. Reptilia (D) 16(89): 35-38.

Naja mossambica

Anonymous. 1958. The races of Naja nigricollis occurring in Rhodesia. Journal of the Herpetological Association of Rhodesia 2(1): 5-6.

Broadley, D.G. 1974. A review of the cobras of the Naja nigricollis complex in southwestern Africa (Serpentes: Elapidae). Cimbebasia 2(14): 155-162.

Edwards, S.J. 1988. African fish eagle preying on snake. Honeyguide 34(3): 123.

Egan, L.; Alexander, G.J. 1997. Fang structure of Hemachatus haemachatus (Serpentes: Elapidae) - a comparison to another spitter and non-spitter. pp. 157-158. In: Wyk, J.H. van (ed.). Proceedings of the FitzSimons Commemorative Symposium (South African lizards: 50 years of progress) and Third H.A.A. Symposium on African herpetology: held at the Transvaal Museum, Pretoria, South Africa, 11-15 October 1993. Herpetological Association of Africa, Matieland. 227 pp.

Hartmann, P.; Steiner, B. 1985. Naja mossambica. Litteratura Serpentium 5(5): 203.

Lilley, J.G. 1986. Captive breeding of the Mozambique spitting cobra (Naja mossambica) at the Poole Serpentarium. Herptile 11(3): 112-114.

Marais, J.; Liebenberg, A.G. 1980. Hybridization between the black-necked spitting cobra and the western barred spitting cobra. Journal of the Herpetological Association of Africa 24(1): 22-25.

Mavromichalis, J.; Bloem, S. 1995. A successful breeding attempt with Naja mossambica. Litteratura Serpentium (English Edition) 15(6): 138-141.

Pauwels, O.S.G.; David, P. 2000. Naja mossambica (Peters, 1854). Mozambique Spitting Cobra. African Herp News 31: 5-6.

Schultz, D.; Massyn, D. 2008. Naja mossambica Peters, 1854. Mozambique Spitting Cobra. African Herp News 46: 20-21.

Wyk, J.H. van; Haacke, W.; Macfadyen, D. 2001. Naja mossambica. Mozambique Spitting Cobra or M'fezi. African Herp News 33: 15.

Young, B.A.; O'Shea, M. 2005. Analyses of venom spitting in African cobras (Elapidae: Serpentes). African Zoology 40(1): 71-76.

Naja multifasciata

Bouillon, A. 1965. Serpentes danger! Zooleo 2: 1-76.

Laurent, R.F. 1956. Notes herpétologiques africaines. Revue de Zoologie et Botanique Africaine (Bruxelles) 53(3-4): 229-256.

Witte, G.F. de 1933. Contribution a la faune herpetologique du Congo Belge a propos du type de l'Elapechis multifasciatus (Werner). Ann. Soc. Scient. Bruxelles 53B: 70-73.

Naja naja

Anonymous. 1957. Cobra wins battle with viper. Loris 7(5): 421.

Anonymous. 1967. A cobra for a playmate. Loris 11: 123-124.

Anonymous. 1967. The cobra and the Polanga. Loris 11: 124.

Anonymous. 1967. White cobra. Loris 11: 125-126.

Addoor, S.N.R. 1993. Additional markings above hood marking in Indian (binocellate) cobra, Naja naja naja. Journal of the Bombay Natural History Society 90(2): 298.

Agrawal, H.P. 1979. Food and feeding habits of Naja naja naja Linn. Journal of Animal Morphology and Physiology 26(1-2): 272-275.

Barker, P.E. 1936. The common cobra in the Dooars. Journal of the Darjeeling Natural History Society 11(2): 81.

Brown, S.B. 1992. Microhabitat relations of some snakes and lizards in Tamil Nadu, south India. Hamadryad 17: 35-38.

Campbell, J.A.; Quinn, H.R. 1975. Reproduction in a pair of asiatic cobras, Naja naja (Serpentes, Elapidae). Journal of Herpetology 9: 229-233

Cheke, A.S. 1973. Snakes at Chiang Mai University. Natural History Bulletin of the Siam Society 24(3-4): 469-471.

Chhangani, A.K. 2004. Death of a blue bull Boselaphus tragocamelus due to snakebite. Journal of the Bombay Natural History Society 101(1): 150.

Cochran, D.M. 1944. Dangerous reptiles (popular). Rep. Smithson. Instn. 1943: 275-325.

Cuylenburg, R. van 1961. Hood of a Ceylon cobra Naja naja (L) also known as the spectacled cobra. Science 133: 665.

Damm, M. 1996. Naja naja (Linnaeus). Sauria (Berlin) 18(3): 361-364.

Daniels, R.J.R. 2011. Snake abundance: the limits of occupancy-based estimates. Current Science (Bangalore) 101(2): 137-138.

Dastidar, D.G.; Mani, S.D. 2008. Translocation of snakes from the NTPC campus at Alappuzha, Kerala, to the Konni Reserve Forest in Kerala. Cobra (Chennai) 2(1): 13-21.

Datta, A.K. 2014. Some death incidents of Spectacled Cobra (Naja naja) at Jahangirnagar University Campus, Savar, Dhaka, Bangladesh. Zoos' Print Magazine 29(5): 20-21.

Deraniyagala, P.E.P. 1940. A new colour variety of cobra from Ceylon and South India. Ceylon Journal of Science (B) 21: 233-235.

Deraniyagala, P.E.P. 1945. Some new races of the Python, Chrysopelea, binocellate cobra and tith-polonga inhabiting Ceylon and India. Spolia Zeylanica 24(2): 103-112.

Digana, P.M.C.; Yapa, W.B.; Randeniya, P.V.; Ratnasooriya, W.D. 2000. Predators of Sri Lankan bats. Loris 22(4): 46-48.

Digana, P.M.C.; Yapa, W.B.; Ratnasooriya, W.D. 2005. Pup predation of Schreibers' long-fingered bat, Miniopterus schreibersii (Microchiroptera) by two species of reptiles in the Wavulpena Cave in Sri Lanka. Lyriocephalus 6(1-2): 307-310.

Dubey, B.; Meganathan, P.R.; Haque, I. 2009. Multiplex PCR assay for rapid identification of three endangered snake species of India. Conservation Genetics 10(6): 1861-1864.

Einfalt, P. 2012. Die Brillenschlange - Naja naja. Natur und Tier-Verlag, Münster. 62 pp.

Elliot, R.H. 1933. 'Hooded Death'. Blackwood's 1933: 197-203.

Erdelen, W. 1998. Trade in lizards and snakes in Indonesia - biogeography, ignorance, and sustainability -. Mertensiella 9: 69-83.

Fleming, R.L.; Fleming, R.L. 1973. Some snakes from Nepal. Journal of the Bombay Natural History Society 70(3): 426-437.

Flowers, H.H. 1963. Active immunization of a human being against cobra (Naja naja) venom. Nature (London) 200: 1017-1018.

Flowers, H.H. 1963. Active immunization of human against Naja naja venom. Rep. US Arm. Med. Res Lab. 594: 1-5.

Ganesh, S.R.; Arumugam, M. 2016. Species richness of montane herpetofauna of southern Eastern Ghats, India: a historical resume and a descriptive checklist. Russian Journal of Herpetology 23(1): 7-24.

Glenn, J.L. 1982. Life history notes: Naja naja naja (Indian Cobra). Coloration. Herpetological Review 13(3): 96.

Hafeez, M.A.; Naz, Y.; Asmatullah; Ali, S.S.; Tariq, S. 1999. Comparative morphology of the pineal complex of Lacertilia and Ophidia (Squamata, Reptilia): light and electron microscopic study. Pakistan Journal of Zoology 31(1): 45-63.

Jadhav, S.S.; Disale, D.S.; Mukherjee, S. 2007. Tall amputation in Indian spectacled cobra Naja naja - A case study. Zoos' Print Journal 22(12): 2950.

Jennison, G. 1931. Cobras bred at Belle Vue Zoological Gardens, Manchester. Proceedings of the Zoological Society of London 101(4): 1413.

Johnsingh, A.J.T. 1976. Peacocks and cobra. Journal of the Bombay Natural History Society 73(1): 214.

Joseph, P.; Mathew, J.P.; Thomas, V.C. 2004. Locomotory behaviour in Xenochrophis piscator (Schneider), Naja naja (Linn) and Eryx johnii (Russell) under captive conditions. Cobra (Chennai) 56: 5-9.

Joseph, P.; Mathew, J.P.; Thomas, V.C. 2007. Scale morphology, arrangement and micro-ornamentation in Xenochrophis piscator (Schneider), Naja naja (Linn), and Eryx johni (Russell). Zoos' Print Journal 22(12): 2909-2912.

Kandambi, D.; Karunarathna, S.; Surasinghe, T.; Madawala, M. 2018. Natural history notes: Naja naja (Indian Cobra). Juvenile coloration. Herpetological Review 49(2): 347-348.

Kauffeld, C.F. 1947. The Hooden Clan. Fauna (Philadelphia) 9: 82-87.

Kauffeld, C.F. 1963. More at home with cobras. Bulletin of the Philadelphia Herpetological Society 11: 13-18.

Khaire, N. 1977. Snake collecting in Matheran. Newsletter of the Madras Snake Park Trust 2(1): 10-11.

Leigh, C. 1926. Notes on snakes. Journal of the Bombay Natural History Society 31: 227-228.

Li, D. 1989. A survey of reptiles in Leigongshan area. pp. 269-275. In: Matsui, M.; Hikida, T. & Goris, R.C. (eds.). Current Herpetology in East Asia. Proceedings of the Second Japan-China Herpetological Symposium Kyoto, July 1988. Herpetological Society of Japan, Kyoto, Japan. 521 pp.

Lim, B.L. 1974. Snakes as natural predators of rats in an oil palm estate. Malayan Nature Journal 27(3-4): 114-117.

Lim, B.L.; Waters, J.P.J.; Omar, A.R. bin 1960. Observations on some captive snakes. Malayan Nature Journal 14: 181-187.

Lin, L. 1983. [Some flavicant variants of reptiles (Plate 1)]. (In Chinese). Acta Herpetologica Sinica 2(1): 77.

Miller, H. 1970. The cobra, India's "good snake". National Geographic Magazine 138: 392-409.

Miller, N.C.E.; Pagden, H.T. 1931. Insect remains in the gut of a cobra, Naia tripudians. Nature (London) 127(3210): 706.

Mirza, Z.A.; Ullalkar, K.K.; Desouza, G.Q. 2007. Note on the predation of flying fox (Pteropus giganteus) by spectacled cobra Naja naja (Linnaeus, 1758). Reptile RAP 8: 9-10.

Mukherjee, D.; Bhupathy, S. 2004. Uropeltis ellioti in the diet of Naja naja. Hamadryad 28(1-2): 109-110.

O'Connell, B.; Greenlee, R.; Bacon, J.; Smith, H.M.; Chiszar, D. 1985. Strike-induced chemosensory searching in elapid snakes (cobras, taipans, tiger snakes and death adders) at San Diego Zoo. Psychological Record 35(4): 431-436.

Pal, A.; Dey, S.; Roy, U.S. 2012. Seasonal diversity and abundance of herpetofauna in and around an industrial city of West Bengal, India. Journal of Applied Sciences in Environmental Sanitation 7(4): 281-286.

Petzold, H.G. 1968. Zur Fortpflanzungsbiologie asiatischer Kobras (Naja naja). Zoologische Garten (Leipzig) 36: 133-146.

Procter, J.B. 1924. Unrecorded characters seen in living snakes and description of a new tree-frog. Proceedings of the Zoological Society of London 1924: 1125-1129.

Purandare, K. 2002. Attempt by the crested serpent-eagle Spilornis cheela to seize the Indian cobra Naja naja. Journal of the Bombay Natural History Society 99(2): 299.

Quinn, H.R.; Hulsey, T.G. 1978. Growth of the cobra Naja naja (Serpentes, Elapidae). Herpetological Review 9(4): 138-139.

Rajadurai, A. 1967. Mongoose and the cobra. Loris 11: 122-123.

Rajendran, M.V. 1953. Fear and prejudice. Journal Bengal Nat. Hist. Soc. 26(1): 21-27.

Rameshwaran, M. 2014. Egg-eating behavior of a Spectacled Cobra Naja naja. Cobra (Chennai) 8(1): 31-32.

Ranjitha, J.; Shoba, S. 2013. Diurnal behavioural profile of captive Indian cobras Naja naja (Linnaeus, 1758) during the dry season. Cobra (Chennai) 7(2): 23-26.

Sharma, O.P.; Kour, G. 2005. Karyology of four Indian snakes. Caryologia 58(4): 380-386.

Silva, A. de 1990. First record of a cobra (Naja naja) with a monocellate marking from Sri Lanka. The Snake 22(2): 134-135.

Simon, E.S. 1944. The breeding habits of the cobra (Naia tripudians Merrem) and the green whipsnake (Dryophis mycterizans). Journal of the Bombay Natural History Society 44(3): 480-1.

Sinha, C.K. 1996. On a new species of Hepatozoon from the blood and organs of Naja naja (Linnaeus). Geobios New Reports 15(2): 73-76.

Smith, M.A. 1937. Breeding habits of the Indian cobra. Journal Siam Soc. (Nat. Hist. Suppl.) 11(1): 62-63.

Soud, R.; Sutradhar, S.; Roy, B.C.; Talukdar, S. 2006. Extended distribution of the spectacled cobra Naja naja (Linnaeus, 1758) (Family: Elapidae) in western Assam, India. Cobra (Chennai) 64: 13.

Staedeli, J.H. 1972. Indian cobra. Zoonooz 45(8): 11.

Stejneger, L.H. 1936. Types of the amphibian and reptilian genera proposed by Laurenti in 1768. Copeia 1936(3): 133-141.

Stemmler, O. 1967. Junge Brillenschlangen Naja naja naja (Linnaeus) 1758. Aqua Terra 1967: 97-100.

Sundersingh, P.P.H. 1960. Reptilian fauna of Pilani with a note on the food contents. Part I. Ophidia. Proceedings Rajasthan Acad. Sci. 7: 52-59.

Tare, T.G.; Renapurkar, D.M. 1991. Observations on cannibalism in cobra Naja naja in captivity. The Snake 23(1): 42-43.

Theobald, W.H. 1926. An interesting encounter with a cobra. Journal of the Bombay Natural History Society 31: 528-529.

Vanderhaege, M. 1972. Variationen in der Brillenzeichnung bei nachgezüchteten Jungtieren von Naja n. naja (Serpentes, Elapidae). Salamandra 8(2): 95-96.

Wall, F. 1907. Peculiar progression of a cobra (Naia tripudians). Journal of the Bombay Natural History Society 18: 209-210.

Wall, F. 1910. A variety of the common cobra (Naia tripudians) from Chitral. Journal of the Bombay Natural History Society 19: 1001-1002.

Wall, F. 1922. Hatching of Cobras (Naia tripudians) with remarks on oodont, genetalia etc. Acquisition of four more specimens of Brachyophium rhodogaster Wall. Journal of the Bombay Natural History Society 28: 553-557.

Whitaker, R. 1970. Cobra and monitor together in the same burrow. Journal of the Bombay Natural History Society 66(3): 364.

Whitaker, R.; Andrews, H.V. 1996. Snake capture and venom extraction in Tamil Nadu, India. Occasional Papers of the IUCN Species Survival Commission 12: 12-13, 40-46.

Wüster, W. 1998. The cobras of the genus Naja in India. Hamadryad 23(1): 15-32.

Zylwa, C.L. de 1933. A tic polonga (Vipera russelli) swallowed by a cobra (Naia naia). Ceylon Journal of Science (B) 18(1): 143.

Naja nigricincta

Barts, M. 2014. Das Porträt zum Titelbild: Naja nigricincta nigricincta Bogert, 1940. Zebraschlange, Namibia Speikobra, Western Barred Cobra, Zebra Cobra. Sauria (Berlin) 36(1): 1-2.

Boycott, R.C.; Haacke, W.D. 1979. Note on the type-locality, distribution and juvenile coloration of Naja nigricollis woodi (Serpentes: Elapidae) and an account of the colour-pattern variation in intergrade populations. Annals of the Cape Provincial Museums Natural History 13(2): 31-38.

Cunningham, P.L.; Shilongo, K.E. 2002. Naja nigricollis nigrlcincta [nigricincta]. Western Barred Spitting Cobra or Zebra Snake. African Herp News 35: 12-13.

Cunningham, P.L.; Waal, C. van der 2010. Natural history notes: Naja nigricincta nigricincta Bogert, 1940. Western Barred Spitting Cobra. Diet. African Herp News 51: 9-11.

Elzen, P. van den 1983. Zur herpetofauna des Brandberges, Südwest-Afrika. Bonner Zoologische Beiträge 34(1-3): 293-309.

Haagner, G.V.; Carpenter, G. 1992. Notes on the captive breeding of the black spitting cobra, Naja nigricollis woodi (Serpentes: Elapidae). Journal of the Herpetological Association of Africa 41: 22-24.

Hoffman, L. 1987. Notes on snakes from Gobabeb, Namib-Naukluft Park. Journal of the Herpetological Association of Africa 33(1): 39.

Loehr, V.J.T. 2005. Naja nigricollis woodi Pringle, 1955. Black Spitting Cobra. Scavenging. African Herp News 38: 27.

Marais, J.; Liebenberg, A.G. 1980. Hybridization between the black-necked spitting cobra and the western barred spitting cobra. Journal of the Herpetological Association of Africa 24(1): 22-25.

Maritz, B. 2013. Natural history notes: Bitis arietans arietans (Merrem, 1820). Predation. African Herp News 59: 21-28.

Pringle, J.A. 1955. A new subspecies of the spitting cobra, Naja nigricollis Reinhardt, from the Cape Province. Annals of the Natal Museum 13: 253-254.

Naja nigricollis

Barry, D.H. 1963. West African snakes (Naja nigricollis). Nigerian Field 28: 142.

Bavay, A. 1895. Le serpent cracheur de la cote occidentale d'Afrique. Bulletin de la Société Zoologique de France 20: 210-213.

Berthe, R.A.; Pury, S. de; Bleckmann, H.; Westhoff, G. 2009. Spitting cobras adjust their venom distribution to target distance. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 195(8): 753-757.

Berthe, R.A.; Westhoff, G.; Bleckmann, H. 2013. Potential targets aimed at by spitting cobras when deterring predators from attacking. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 199(5): 335-340.

Bogert, C.M. 1954. Snakes that spit their venom. Animal Kingdom 57: 66-70.

Broadley, D.G. 1974. A review of the cobras of the Naja nigricollis complex in southwestern Africa (Serpentes: Elapidae). Cimbebasia 2(14): 155-162.

Eniang, E.A.; Ebin, C.O.; Luiselli, L. 2002. On the composition of the snake fauna of Okwangwo division of Cross River National Park, a hilly forest-savanna transition zone in south-eastern Nigeria (Squamata: Serpentes). Herpetozoa 15(1-2): 79-82.

FitzSimons, V. 1956. Reptiles and Amphibians of the Wonderboom Nature Reserve. Fauna and Flora (Pretoria) 7: 89-106.

Frade, F.; Manacas, S. 1955. Serpentes do Ultramar portugues. Garcia de Orta Sér. Zool. (Lisboa) 3(4): 547-553.

Freyvogel, T.A. 1965. Der 'Speiakt' von Naja nigricollis (Speikobra). Revue Suisse de Zoologie 72: 593-594.

Freyvogel, T.A.; Honegger, C.G. 1965. Der 'Speiakt' von Naja nigricollis. Acta Tropica (Basel) 22: 289-302.

Harper, J.B. 1963. Captive snakes in Ghana. British Journal of Herpetology 3: 71-74.

Hayes, W.K.; Herbert, S.S.; Harrison, J.R.; Wiley, K.L. 2008. Spitting versus biting: Differential venom gland contraction regulates venom expenditure in the black-necked spitting cobra, Naja nigricollis nigricollis. Journal of Herpetology 42(3): 453-460.

Hoier, R. 1950. A travers plaines et volcans au Pare National Albert. Reptiles. pp. 119-127. Bruxelles. 174 pp.

Horton, C.W. 1949. On the mechanics of the African spitting cobras, Naja nigricollis and Haemachatus haemachatus. Texas Journal of Science 1: 98.

Hughes, D.F.; Scharsu, C.; Behangana, M.; Wilber, L. 2017. Geographic distribution: Naja nigricollis (Black-necked Spitting Cobra). Herpetological Review 48(2): 392.

Ionides, C.J.P. 1946. Notes on snakes. Journal of the East Africa Natural History Society 19(3-4): 136-137.

Ionides, C.J.P. 1952. Cobra feigning death. British Journal of Herpetology 1: 114-115.

Ionides, C.J.P. 1953. Notes on mamba (genus Dendroaspis) in the southern province of Tanganyika Territory: notes on black-necked "spitting" cobra (Naja nigricollis Reinhardt) in southeast Tanganyika Territory. African Wild Life 7: 303-314.

Kauffeld, C.F. 1963. More at home with cobras. Bulletin of the Philadelphia Herpetological Society 11: 13-18.

Koch, M.; Sachs, W.B. 1927. Über zwei giftspeichende Schlangen, Sepedon haemachates und Naia nigricollis. Zoologischer Anzeiger 70: 155-159.

Lambiris, A.J. 1967. Observations on the feeding behaviour of Naja haje and Naja nigricollis. Journal of the Herpetological Association of Africa 3: 36-37.

Laurent, R.F. 1955. Diagnoses preliminaires de quelques serpents venimeux. Revue de Zoologie et Botanique Africaine (Bruxelles) 51: 127-139.

Laurent, R.F. 1973. Le concept de sous-espèce et la variation géographique chez Naja melanoleuca et Naja nigricollis (Reptilia, Ophidia). Revue de Zoologie et Botanique Africaine (Bruxelles) 87(1): 198-204.

Lea, J.M.; Politano, E.; Luiselli, L. 2003. Changes in the herpetofauna of a fresh water river in Southern Nigeria, after 20 years of development. Russian Journal of Herpetology 10(3): 191-198.

Luiselli, L. 2001. The ghost of a recent invasion in the reduced feeding rates of spitting cobras during the dry season in a rainforest region of tropical Africa? Acta Oecologica 22(5-6): 311-314.

Luiselli, L. 2002. Life-history correlates of suboptimal adaptation to rainforest biota by spitting cobras, Naja nigricollis, in southern Nigeria: comparative evidences with sympatric Forest Cobras, Naja melanoleuca. Revue d'Ecologie la Terre et la Vie 57(2): 123-133.

Luiselli, L.; Angelici, F.M.; Akani, G.C. 2002. Comparative feeding strategies and dietary plasticity of the sympatric cobras Naja melanoleuca and Naja nigricollis in three diverging Afrotropical habitats. Canadian Journal of Zoology 80(1): 55-63.

Luiselli, L.; Bikikoro, L.; Chiyere Ude, L. 1997. Fish-eating by the cobras Naja melanoleuca and Naja nigricollis (Elapidae) in the Niger Delta (Port Harcourt, Rivers State, Nigeria). British Herpetological Society Bulletin 59: 13-14.

Maclaud, M. 1896. Notes sur le Naja cracheur et le serum antivenimeux du Dr. Calmette. Arch. Medecine Nav. et Colon. 1896: 392-398.

Meier, J.; Freyvogel, T.A. 1980. Comparative studies on venoms of the fer-de-lance (Bothrops atrox), carpet viper (Echis carinatus) and spitting cobra (Naja nigricollis) snakes at different ages. Toxicon 18(5-6): 661-662.

Menzies, J.I. 1963. West African snakes (Naja nigricollis and N. melanoleuca). Nigerian Field 28: 143.

Pauwels, O.S.G.; Carlino, P.; Chirio, L.; Meunier, Q.; Okouyi Okouyi, J.V.; Orbell, C.; Rousseaux, D.; Testa, O. 2017. Miscellanea Herpetologica Gabonica IX. Bulletin of the Chicago Herpetological Society 52(6): 97-102.

Petit, L. 1895. Sur le serpent cracheur. Bulletin de la Société Zoologique de France 20: 239.

Phillips, J.A.; Alberts, A.C. 1992. Naive ophiophagus [sic] lizards recognize and avoid venomous snakes using chemical cues. Journal of Chemical Ecology 18(10): 1775-1783.

Rasmussen, J.B.; Hughes, B. 1997. Description of some new snake species. 1. Th. Reinhardt. Vid. Sel. naturvid. og mathem. Afh. (1843) 10 Part: 233-279. Steenstrupia 22 [1996]: 13-39.

Rose, W. 1955. The way of a serpent on a rock. African Wild Life 9: 170.

Scortecci, G. 1955. Viaggio di esplorazione biologica sulle montagne della Migiurtinia compiuto con il contribute del consiglio Nazionale delle Ricerche. Bollettino dei Musei e degli Istituti Biologici dell'Universita di Genova 25 [1953-1955]: 43-105.

Stanley, W.T.; Gara, I.J.; Msuya, C.A. 2007. Naja n. nigricollis Reinhardt, 1934. Black-necked Spitting Cobra. Consumption of museum voucher specimens. African Herp News 43: 23-24.

Villiers, A. 1950. Contribution a l'étude de l'Air (Mission L. Chopard et A. Villiers). Reptiles. Ophidiens et cheloniens. Mem. Inst. Franc. Afr. Noire 10: 337-344.

Villiers, A. 1956. Le Parc National du Niokolo-Koba, Reptiles. Memoires de l'Institut Fondamental D'Afrique Noire 48: 143-162.

Westhoff, G.; Boetig, M.; Bleckmann, H.; Young, B.A. 2010. Target tracking during venom 'spitting' by cobras. Journal of Experimental Biology 213(11): 1797-1802.

Westhoff, G.; Tzschaetzsch, K.; Bleckmann, H. 2005. The spitting behavior of two species of spitting cobras. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 191(10): 873-881.

Wüster, W.; Broadley, D.G. 2007. Get an eyeful of this: a new species of giant spitting cobra from eastern and north-eastern Africa (Squamata: Serpentes: Elapidae: Naja). Zootaxa 1532: 51-68.

Young, B.A.; O'Shea, M. 2005. Analyses of venom spitting in African cobras (Elapidae: Serpentes). African Zoology 40(1): 71-76.

Naja nivea

Broadley, D.G.; Wüster, W. 2004. A review of the southern African 'non-spitting' cobras (Serpentes: Elapidae: Naja). African Journal of Herpetology 53(2): 101-122.

FitzSimons, V.; Brain, C.K. 1958. A short account of the reptiles of the Kalahari Gemsbok National Park. Koedoe 1: 99-104.

Goldberg, S.R. 2010. Natural history notes: Naja nivea Linnaeus, 1758. Cape Cobra. Reproduction. African Herp News 50: 28.

Layloo, I.; Smith, C.; Maritz, B. 2017. Diet and feeding in the Cape Cobra, Naja nivea. African Journal of Herpetology 66(2): 147-153.

Morgan, D.R.; Haagner, G.V. 1992. Husbandry and propagation of the cape cobra (Naja nivea) at the Manyeleti Reptile Centre. Journal of the Herpetological Association of Africa 40: 87-89.

Phelps, T. 2006. Naja nivea (Linnaeus, 1758). Cape Cobra. Scavenging. African Herp News 40: 24.

Phelps, T. 2007. Naja nivea (Linnaeus, 1758). Cape Cobra. Maximum size. African Herp News 42: 23.

Phelps, T. 2007. Observations of the Cape cobra, Naja nivea (Serpentes: Elapidae) in the DeHoop Nature Reserve, Western Cape Province, South Africa. Herpetological Bulletin 99: 29-35.

Siebert, S.; Siebert, P. 2003. Pale chanting goshawk following Cape cobra. Promerops 254: 19.

Smythe, R. 2003. Unusual prey item. Promerops 254: 19.

Strassmann, W. 2001. Die Kapkobra Naja nivea. Haltung und Vermehrung. Reptilia (D) 6(27): 74-76.

Trutnau, L. 1984. Erfahrungen bei der Pflege und Nachzucht von zwei afrikanischen Naja-Arten: der Schwarzweissen Hutschlange Naja melanoleuca und der Kapkobra Naja nivea. Herpetofauna (Weinstadt) 6(31): 15-26.

Vanderwalt, B. 2006. Interaction between Cape cobra & pale chanting goshawk. Promerops 265: 10.

Visser, J. 2012. Geographical distribution: Naja nivea Linaeus, 1758. Cape Cobra. Namibia. African Herp News 56: 32-33.

Witberg, M. 2007. Naja nivea (Linnaeus, 1758). Cape Cobra. Juvenile markings. African Herp News 42: 21-22.

Witberg, M. 2011. Natural history notes: Naja nivea (Linnaeus, 1758). Cape Cobra. Juvenile colouration. African Herp News 53: 46-47.

Naja nubiae

Watkins-Colwell, G.J.; Dornburg, A.; Hawlena, D.; Moore, J.A. 2010. The amphibians and reptiles of the 1962-1965 Yale University Prehistoric Expedition to Nubia. Bulletin Peabody Museum of Natural History Yale University 51(2): 179-197.

Wüster, W.; Broadley, D.G. 2003. A new species of spitting cobra (Naja) from north-eastern Africa (Serpentes: Elapidae). Journal of Zoology (London) 259(4): 345-359.

Naja oxiana

Akhtar, S.A.; Tiwari, J.K. 1991. Extension of range of the black cobra Naja naja oxiana. Journal of the Bombay Natural History Society 88(1): 123.

Allen, G.S.; Fortyn, K. 1992. The Central Asian cobra, Naja oxiana (Eichwald, 1831): maintenance in captivity and a report of a first captive breeding. Bulletin of the Chicago Herpetological Society 27(6): 129-131.

Chernikin, E.M. 1963. Cobras in western Turkmenia. Priroda (Moscow) 1963(5): 115-116.

Danov, R.A. 1984. [Some data on the biology of the central Asian cobra (Naja oxiana Eichw., 1831) in Turkmenistan]. (In Russian). Izvestiya Akademii Nauk Turkmenskoi SSR Seriya Biologicheskikh Nauk 1984(1): 69-72.

Kamelin, Y.; Lukin, Y. 1997. Breeding Naja oxiana in captivity. pp. 121-125. In: Shilo, R.A. (ed.). Research work done in zoos. 8. Collection of oral papers of the International Conference devoted to the 50th Anniversary of Novosibirsk Zoo July 21-28, 1997. Novosibirsk Zoo, Novosibirsk. 135 pp.

Makeyev, V.M. 1969. A contribution to the comparative ecology of the kafi (Vipera lebetina) and the cobra (Naja oxiana) in the southwest Kopet-Dag. Zoologicheskii Zhurnal 48: 1832-1837.

Makeyev, V.M. 1972. The activity and behaviour of Naja oxiana (Eichwald) in captivity. Ekologiya 1972(6): 89-91.

Makeyev, V.M.; Zemlyanova, N.A. 1968. On the breeding of the Middle-Asian cobra (Naja oxiana Eichw.). Zoologicheskii Zhurnal 47: 1872-1874.

Nasoori, A.; Taghipour, A.; Delavar, S.; Aminirissehei, A.; Moghaddam, S. 2014. Heart place and tail length evaluation in Naja oxiana, Macrovipera lebetina, and Montivipera latifii. Asian Pacific Journal of Tropical Medicine 7: S137-S142.

Strauch, A. 1868. Über Eichwald's Tomyris oxiana, eine Giftschlange aus der Familie der Elapiden. Bull. Ac. Sc. St. Petersb. l° 13: 81-94.

Sundersingh, P.P.H. 1960. Reptilian fauna of Pilani with a note on the food contents. Part I. Ophidia. Proceedings Rajasthan Acad. Sci. 7: 52-59.

Vashetko, E.V.; Khodzhaev, A.F.; Fayazova, S. 1985. [Some problems in the ecology of Naja oxiana and histological changes in its internal organs in captivity]. pp. 39-45. (In Russian?). In: Persianova, L.A. (ed.). [Ecology and preservation of rare and threatened vertebrates of Uzbekistan]. Fan, Tashkent. 51 pp.

Wüster, W. 1998. The cobras of the genus Naja in India. Hamadryad 23(1): 15-32.

Young, B.A. 2018. Male reproductive behaviour of Naja oxiana (Eichwald, 1831) in captivity, with a case of unilateral hemipenile prolapse. Herpetology Notes 11: 1061-1064

Naja pallida

Attmarsson, E. 2004. Naja pallida captive care and reproduction of the red spitting cobra. Reptilia (GB) 33: 72-76.

Berthe, R.A.; Pury, S. de; Bleckmann, H.; Westhoff, G. 2009. Spitting cobras adjust their venom distribution to target distance. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 195(8): 753-757.

Berthe, R.A.; Westhoff, G.; Bleckmann, H. 2013. Potential targets aimed at by spitting cobras when deterring predators from attacking. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 199(5): 335-340.

Chiszar, D.; Castro, C.; Murphy, J.B.; Smith, H.M. 1989. Discriminations between thermally different rodent carcasses by Bull Snakes (Pituophis melanoleucus) and cobras (Naja pallida and Aspidelaps scutatus). Bulletin of the Chicago Herpetological Society 24(10): 181-183.

Chiszar, D.; Radcliffe, C.W.; Byers, T.; Stoops, R. 1986. Prey capture behavior in nine species of venomous snakes. Psychological Record 36(4): 433-438.

Dathe, F. 1997. Pflege und Vermehrung der Roten Speikobra (Naja pallida Boulenger, 1896) im Tierpark Berlin-Friedrichsfelde. Milu 9(2): 144-150.

Kardong, K.V.; Kiene, T.L.; Johnson, E.K. 1997. Proximate factors affecting the predatory behavior of the red spitting cobra, Naja mossambica pallida. Journal of Herpetology 31(1): 66-71.

Radcliffe, C.W.; Estep, K.; Boyer, T.; Chiszar, D. 1986. Stimulus control of predatory behaviour in red spitting cobras (Naja mossambica pallida) and Prairie Rattlesnakes (Crotalus v. viridis). Animal Behaviour 34(3): 804-814.

Rasmussen, J.B. 1996. Afrikanske slanger. 9. Naja pallida [African snakes. 9. Naja pallida]. (In Danish, English summary). Nordisk Herpetologisk Forening 39(5): 117-120.

Westhoff, G.; Boetig, M.; Bleckmann, H.; Young, B.A. 2010. Target tracking during venom 'spitting' by cobras. Journal of Experimental Biology 213(11): 1797-1802.

Westhoff, G.; Tzschaetzsch, K.; Bleckmann, H. 2005. The spitting behavior of two species of spitting cobras. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 191(10): 873-881.

Young, B.A.; Boetig, M.; Westhoff, G. 2009. Spitting behaviour of hatchling red spitting cobras (Naja pallida). Herpetological Journal 19(4): 185-191.

Young, B.A.; O'Shea, M. 2005. Analyses of venom spitting in African cobras (Elapidae: Serpentes). African Zoology 40(1): 71-76.

Naja peroescobari

Ceríaco, L.M.P.; Marques, M.P.; Schmitz, A.; Bauer, A.M. 2017. The “Cobra-preta” of São Tomé Island, Gulf of Guinea, is a new species of Naja Laurenti, 1768 (Squamata: Elapidae). Zootaxa 4324(1): 121–141.

Wüster, W.; Chirio, L.; Trape, J.F.; Ineich, I.; Jackson, K.; Greenbaum, E.; Barron, C.; Kusamba, C.; Nagy, Z.T.; Storey, R.; Hall, C.; Wüster, C.E.; Barlow, A.; Broadley, D.G. 2018. Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455(1): 68–98.

Naja philippinensis

Lutz, M. 2006. Die Kobras des philippinischen Archipels. I. Die Philippinen-Kobra, Naja philippinensis Taylor, 1922. Sauria (Berlin) 28(3): 5-11.

Salafranca, E.S. 1967. Longevity of the Naja naja philippinensis under stress of venom extraction. Zoologica (New York) 52: 41-47.

Sy, E.Y.; Balete, D.S. 2017. Geographic Distribution: Naja philippinensis (Northern Philippine Cobra). Herpetological Review 48(3): 590.

Sy, E.Y.; Gerard, T. 2014. Geographic distribution: Naja philippinensis (Northern Philippine Cobra). Herpetological Review 45(4): 664-665.

Sy, E.Y.; Vargas, M.G. 2017. Geographic Distribution: Naja philippinensis (Northern Philippine Cobra). Herpetological Review 48(3): 590.

Wüster, W.; Thorpe, R.S. 1990. Systematics and biogeography of the Asiatic cobra (Naja naja) species complex in the Philippine Islands. pp. 333-344. In: Peters, G. & Hütterer, R. (eds.). Vertebrates in the Tropics. Proceedings of the International Symposium on Vertebrate Biogeography and Systematics in the Tropics, Bonn, June 5-8, 1989. Alexander Koenig Zoological Research Institute and Zoological Museum, Bonn. 424 pp.

Naja sagittifera

Wüster, W. 1998. The cobras of the genus Naja in India. Hamadryad 23(1): 15-32.

Naja samarensis

Froemberg, C. 2010. Haltung und Nachzucht der Samar-Kobra (Naja samarensis). Reptilia (D) 15(81): 64-73.

Gressitt, J.L. 1937. Note on a Philippine cobra. Copeia 1937: 73.

Wüster, W.; Thorpe, R.S. 1990. Systematics and biogeography of the Asiatic cobra (Naja naja) species complex in the Philippine Islands. pp. 333-344. In: Peters, G. & Hütterer, R. (eds.). Vertebrates in the Tropics. Proceedings of the International Symposium on Vertebrate Biogeography and Systematics in the Tropics, Bonn, June 5-8, 1989. Alexander Koenig Zoological Research Institute and Zoological Museum, Bonn. 424 pp.

Naja savannula

Wüster, W.; Chirio, L.; Trape, J.F.; Ineich, I.; Jackson, K.; Greenbaum, E.; Barron, C.; Kusamba, C.; Nagy, Z.T.; Storey, R.; Hall, C.; Wüster, C.E.; Barlow, A.; Broadley, D.G. 2018. Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455(1): 68–98.

Naja senegalensis

Trape, J.F.; Baldé, C. 2014. A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa 3900(3): 301-338.

Trape, J.F.; Chirio, L.; Broadley, D.G.; Wüster, W. 2009. Phylogeography and systematic revision of the Egyptian cobra (Serpentes: Elapidae: Naja haje) species complex, with the description of a new species from West Africa. Zootaxa 2236: 1-25.

Naja siamensis

Berthe, R.A.; Westhoff, G.; Bleckmann, H. 2013. Potential targets aimed at by spitting cobras when deterring predators from attacking. Journal of Comparative Physiology A Sensory Neural and Behavioral Physiology 199(5): 335-340.

Chan-ard, T.; Stuart, B.L.; Wüster, W. 2001. First record of Indochinese spitting cobra Naja siamensis Laurenti (Serpentes: Elapidae) from Laos, with comments on the genus in the country. Natural History Bulletin of the Siam Society 48(1): 149-152.

Gaur, A.; Singh, C.S.; Sreenivas, A.; Singh, L. 2012. DNA-based identification of a snake in a wine bottle using universal primers: a case of mistaken identity. Forensic Science International 214(1-3): e51-e53.

Orlov, N.L.; Ryabov, S.A.; Nguyen, V.S.; Nguyen, Q.T. 2003. New records and data on the poorly known snakes of Vietnam. Russian Journal of Herpetology 10(3): 217-240.

Teynie, A.; David, P. 2007. Additions to the snake fauna of southern Laos, with the second Laotian specimen of Naja siamensis (Laurenti, 1768) and the first country record of Oligodon taeniatus (Gunther, 1861) (Squamata, Serpentes). Russian Journal of Herpetology 14(1): 39-44.

Westhoff, G.; Boetig, M.; Bleckmann, H.; Young, B.A. 2010. Target tracking during venom 'spitting' by cobras. Journal of Experimental Biology 213(11): 1797-1802.

Wüster, W.; Thorpe, R.S. 1994. Naja siamensis, a cryptic species of venomous snake revealed by mtDNA sequencing. Experientia (Basel) 50(1): 75-79.

Wüster, W.; Warrell, D.A.; Cox, M.J.; Jintakune, P.; Nabhitabhata, J. 1997. Redescription of Naja siamensis (Serpentes: Elapidae), a widely overlooked spitting cobra from S.E. Asia: geographic variation, medical importance and designation of a neotype. Journal of Zoology (London) 243(4): 771-788.

Naja sputatrix

Boeadi; Shine, R.; Sugardijto, J.; Amir, M.; Sinaga, M.H. 1998. Biology of the commercially-harvested Rat Snake (Ptyas mucosus) and cobra (Naja sputatrix) in central Java. Mertensiella 9: 99-104.

Fujimi, T.J.; Tsuchiya, T.; Tamiya, T. 2002. A comparative analysis of invaded sequences from group IA phospholipase A2 genes provides evidence about the divergence period of genes groups and snake families. Toxicon 40(7): 873-884.

Hamidy, A. 2007. Head scalation in Naja sputatrix Boie, 1827 - discussion on the presence of a small accessory scale. Hamadryad 31(2): 361-365.

Kopstein, F. 1930. Herpetologische Notizen. II. Oologische Beobachtungen an West-Javanischen Reptilien. Treubia 11: 301-307.

Sugardjito, J.; Boeadi; Amir, M.; Sinaga, M.H. 1998. Assessment of harvest levels and status of the spitting cobra (Naja sputatrix) and the Rat Snake (Ptyas mucosus) in central Java. Mertensiella 9: 105-110.

Wegner, A.M.R. 1953. Slangen. III. (Die slangen van de Indo-Australisches Archipel). De Tropische Natuur 23(1): 3-9.

Wüster, W. 1996. The status of the cobras of the genus Naja Laurenti, 1768 (Reptilia: Serpentes: Elapidae) on the island of Sulawesi. The Snake 27(2): 85-90.

Naja subfulva

Armitage, W.W. 1961. Some notes on a Naja melanoleuca at Durban Snake Park. Journal of the Herpetological Association of Rhodesia 16(1): 13-14.

Broadley, D.G. 1957. Occurrence of the forest cobra (Naja melanoleuca) in Rhodesia. Journal of the Herpetological Association of Rhodesia 1: 4-5.

Broadley, D.G. 1962. The herpetofauna of the Inyanga National Park. Journal of the Herpetological Association of Rhodesia 17/18: 8-10.

Broadley, D.G. 1968. An exceptional forest cobra from Rhodesia. Journal of the Herpetological Association of Africa 4(1): 30.

Bruno, S. 1967. Una notevole conferma per la fauna Somalia: Naja melanoleuca Hallowell 1858. Doriana 4(183): 1-9.

FitzSimons, V. 1947. Descriptions of new Species and Subspecies of reptiles and Amphibians from Natal, together with Notes on some other Little Known Species. Annals of the Natal Museum (Pietermaritzburg) 11(1): 111-137.

Flower, S.S. 1931. The occurrence of the "black-and-white cobra", Naja melanoleuca, in the Sudan. Annals and Magazine of Natural History (Ser. 10) 7: 499-500.

Hitchins, P.M.; Moll, E.J. 1967. A record of forest cobra, Naja melanoleuca, from S. of Tugeia river, Natal. Journal of the Herpetological Association of Africa 3: 13.

Laurent, R.F. 1955. Diagnoses preliminaires de quelques serpents venimeux. Revue de Zoologie et Botanique Africaine (Bruxelles) 51: 127-139.

Laurent, R.F. 1958. Notes herpetologiques africaines 2. Revue de Zoologie et Botanique Africaine (Bruxelles) 58: 115-128.

Marais, J.; Jubber, W. 2010. Geographical distribution: Naja melanoleuca Hallowell, 1857. Forest Cobra. South Africa: Limpopo. African Herp News 52: 24-25.

Newby, J.E. 1961. Snake catching at Lake Tanganyika. Nat. Soc. Cent. Afr. J. 2: 11-12.

Perret, J.L. 1961. Études herpétologiques Africaines. III. Bulletin Soc. Neuchâtel. Sci. Nat. (3) 84: 133-138.

Rose, W. 1962. Southern limit of Naja melanoleuca. Journal of the Herpetological Association of Rhodesia 17-18: 13.

Warren, B. 1960. A 4½ foot Naja melanoleuca in S. Rhodesia. Journal of the Herpetological Association of Rhodesia 11: 10.

Wüster, W.; Chirio, L.; Trape, J.F.; Ineich, I.; Jackson, K.; Greenbaum, E.; Barron, C.; Kusamba, C.; Nagy, Z.T.; Storey, R.; Hall, C.; Wüster, C.E.; Barlow, A.; Broadley, D.G. 2018. Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455(1): 68–98.

Naja sumatrana

Asad, S.; Siku, J.; Shabrani, A.; Wilting, A.; Rödel, M.O. 2018. Natural history notes: Naja sumatrana (Sumatran Spitting Cobra). Diet. Herpetological Review 49(1): 134-135.

Kopstein, F. 1935. Bemerkungen über Naja naja leucodira. Treubia 15: 151-155.

Leong, T.M.; Shunari, M. 2010. Attempted predation on a dog-toothed cat snake, Boiga cynodon by a black spitting cobra, Naja sumatrana in Singapore (Reptilia: Squamata). Nature in Singapore 3: 269-271.

Sy, E.Y.; Dycoco, Q.S.; Rodriguez, A.Z.A.; Du, S.G.; Wisco, M.A.S.; Pangilinan, C.M.M.; Banaguas, G.S. 2016. Geographic distribution: Naja sumatrana (Equatorial Spitting Cobra). Herpetological Review 47(3): 427-428.

Wüster, W.; Thorpe, R.S. 1990. Systematics and biogeography of the Asiatic cobra (Naja naja) species complex in the Philippine Islands. pp. 333-344. In: Peters, G. & Hütterer, R. (eds.). Vertebrates in the Tropics. Proceedings of the International Symposium on Vertebrate Biogeography and Systematics in the Tropics, Bonn, June 5-8, 1989. Alexander Koenig Zoological Research Institute and Zoological Museum, Bonn. 424 pp.