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© Rune Midtgaard


Taxonomy of the family Cordylidae
Bibliography of the genus Smaug
Biodiversity of the family Cordylidae








Genus
Smaug

Dragon Girdled Lizards, Dragon Lizards

Drachen-Gürtelschweife

Dragebæltehaler

2011 Smaug Stanley, Bauer, Jackman, Branch & Mouton (type species: Cordylus giganteus Smith 1844)
Contents: 9 species, of which 6 (66.7%) are endemic.
Distribution: Eswatini, Mozambique, South Africa (Free State, Gauteng, KwaZulu-Natal, Limpopo, Mpumalanga), Zimbabwe.

Smaug barbertonensis

Barberton Girdled Lizard, Barberton Dragon Lizard

Barberton-Gürtelschweif

Barberton-bæltehale

1921 Zonurus barbertonensis Van Dam
Cordylus warreni barbertonensis FitzSimons 1943
Smaug warreni barbertonensis Stanley, Bauer, Jackman, Branch & Mouton 2011
Smaug barbertonensis Stanley & Bates 2014

Remarks: Now restricted to Mpumalanga, South Africa. Previously included the population now assigned to swazicus (Bates & Stanley 2020).
Distribution: South Africa (Mpumalanga).


Smaug breyeri

Waterberg Girdled Lizard, Waterberg Dragon Lizard

Waterberg-Gürtelschweif

Waterberg-bæltehale

1921 Zonurus breyeri Van Dam
Cordylus warreni breyeri FitzSimons 1943
Cordylus breyeri Branch 1998
Smaug breyeri Stanley, Bauer, Jackman, Branch & Mouton 2011

Distribution: South Africa (Limpopo).


Smaug depressus

Soutpansberg Girdled Lizard, Soutpansberg Dragon Lizard, Flat Girdled Lizard

Soutpansberg-Gürtelschweif

Soutpansberg-bæltehale

1930 Zonurus barbertonensis depressus Fitzsimons
Cordylus warreni depressus Fitzsimons 1943
Smaug warreni depressus Stanley, Bauer, Jackman, Branch & Mouton 2011
Smaug depressus Stanley & Bates 2014

Distribution: South Africa (Limpopo).


Smaug giganteus

Giant Girdled Lizard, Sungazer

Riesen-Gürtelschweif

Kæmpebæltehale, Sydafrikansk Solkigger

1844 Cordylus giganteus Smith
Zonurus giganteus Boulenger 1885
Smaug giganteus Stanley, Bauer, Jackman, Branch & Mouton 2011
1845 Zonurus derbianus Gray (Boulenger 1885)

Remarks: Although reported to occur in KwaZulu-Natal (e.g., Branch 1998), it seems that no verified records exist for that province (Armstrong 2011).
Distribution: South Africa (Free State, Mpumalanga).


Smaug giganteus
© Rune Midtgaard

Smaug mossambicus

Gorongoza Girdled Lizard, Gorongoza Dragon Lizard

Gorongoza-Gürtelschweif

Gorongoza-bæltehale

1958 Cordylus warreni mossambicus Fitzsimons
Cordylus mossambicus Branch 1998
Smaug mossambicus Stanley, Bauer, Jackman, Branch & Mouton 2011

Introduced to: USA (Florida).
Distribution: Mozambique, Zimbabwe.


Smaug mossambicus
© Rune Midtgaard

Smaug regius

Regal Girdled Lizard, Regal Dragon Lizard

König-Gürtelschweif

Kongebæltehale

1962 Cordylus warreni regius Broadley
Cordylus regius Branch 1998
Smaug regius Stanley, Bauer, Jackman, Branch & Mouton 2011

Distribution: Zimbabwe.


Smaug swazicus

Swazi Girdled Lizard, Swazi Dragon Lizard

Swazi-Gürtelschweif

Swazi-bæltehale

2020 Smaug swazicus Bates & Stanley

Remarks: Previously regarded as a population of barbertonensis.
Distribution: Eswatini, South Africa (KwaZulu-Natal, Mpumalanga).





Smaug vandami

Gravelotte Girdled Lizard, Gravelotte Dragon Lizard, (Van Dam's Girdled Lizard)

Gravelotte-Gürtelschweif

Gravelotte-bæltehale

1930 Zonurus vandami Fitzsimons
Cordylus vandami FitzSimons 1943
Cordylus warreni vandami Loveridge 1944
Cordylus vandami Jacobsen 1989
Smaug vandami Stanley, Bauer, Jackman, Branch & Mouton 2011

Distribution: South Africa (Gauteng, Limpopo, Mpumalanga).


Smaug warreni

Lebombo Girdled Lizard, Lebombo Dragon Lizard, (Warren's Girdled Lizard)

Lebombo-Gürtelschweif, (Warrens Gürtelschweif)

Lebombo-bæltehale

1908 Zonurus warreni Boulenger
Cordylus warreni FitzSimons 1943
Smaug warreni Stanley, Bauer, Jackman, Branch & Mouton 2011
1930 Zonurus vandami perkoensis Fitzsimons (Branch 1988)
Cordylus vandami perkoensis Fitzsimons 1943
Cordylus warreni perkoensis Loveridge 1944
1933 Zonurus laevigatus Fitzsimons (Branch 1988)
Cordylus laevigatus Fitzsimons 1943
Cordylus warreni laevigatus Loveridge 1944

Remarks: Previously included barbertonensis, breyeri, depressus, mossambicus, regius, and vandami, as well as the population now assigned to swazicus.
Distribution: Eswatini, Mozambique, South Africa (KwaZulu-Natal, Mpumalanga).