Reptiles of Zimbabwe  (formerly Southern Rhodesia)   Bibliography of the reptiles of Zimbabwe Taxonomic history of the reptiles of Zimbabwe Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Chirindia (Pink Round-headed Worm Lizards)   Chirindia swynnertoni Chirinda Forest Round-headed Worm Lizard Branch 1998 Broadley & Howell 1991 Broadley 1988 Gans 2005 Spawls, Howell, Drewes & Ashe 2002 Genus Dalophia (Blunt-tailed Worm Lizards)   Dalophia longicauda Long-tailed Worm Lizard Broadley 1988 Broadley, Gans & Visser 1976 Gans 2005   Dalophia pistillum Zambezi Blunt-tailed Worm Lizard Broadley 1988 Broadley, Gans & Visser 1976 Genus Monopeltis (Spade-snouted Worm Lizards)   Monopeltis capensis Cape Spade-snouted Worm Lizard Broadley 1988   Monopeltis decosteri Delagoa Bay Worm Lizard Branch 1998 Broadley 1997 Broadley, Gans & Visser 1976 [as capensis "Group C"] Gans 2005   Monopeltis infuscata Dusky Spade-snouted Worm Lizard Branch 1998 Broadley 1997   Monopeltis leonhardi Kalahari Spade-snouted Worm Lizard Branch 1998 Broadley 1988 Broadley, Gans & Visser 1976   Monopeltis mauricei Botswana Spade-snouted Worm Lizard Broadley 1988   Monopeltis rhodesiana Zimbabwe Spade-snouted Worm Lizard Branch 1998 Broadley 1988 Broadley 1997 Gans 2005   Monopeltis zambezensis Zambezi Spade-snouted Worm Lizard Branch 1998 Broadley 1988 Broadley 1997 Broadley, Gans & Visser 1976 Gans 2005 Genus Zygaspis (Purple Round-headed Worm Lizards)   Zygaspis arenicola Maputo Round-headed Worm Lizard Branch 1998 Broadley & Broadley 1997   * Zygaspis ferox Ferocious Round-headed Worm Lizard Branch 1998 Broadley & Broadley 1997 Gans 2005   Zygaspis quadrifrons Kalahari Round-headed Worm Lizard Broadley & Broadley 1997 Broadley 1988 Gans 2005   Zygaspis violacea Violet Round-headed Worm Lizard Broadley 1988

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus niloticus Nile Crocodile Broadley 1988 King & Burke 1989 Wermuth & Mertens 1977

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthocercus (Ridgeback Agamas)   Acanthocercus atricollis Black-necked Ridgeback Agama Broadley 1988 Broadley 2008 Wagner, Greenbaum, Bauer, Kusamba & Leache 2018 Genus Agama (Typical Agamas)   Agama armata Zambezi Spiny Agama Branch 1998 Broadley 1988 Loveridge 1957   Agama kirkii Zambesi Rock Agama Branch 1998 Broadley 1988 Wermuth 1967   Agama mossambica Mozambique Agama Branch 1998 Broadley & Howell 1991 Broadley 1988  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo dilepis Flap-necked Chameleon Broadley 1988 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010 Genus Rhampholeon (African Leaf Chameleons)   Rhampholeon marshalli Chirinda Leaf Chameleon Branch 1998 Broadley 1988 Glaw 2015 Klaver & Boehme 1997 Tilbury 2010  Family Cordylidae (Girdled Lizards) Genus Chamaesaura (African Grass Lizards)   Chamaesaura macrolepis Large-scaled Grass Lizard Branch 1998 Broadley 1988 Genus Cordylus (Girdled Lizards)   Cordylus jonesii Limpopo Girdled Lizard Broadley & Branch 2002 Broadley 1988 Wermuth 1968   Cordylus rhodesianus Zimbabwe Girdled Lizard Branch 1998 Broadley & Branch 2002 Broadley 1988 Wermuth 1968   Cordylus tropidosternum Tropical Girdled Lizard Branch 1998 Broadley & Branch 2002 Broadley 1988 Wermuth 1968 Genus Platysaurus (Flat Lizards)   Platysaurus imperator Emperor Flat Lizard Branch 1998 Broadley 1978 Broadley 1988   Platysaurus intermedius Common Flat Lizard Branch 1998 Broadley 1978 Broadley 1988   Platysaurus ocellatus Ocellated Flat Lizard Branch 1998 Broadley 1978 Broadley 1988   Platysaurus pungweensis Pungwe Flat Lizard Branch 1998 Broadley 1978 Broadley 1988   Platysaurus torquatus Striped Flat Lizard Branch 1998 Broadley 1978 Broadley 1988 Genus Smaug (Dragon Girdled Lizards)   Smaug mossambicus Gorongoza Girdled Lizard Branch 1998 Broadley 1988   * Smaug regius Regal Girdled Lizard Branch 1998 Broadley 1988 Stanley & Bates 2014  Family Gekkonidae (Typical Geckos) Genus Afroedura (Flat Geckos)   Afroedura transvaalica Transvaal Flat Gecko Branch 1998 Branch, Guyton, Schmitz, Barej, Naskrecki, Farooq, Verburgt & Roedel 2017 Broadley 1988 Jacobsen, Kuhn, Jackman & Bauer 2014 Genus Chondrodactylus (Giant Ground Gecko and Thick-toed Geckos)   Chondrodactylus laevigatus Karas Thick-toed Gecko Heinz, Brennan, Jackman & Bauer 2021 Genus Elasmodactylus (Tropical Thick-toed Geckos)   Elasmodactylus tetensis Tete Thick-toed Gecko Broadley 1988 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus mabouia Cosmopolitan House Gecko Branch 1998 Broadley 1977 Broadley 1988   Hemidactylus platycephalus Flat-headed Tropical House Gecko Branch 1998 Broadley & Howell 1991 Broadley 1977 Broadley 1988   Hemidactylus tasmani Zimbabwean Half-toed Gecko Branch 1998 Broadley 1977 Broadley 1988 Wermuth 1965 Genus Homopholis (African Velvet Geckos)   Homopholis arnoldi Mahalapye Velvet Gecko Branch 1998 [as wahlbergii] Broadley 1988 [as wahlbergii] Broadley, Jackman & Bauer 2014 Loveridge 1947 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus angolensis Angolan Dwarf Gecko Branch 1998 Broadley & Howell 1991 Broadley 1988 Wermuth 1965   * Lygodactylus bernardi Zimbabwe Dwarf Gecko Broadley 1988 Travers, Jackman & Bauer 2014 Wermuth 1965   Lygodactylus bradfieldi Namibian Dwarf Gecko Branch 1998 Broadley 1991 Broadley 1992   Lygodactylus capensis Cape Dwarf Gecko Broadley 1988   Lygodactylus chobiensis Chobe Dwarf Gecko Branch 1998 Broadley 1988 Travers, Jackman & Bauer 2014   * Lygodactylus stevensoni Khami Ruins Dwarf Gecko Broadley 1988 Broadley 1992 Travers, Jackman & Bauer 2014 Genus Pachydactylus (Typical Thick-toed Geckos)   Pachydactylus oshaughnessyi Lake Nyassa Thick-toed Gecko Bauer & Lamb 2002 Branch 1998 Broadley 1977 Broadley 1988   Pachydactylus punctatus Speckled Thick-toed Gecko Bauer & Branch 1995 Branch 1998 Broadley 1988   Pachydactylus tigrinus Tiger Thick-toed Gecko Bauer & Lamb 2002 Branch 1998 Broadley 1977 Broadley 1988 Wermuth 1965   Pachydactylus vansoni Soutpansberg Thick-toed Gecko Branch 1998 Broadley 1977 Broadley 1988   Pachydactylus wahlbergii Kalahari Ground Gecko Branch 1998 Broadley & Spawls 1991 Genus Ptenopus (African Barking Geckos)   Ptenopus garrulus Common Barking Gecko Branch 1998 Broadley 1992  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Broadley 1987 Broadley 1988 Wermuth 1968 Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus auritus Kalahari Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch 1998 Broadley 1988   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Broadley 1988   Gerrhosaurus intermedius Eastern Black-lined Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch 1998 [as nigrolineatus] Broadley 1988 [as nigrolineatus] Genus Matobosaurus (Giant Plated Lizards)   Matobosaurus validus Eastern Giant Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch 1998 Broadley 1988 Wermuth 1968  Family Lacertidae (Typical Lizards) Genus Heliobolus (Bushveld Lizards)   Heliobolus lugubris Southern African Bushveld Lizard Branch 1998 Broadley 1988 Szczerbak 1975 Genus Holaspis (Blue-tailed Gliding Lizards)   Holaspis laevis Eastern Blue-tailed Gliding Lizard Broadley & Wursten 2007 Genus Ichnotropis (Rough-scaled Lizards)   Ichnotropis capensis Southern Rough-scaled Lizard Branch 1998 Broadley 1988 Genus Latastia (African Long-tailed Lizards)   Latastia johnstonii Malawi Long-tailed Lizard Bischoff 1991 Broadley 1988 Loveridge 1957 Genus Meroles (Southwest African Desert Lizards)   Meroles squamulosus Rough-scaled Desert Lizard Branch 1998 Broadley 1988 Genus Nucras (Sandveld Lizards)   Nucras caesicaudata Blue-tailed Sandveld Lizard Branch 1998 Broadley 1972 Broadley 1988   Nucras holubi Southern Sandveld Lizard Branch 1998   Nucras intertexta Spotted Sandveld Lizard Broadley 1972 Broadley 1988   Nucras ornata Ornate Sandveld Lizard Branch 1998 Broadley 1972 Broadley 1988  Family Scincidae (Skinks) Genus Acontias (Greater Legless Skinks)   Acontias aurantiacus Golden Blind Legless Skink Branch 1998 Broadley 1968 Broadley 1988   * Acontias bicolor Zimbabwe Blind Legless Skink Branch 1998 Broadley 1968 Broadley 1988 Lamb, Biswas & Bauer 2010   Acontias occidentalis Western Black Legless Skink Branch 1998 Broadley & Greer 1969 Broadley 1988 Lamb, Biswas & Bauer 2010   Acontias plumbeus Giant Legless Skink Branch 1998 Broadley & Greer 1969 Broadley 1988 Zhao, Verdu-Ricoy, Mohlakoana, Jordaan, Conradie & Heideman 2019 Genus Mochlus (African Supple Skinks)   Mochlus sundevallii Common Savanna Supple Skink Broadley 1966 Broadley 1988 Genus Panaspis (African Snake-eyed Skinks)   Panaspis maculicollus Spotted-neck Snake-eyed Skink Jacobsen & Broadley 2000   Panaspis wahlbergii Savanna Snake-eyed Skink Broadley 1988 Jacobsen & Broadley 2000 Genus Proscelotes (African Slender Skinks)   * Proscelotes arnoldi Zimbabwe Slender Skink Branch 1998 Broadley 1988 Genus Scelotes (Dwarf Burrowing Skinks)   Scelotes limpopoensis Limpopo Dwarf Burrowing Skink Branch 1998 Broadley 1988 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis boulengeri Lindi Speckled-lipped Skink Branch 1998 Broadley 1974 Broadley 1988 Broadley 2000   Trachylepis capensis Cape Skink Branch 1998 Broadley 1988 Broadley 2000   Trachylepis damarana Damara Variable Skink Weinell & Bauer 2018   Trachylepis depressa Eastern Coastal Skink Branch 1998 Broadley 1988 Broadley 2000   Trachylepis lacertiformis Bronze Rock Skink Branch 1998 Broadley 1975 Broadley 1988 Broadley 2000   Trachylepis margaritifer African Rainbow Skink Branch 1998 Broadley & Bauer 1998 Broadley 1988 Broadley 2000   Trachylepis punctatissima Montane Speckled Skink Branch 1998 Broadley 1977 Broadley 1988 Broadley 2000   Trachylepis punctulata Eastern Variegated Skink Broadley 1975 Broadley 1988 Broadley 2000 Portik & Bauer 2012   Trachylepis striata Eastern Striped Skink Branch 1998 Broadley & Howell 1991 Broadley 1977 Broadley 1988 Broadley 2000   Trachylepis varia Savanna Variable Skink Broadley 1988 Weinell & Bauer 2018   Trachylepis wahlbergii Mopane Skink Branch 1998 Broadley 1977 Broadley 1988 Broadley 2000 Genus Typhlacontias (Western Burrowing Skinks)   Typhlacontias rohani Kalahari Burrowing Skink Branch 1998 Broadley 1988 Haacke 1997  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bayless 2002 Broadley 1988 Mertens 1963   Varanus niloticus Nile Monitor Bayless 2002 Broadley 1988

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Genus Dasypeltis (African Egg Eaters)   Dasypeltis medici East African Egg Eater Bates & Broadley 2018 Branch 1998 Broadley & Blaylock 2013 Broadley 1988   Dasypeltis scabra Common African Egg Eater Broadley & Blaylock 2013 Broadley 1988 Gans 1959 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa aulica Cross-barred Nocturnal Tree Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Genus Dispholidus (Boomslangs)   Dispholidus viridis Green Boomslang Broadley & Blaylock 2013 Genus Limnophis (Striped Swamp Snakes)   Limnophis bangweolicus Eastern Striped Swamp Snake Broadley & Blaylock 2013 Broadley 1988 Genus Meizodon (African Smooth Snakes)   Meizodon semiornatus Semi-ornate Smooth Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Schatti 1985 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1966 Broadley 1988   Natriciteres sylvatica Eastern Forest Marsh Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1966 Broadley 1988 Spawls, Howell, Drewes & Ashe 2002 Genus Philothamnus (African Green Snakes)   Philothamnus angolensis Angolan Green Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Loveridge 1957 [as P. i. irregularis]   Philothamnus hoplogaster Southeastern Green Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988   Philothamnus natalensis Eastern Natal Green Snake Broadley & Blaylock 2013 Broadley 1988   Philothamnus occidentalis Western Natal Green Snake Broadley & Blaylock 2013 Broadley 1988   Philothamnus ornatus Ornate Green Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988   Philothamnus semivariegatus Variegated Bush Snake Broadley & Blaylock 2013 Broadley 1988 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus semiannulatus Southern Tiger Snake Broadley & Blaylock 2013 Broadley 1988 Genus Thelotornis (African Vine Snakes)   Thelotornis capensis Southern Vine Snake Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2002 Broadley 1988 Broadley 2001 Spawls & Branch 1995   Thelotornis mossambicanus Eastern Vine Snake Broadley & Blaylock 2013 Broadley & Wallach 2002 Broadley 1988 Broadley 2001  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Aspidelaps (Shield-nosed Cobras)   Aspidelaps scutatus Shield-nosed Coral Snake Branch 1998 Broadley & Baldwin 2006 Broadley & Blaylock 2013 Broadley 1988 Wallach, Williams & Boundy 2014 Genus Dendroaspis (Mambas)   Dendroaspis intermedius Eastern Green Mamba Branch 1998 [as D. angusticeps] Broadley & Blaylock 2013 Broadley & Howell 1991 [as D. angusticeps] Broadley 1988 [as D. angusticeps] David & Ineich 1999 [as D. angusticeps] Spawls & Branch 1995 [as D. angusticeps]   Dendroaspis polylepis Black Mamba Broadley & Blaylock 2013 Broadley 1988 David & Ineich 1999 Haakansson & Madsen 1983 Spawls & Branch 1995 Genus Elapsoidea (African Garter Snakes)   Elapsoidea boulengeri Zambezi Garter Snake Broadley & Blaylock 2013 Broadley 1971 Broadley 1988 David & Ineich 1999 Spawls & Branch 1995   Elapsoidea guentherii Western Garter Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1988 David & Ineich 1999 Jakobsen 1997 Spawls & Branch 1995   Elapsoidea sundevallii Southern African Garter Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1988 David & Ineich 1999 Spawls & Branch 1995 Genus Hemachatus (Ring-necked Spitting Cobra)   Hemachatus haemachates Ring-necked Spitting Cobra Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Spawls & Branch 1995 Genus Naja (Typical Cobras)   Naja anchietae Southwest African Cobra Branch 1998 Broadley & Blaylock 2013 Broadley & Wuster 2004 Broadley 1968 Broadley 1988 Broadley 1995   Naja annulifera African Banded Cobra Broadley & Blaylock 2013 Broadley & Wuster 2004 Broadley 1968 Broadley 1988 Broadley 1995 Spawls & Branch 1995   Naja mossambica Mozambique Spitting Cobra Broadley & Blaylock 2013 Broadley 1988 David & Ineich 1999 Spawls & Branch 1995 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja subfulva Brown Forest Cobra Branch 1998 [as N. melanoleuca] Broadley & Blaylock 2013 Broadley & Howell 1991 [as N. melanoleuca] Broadley 1968 [as N. melanoleuca] Broadley 1988 [as N. melanoleuca] David & Ineich 1999 [as N. melanoleuca] Spawls & Branch 1995 [as N. melanoleuca] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas bicolor Bicoloured Quill-snouted Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1988   Amblyodipsas mechowii Elongate Quill-snouted Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1988   Amblyodipsas polylepis Common Purple-glossed Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1988 Wallach, Williams & Boundy 2014   Amblyodipsas sabiensis Sabi Quill-snouted Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1988   Amblyodipsas ventrimaculata Kalahari Purple-glossed Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1971 Broadley 1988 Genus Amplorhinus (Many-spotted Snake)   Amplorhinus multimaculatus Many-spotted Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus capensis Cape Centipede Eater Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Wallach, Williams & Boundy 2014   Aparallactus guentheri East African Black Centipede Eater Broadley & Blaylock 2013 Broadley & Howell 1991 Broadley 1988 Loveridge 1957 Wallach, Williams & Boundy 2014   Aparallactus lunulatus Reticulated Centipede Eater Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Loveridge 1957 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis bibronii Southern Stiletto Snake Broadley & Blaylock 2013 Broadley 1988 Broadley 1991 David & Ineich 1999 Spawls & Branch 1995 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon capensis Brown House Snake Broadley & Blaylock 2013 Broadley 1988 [as fuliginosus] Genus Chilorhinophis (Black-and-Yellow Burrowing Snakes)   Chilorhinophis gerardi Katanga Black-and-Yellow Burrowing Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Spawls, Howell, Drewes & Ashe 2002 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Hughes 2004 Genus Duberria (African Slug Eaters)   Duberria lutrix Common Slug Eater Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Genus Gracililima (Black File Snake)   Gracililima nyassae Black File Snake Broadley & Blaylock 2013 Broadley 1988 Loveridge 1957 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis nototaenia Southeastern Bark Snake Broadley & Blaylock 2013 Broadley & Hughes 2000 Broadley 1988 Genus Limaformosa (Venus File Snakes)   Limaformosa capensis Cape File Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Genus Lycodonomorphus (Southern African Water Snakes)   Lycodonomorphus mlanjensis Mulanje White-bellied Water Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1967 Broadley 1988   Lycodonomorphus obscuriventris Floodplain Water Snake Broadley & Blaylock 2013 Broadley 1988 Kyle, Alexander & Preez 2021   Lycodonomorphus rufulus Common Brown Water Snake Broadley & Blaylock 2013 Broadley 1967 Broadley 1988 Genus Lycophidion (African Wolf Snakes)   Lycophidion capense Cape Wolf Snake Branch 1976 Broadley & Blaylock 2013 Broadley 1988 Broadley 1996   Lycophidion nanum Dwarf Wolf Snake Broadley & Blaylock 2013 Broadley 1988 Broadley 1996   Lycophidion variegatum Variegated Wolf Snake Branch 1976 Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Broadley 1996 Genus Prosymna (Shovel-snout Snakes)   Prosymna angolensis Angolan Shovel-snout Branch 1998 Broadley & Blaylock 2013 Conradie, Keates, Baptista & Lobon-Rovira 2022   Prosymna bivittata Two-striped Shovel-snout Branch 1998 Broadley & Blaylock 2013 Broadley 1980 Broadley 1988   Prosymna lineata Lineolate Shovel-snout Branch 1998 Broadley & Blaylock 2013 Broadley 1980 Broadley 1988   Prosymna stuhlmannii East African Shovel-snout Branch 1998 Broadley & Blaylock 2013 Broadley 1980 Broadley 1988 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis angolensis Dwarf Sand Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1977 Broadley 1988 Broadley 2002   Psammophis brevirostris Short-snouted Grass Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Broadley 2002   Psammophis crucifer Cross-marked Grass Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1977 Broadley 1988 Broadley 2002   Psammophis jallae Zimbabwe Sand Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1977 Broadley 1988 Broadley 2002   Psammophis mossambicus East African Olive Grass Snake Broadley & Blaylock 2013 [as P. phillipsii] Broadley 1988 [as P. phillipsi] Broadley 2002 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis orientalis Eastern Stripe-bellied Sand Snake Broadley & Blaylock 2013 Broadley 1977 Broadley 1988 Broadley 2002 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis subtaeniatus Western Stripe-bellied Sand Snake Broadley & Blaylock 2013 Broadley 1988 Broadley 2002 Genus Psammophylax (Skaapstekers)   Psammophylax tritaeniatus Southern Striped Skaapsteker Branch 1998 Broadley & Blaylock 2013 Broadley 1976 Broadley 1988 Keates, Conradie, Greenbaum & Edwards 2019 Genus Pseudaspis (African Mole Snake)   Pseudaspis cana African Mole Snake Broadley & Blaylock 2013 Broadley 1988 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis rostratus Rufous Beaked Snake Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Chirio & Ineich 1991 Lanza 1990  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops incognitus Incognito Thread Snake Branch 1998 Broadley & Blaylock 2013 Broadley & Broadley 1999 Broadley & Wallach 1997 Broadley & Watson 1976 Broadley 1983 Broadley 1988 Hahn 1980   Leptotyphlops scutifrons Zambezi Thread Snake Broadley & Blaylock 2013 Broadley & Broadley 1999 Broadley & Watson 1976 Broadley 1988 Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis longicauda Southern African Long-tailed Thread Snake Branch 1998 Broadley & Blaylock 2013 Broadley & Broadley 1999 Broadley & Howell 1991 Broadley & Watson 1976 Broadley 1988 Hahn 1980 McDiarmid, Campbell & Toure 1999  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python natalensis Southern Rock Python Broadley & Blaylock 2013 Broadley 1984 Broadley 1988 Spawls & Branch 1995 [as sebae]  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops bibronii Southern African Blind Snake Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1988 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014   Afrotyphlops fornasinii East African Blind Snake Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1988 Hahn 1980 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014   Afrotyphlops mucruso Zambezi Giant Blind Snake Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1988   Afrotyphlops obtusus Southern Gracile Blind Snake Broadley & Blaylock 2013 Broadley & Wallach 2007 Broadley 1988 McDiarmid, Campbell & Toure 1999 Genus Letheobia (African Gracile Blind Snakes)   Letheobia gracilis Urungu Gracile Blind Snake McDiarmid, Campbell & Toure 1999 Genus Rhinotyphlops (African Beaked Blind Snakes)   Rhinotyphlops lalandei Cape Blind Snake Branch 1998 Broadley & Blaylock 2013 Broadley & Wallach 2009 Broadley 1988 Hahn 1980 McDiarmid, Campbell & Toure 1999  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Branch 1998 Broadley & Blaylock 2013 Broadley 1988 David & Ineich 1999 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995   Bitis atropos Cape Mountain Adder Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Spawls & Branch 1995   Bitis caudalis Horned Adder Branch 1998 Broadley & Blaylock 2013 Broadley 1988 Spawls & Branch 1995   Bitis gabonica Gaboon Adder Branch 1998 Broadley & Blaylock 2013 Broadley & Howell 1991 Broadley 1988 Spawls & Branch 1995 Genus Causus (Night Adders)   Causus defilippii Snouted Night Adder Branch 1998 Broadley & Blaylock 2013 Broadley & Howell 1991 Broadley 1988 David & Ineich 1999 Spawls & Branch 1995   Causus rhombeatus Rhombic Night Adder Branch 1998 Broadley & Blaylock 2013 Broadley 1988 David & Ineich 1999 Spawls & Branch 1995

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa subrufa Common Helmeted Turtle Boycott & Bourquin 2008 Broadley 1988 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios bechuanicus Okavango Mud Turtle Broadley 1988 Fritz & Havas 2006 Iverson 1992 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios rhodesianus Zambian Mud Turtle Branch 1998 Broadley & Boycott 2008 Broadley 1988 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios sinuatus East African Serrated Mud Turtle Broadley & Boycott 2009 Broadley 1988 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelusios subniger East African Black Mud Turtle Branch 1998 Broadley 1988 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys spekii Eastern Savanna Hinge-back Tortoise Branch 1998 Broadley 1988 Hailey, Broadley, Boycott & Spawls 2021 Ihlow, Farooq, Gvozdik, Hofmeyr, Conradie, Campbell, Harvey, Verburgt & Fritz 2019 Kindler, Branch, Hofmeyr, Maran, Siroky, Vences, Harvey, Hauswaldt, Schleicher, Stuckas & Fritz 2012 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Kinixys zombensis Southeastern Hinge-back Tortoise Branch 1998 [as belliana] Broadley 1988 [as belliana] Fritz & Havas 2006 [as belliana] King & Burke 1989 [as belliana] McCord, Joseph-Ouni & Tabaka 2005 Genus Psammobates (South African Star Tortoises)   Psammobates oculiferus African Serrated Star Tortoise Broadley, Hunt & Cantle 2010 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Broadley 1988 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Trionychidae (Softshell Turtles) Genus Cycloderma (Central African Flapshell Turtles)   Cycloderma frenatum Zambezi Flapshell Turtle Branch 1998 Broadley & Sachsse 2011 Broadley 1988 Fritz & Havas 2006 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021