Reptiles of Free State [South Africa] Related checklists:  South Africa Bibliography of the reptiles of Free State Taxonomic history of the reptiles of Free State Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Monopeltis (Spade-snouted Worm Lizards)   Monopeltis capensis Cape Spade-snouted Worm Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1997 Broadley, Gans & Visser 1976 [as capensis "Group A"] Waal 1978

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama aculeata Kalahari Spiny Agama Bates 1992 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998   Agama atra Southern Rock Agama Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Wermuth 1967 Waal 1978   Agama hispida Southern Spiny Agama Bates 1992 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Cornelius, Marais & Cunningham 2012 McLachlan 1981 Waal 1978  Family Chamaeleonidae (Chameleons) Genus Bradypodion (Southern African Dwarf Chameleons)   Bradypodion dracomontanum Drakensberg Dwarf Chameleon Bates 1996 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Tilbury 2010 Genus Chamaeleo (Typical Chameleons)   Chamaeleo dilepis Flap-necked Chameleon Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Tilbury 2010 Waal 1978  Family Cordylidae (Girdled Lizards) Genus Chamaesaura (African Grass Lizards)   Chamaesaura aenea Coppery Grass Lizard Bates 1988 Bates 1991 Bates 1992 Bates 1996 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Genus Cordylus (Girdled Lizards)   Cordylus cordylus Cape Girdled Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Diedericks & Daniels 2014 Waal 1978   Cordylus vittifer Transvaal Girdled Lizard Bates & Broadley 2012 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978 Genus Karusasaurus (Karoo Girdled Lizards)   Karusasaurus polyzonus Southern Karoo Girdled Lizard Bates 1997 Branch 1998 Waal 1978 Genus Pseudocordylus (Eastern Crag Lizards)   Pseudocordylus langi Lesotho Crag Lizard Bates 2013 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Pseudocordylus melanotus Common Crag Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Waal 1978   Pseudocordylus microlepidotus Cape Crag Lizard Bates 1997   Pseudocordylus spinosus Spiny Crag Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Waal 1978   Pseudocordylus subviridis Drakensberg Crag Lizard Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978 Genus Smaug (Dragon Girdled Lizards)   Smaug giganteus Giant Girdled Lizard Armstrong 2011 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Waal 1978  Family Gekkonidae (Typical Geckos) Genus Afroedura (Flat Geckos)   Afroedura halli Eastern Cape Flat Gecko Bates 1992 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Jacobsen, Kuhn, Jackman & Bauer 2014 Waal 1978   Afroedura nivaria Drakensberg Flat Gecko Bates 1991 Bates 1992 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Jacobsen, Kuhn, Jackman & Bauer 2014 Waal 1978 Genus Chondrodactylus (Giant Ground Gecko and Thick-toed Geckos)   Chondrodactylus bibronii Tubercled Thick-toed Gecko Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Heinz, Brennan, Jackman & Bauer 2021 Waal 1978 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus capensis Cape Dwarf Gecko Rebelo, Bates, Burger, Branch & Conradie 2019 Genus Pachydactylus (Typical Thick-toed Geckos)   Pachydactylus capensis Common Cape Thick-toed Gecko Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Waal 1978   Pachydactylus mariquensis Marico Thick-toed Gecko Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Heinicke, Jackman & Branch 2011 Branch 1981 Branch 1998 Waal 1978   Pachydactylus oculatus Golden Spotted Thick-toed Gecko Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Waal 1978   Pachydactylus vansoni Soutpansberg Thick-toed Gecko Bates 1992 Bates 1996 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Genus Ptenopus (African Barking Geckos)   Ptenopus garrulus Common Barking Gecko Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978  Family Gerrhosauridae (Plated Lizards) Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Douglas 1992 Waal 1978 Genus Tetradactylus (Plated Snake Lizards)   Tetradactylus africanus African Plated Snake Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978   Tetradactylus breyeri Transvaal Plated Snake Lizard Bates 1996 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Pietersen 2020 Waal 1978   Tetradactylus tetradactylus Four-toed Plated Snake Lizard Bates 1988 Bates 1992 Bates 1996 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998  Family Lacertidae (Typical Lizards) Genus Meroles (Southwest African Desert Lizards)   Meroles squamulosus Rough-scaled Desert Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978 Genus Nucras (Sandveld Lizards)   Nucras holubi Southern Sandveld Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998   Nucras intertexta Spotted Sandveld Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1972   Nucras lalandii Southeastern Sandveld Lizard Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Waal 1978 Genus Pedioplanis (African Sand Lizards)   Pedioplanis burchelli Southern Sand Lizard Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Waal 1978   Pedioplanis lineoocellata Spotted Sand Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Waal 1978   Pedioplanis namaquensis Namaqua Sand Lizard Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978 Genus Tropidosaura (South African Mountain Lizards)   Tropidosaura cottrelli Drakensberg Mountain Lizard Bates 2013 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Tropidosaura essexi Lesotho Mountain Lizard Bates 1992 Bates 1996 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998  Family Scincidae (Skinks) Genus Acontias (Greater Legless Skinks)   Acontias gracilicauda Thin-tailed Legless Skink Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley & Greer 1969 Conradie, Busschau & Edwards 2018 Waal 1978 Genus Panaspis (African Snake-eyed Skinks)   Panaspis wahlbergii Savanna Snake-eyed Skink Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Jacobsen & Broadley 2000 Waal 1978 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis capensis Cape Skink Bates 1997 Waal 1978   Trachylepis homalocephala Red-sided Skink Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978   Trachylepis occidentalis Western Three-striped Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978   Trachylepis punctatissima Montane Speckled Skink Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1977 Waal 1978   Trachylepis punctulata Eastern Variegated Skink Bates 1988 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 2000 Waal 1978   Trachylepis spilogaster Namibian Tree Skink Bates 1997   Trachylepis sulcata Koppie Skink Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 2000 Waal 1978   Trachylepis varia Savanna Variable Skink Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1978   Trachylepis variegata Western Variegated Skink Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1975 Broadley 2000 Waal 1978  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bates 1990 Bates 1997 Waal 1978   Varanus niloticus Nile Monitor Bates 1992 Bates 1997 Bates 2010 Waal 1978

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Waal 1978 Genus Dasypeltis (African Egg Eaters)   Dasypeltis scabra Common African Egg Eater Bates 1991 Bates 1997 Broadley 1983 Waal 1978 Genus Dispholidus (Boomslangs)   Dispholidus viridis Green Boomslang Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as D. typus] Genus Philothamnus (African Green Snakes)   Philothamnus occidentalis Western Natal Green Snake Bates 1991 Bates 1992 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Waal 1978 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus beetzii Namib Tiger Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Waal 1978  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Aspidelaps (Shield-nosed Cobras)   Aspidelaps lubricus African Coral Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley & Baldwin 2006 Broadley 1983 Wallach, Williams & Boundy 2014 Waal 1978 Yeadon & Bates 2000 Genus Elapsoidea (African Garter Snakes)   Elapsoidea sundevallii Southern African Garter Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1971 Broadley 1983 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Waal 1978 Genus Hemachatus (Ring-necked Spitting Cobra)   Hemachatus haemachates Ring-necked Spitting Cobra Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Waal 1978 Genus Naja (Typical Cobras)   Naja nivea Yellow Cobra Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley & Wuster 2004 Broadley 1968 Broadley 1983 Waal 1978  Family Lamprophiidae (House Snakes, etc.) Genus Alopecion (Spotted House Snake)   Alopecion guttatum Spotted House Snake Broadley 1983 Waal 1978 Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas bicolor Bicoloured Quill-snouted Snake Bates 1988 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1971 Broadley 1983 Waal 1978 Genus Aparallactus (African Centipede Eaters)   Aparallactus capensis Cape Centipede Eater Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Wallach, Williams & Boundy 2014 Waal 1978 Genus Atractaspis (Stiletto Snakes)   Atractaspis bibronii Southern Stiletto Snake Bates 1988 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1991 Wallach, Williams & Boundy 2014 Waal 1978 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon capensis Brown House Snake Bates 1997 [as fuliginosus] Broadley 1983 [as fuliginosus] Waal 1978 [as f. fuliginosus] Genus Duberria (African Slug Eaters)   Duberria lutrix Common Slug Eater Bates 1991 Bates 1992 Bates 1997 Broadley 1983 Waal 1978 Genus Homoroselaps (African Harlequin Snakes)   Homoroselaps dorsalis Striped Harlequin Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Waal 1978   Homoroselaps lacteus Spotted Harlequin Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Waal 1978 Genus Lamprophis (SubSaharan House Snakes)   Lamprophis aurora Aurora House Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Waal 1978   Lamprophis fuscus Yellow-bellied House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Waal 1978 Genus Lycodonomorphus (Southern African Water Snakes)   Lycodonomorphus inornatus Olive-brown House Snake Bates 1991 Bates 1992 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Waal 1978   Lycodonomorphus laevissimus Dusky-bellied Water Snake Bates 1996 Bates 1997 Haagner & Branch 1994   Lycodonomorphus rufulus Common Brown Water Snake Bates 1988 Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Waal 1978 Genus Lycophidion (African Wolf Snakes)   Lycophidion capense Cape Wolf Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1976 Broadley 1983 Broadley 1996 Waal 1978 Genus Prosymna (Shovel-snout Snakes)   Prosymna bivittata Two-striped Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1980 Broadley 1983 Waal 1978   Prosymna sundevallii Southern African Shovel-snout Broadley 1980 Broadley 1983 Waal 1978 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis crucifer Cross-marked Grass Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Waal 1978   Psammophis notostictus Karoo Sand Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Waal 1978   Psammophis trinasalis Kalahari Sand Snake Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1977 Broadley 1983 Broadley 2002 Waal 1978 Genus Psammophylax (Skaapstekers)   Psammophylax rhombeatus Spotted Skaapsteker Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1976 Broadley 1983 Waal 1978   Psammophylax tritaeniatus Southern Striped Skaapsteker Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1976 Broadley 1983 Waal 1978 Genus Pseudaspis (African Mole Snake)   Pseudaspis cana African Mole Snake Bates 1991 Bates 1997 Broadley 1983 Waal 1978  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops conjunctus Cape Thread Snake Bates 1992 Bates 1996 Bates 1997 Broadley & Broadley 1999 Broadley & Wallach 1997 Broadley 1983 Waal 1978   Leptotyphlops scutifrons Zambezi Thread Snake Bates 1991 Bates 1992 Bates 1997 Broadley 1983 Waal 1978  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops bibronii Southern African Blind Snake Bates 1991 Bates 1997 Broadley & Wallach 2009 Broadley 1983 Hahn 1980 Wallach, Williams & Boundy 2014 Waal 1978 Genus Rhinotyphlops (African Beaked Blind Snakes)   Rhinotyphlops lalandei Cape Blind Snake Bates 1997 Broadley & Wallach 2009 Broadley 1983 Hahn 1980 Waal 1978  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Bates 1997 Branch 1998 Broadley 1983 Waal 1978   Bitis atropos Cape Mountain Adder Bates 1988 Bates 1991 Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 McDiarmid, Campbell & Toure 1999 Waal 1978 Genus Causus (Night Adders)   Causus rhombeatus Rhombic Night Adder Broadley 1983 Waal 1978

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa galeata South African Helmeted Turtle Bates 1997 [as subrufa] Branch 1998 [as subrufa] Iverson 1992 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Waal 1980 [as subrufa]  Family Testudinidae (Tortoises) Genus Homopus (Four-toed Dwarf Tortoises)   Homopus femoralis Greater Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Fritz & Havas 2006 Waal 1980 Genus Psammobates (South African Star Tortoises)   Psammobates oculiferus African Serrated Star Tortoise Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Fritz & Havas 2006 Waal 1980   Psammobates tentorius African Tent Tortoise Bates 1997 Wermuth & Mertens 1977 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Bates 1997 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Waal 1980