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Taxonomy of the genus Zamenis Biodiversity of the family Colubridae

Bibliography of the genus
Zamenis (Western Palearctic Rat Snakes)

(Reptilia: Serpentes: Colubridae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Zamenis in general

Chen, X.; Lemmon, A.R.; Lemmon, E.M.; Pyron, R.A.; Burbrink, F.T. 2017. Using phylogenomics to understand the link between biogeographic origins and regional diversification in ratsnakes. Molecular Phylogenetics and Evolution 111: 206-218.

Helfenberger, N.; Schulz, K.D. 2013. A checklist of the extinct taxa allied to the Old World ratsnakes, with remarks on the phylogeny and radiation of Elaphe, Rhinechis and Zamenis. pp. 423-427. In: Schulz, K.D. (ed.). Old World ratsnakes: a collection of papers. Bushmaster Publications, Berg. 432 pp.

Hoser, R.T. 2012. A taxonomic revision of the Colubrinae genera Zamenis and Orthriophis with the creation of two new genera (Serpentes: Colubridae). Australasian Journal of Herpetology 11: 59-64.

Utiger, U.; Helfenberger, N.; Schätti, B.; Schmidt, C.; Ruf, M.; Ziswiler, V. 2002. Molecular systematics and phylogeny of Old and New World ratsnakes, Elaphe auct., and related genera (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology 9(2): 105-124.

Zamenis hohenackeri

Afrasiab, S.R.; Mohamad, S.I. 2011. First record of the rat snake, Zamenis hohenackeri (Strauch, 1873), from north-eastern Iraq with notes on other colubrid snakes. Zoology in the Middle East 54: 19-22.

Akhmedov, S.B. 1989. [A new find of Elaphe hohenackeri hohenackeri in Azerbaidjan]. (In Russian). Vestnik Zoologii 1989(6): 84.

Baran, I. 1978. Some rare species of snakes from Turkey. Annalen des Naturhistorischen Museums in Wien 81 [1977]: 261-265.

Bischoff, W. 1993. Elaphe hohenackeri - Transkaukasische Kletternatter. pp. 317-329. In: Böhme, W. (ed.). Handbuch der Reptilien und Amphibien Europas. Band 3/I: Schlangen (Serpentes) 1 (Typhlopidae, Boidae, Colubridae 1: Colubrinae). Aula-Verlag, Wiesbaden. 480 pp.

Bosch, H.A.J. in den; Bischoff, W.; Schmidtler, J.F. 1998. Bemerkenswerter Reptilienfunde im Libanon. Herpetofauna (Weinstadt) 20(117): 19-32.

Bussmann, M.; Kostyra, S. 1996. Erganzende Mitteilungen zur Herpetofauna des Koycegiz-Beckens (Prov. Mugla, Türkei) mit Hinweis auf ein neues Vorkommen von Elaphe hohenackeri. Salamandra 32(1): 59-62.

Hingley, K.J. 1987. Snakes of the genus Elaphe. Their care and breeding in captivity. Part 3. Snake Keeper 1(3): 6-10.

Hofmann, S.; Mebert, K.; Schulz, K.D.; Helfenberger, N.; Göçmen, B.; Böhme, W. 2018. A new subspecies of Zamenis hohenackeri (Strauch, 1873) (Serpentes: Colubridae) based on morphological and molecular data. Zootaxa 4471(1): 137–153.

Hoofien, J.H. 1973. Contributions to the herpetofauna of Mount Hermon. 4. Elaphe hohenackeri (Ophidia: Colubridae). Israel Journal of Zoology 22(2-4): 67-73.

Hraoui-Bloquet, S.; Sadek, R.; Tok, C.V.; Ugurtas, I.H.; Sevinç, M.; Werner, Y.L.; Sterijovski, B.; Nilson, G.; Tuniyev, B.; Akarsu, F. 2009. Zamenis hohenackeri. The IUCN Red List of Threatened Species 2009: e.T157251A5060528. http://dx.doi.org/10.2305/IUCN.UK.2009.RLTS.T157251A5060528.en.

Jandzik, D.; Avci, A.; Gvozdik, V. 2013. Incongruence between taxonomy and genetics: three divergent lineages within two subspecies of the rare Transcaucasian rat snake (Zamenis hohenackeri). Amphibia-Reptilia 34(4): 579-584.

Nilson, G.; Andrén, C. 1984. A taxonomic account of the Iranian ratsnakes of the Elaphe longissima species-group. Amphibia-Reptilia 5(2): 157-171.

Schätti, B.; Baran, I. 1988. Bemerkungen zur Verbreitung von Elaphe hohenackeri (Strauch, 1873) und Vipera xanthina (Gray, 1849) in Süd-Anatolien (Serpentes: Colubridae, Viperidae). Salamandra 24(4): 306-309.

Schreiber, R. 2015. Haltung und Zucht der Transkaukasischen Kletternatter Zamenis hohenackeri (Strauch, 1873), sowie einige Anmerkungen zur Unterart Z. hohenackeri taurica (Werner, 1898). Sauria (Berlin) 37(3): 29-34.

Sikmashvili, N.M. 1975. [A study of the batracho and herpetofauna of the Little Caucasus (Meskhet-Dzhabakheti).] (In Russian). Vestnik Gosudarstvennogo Muzeya Gruzii (A) 28: 368-402.

Zamenis lineatus

Balli, A. 1944. Sulla presenza di Coluber longissimus var. lineata in provincia di Parma. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 33(2): 77-95.

Capula, M.; Luiselli, L.; Valenti, S.; Ceccarelli, A.; Rugiero, L.; Aloise, G. 2006. Are endemic snakes with a narrow distribution more specialist than their wide-ranging counterparts? Evidence from the prey composition and morphometric correlates of the diet in Zamenis lineatus, a rat snake endemic to southern Italy. Amphibia-Reptilia 27(4): 531-537.

Capula, M.; Luiselli, L.M. 2013. Natural history, morphology and ecology of the Italian Aesculapian snake, Zamenis lineatus. pp. 291-304. In: Schulz, K.D. (ed.). Old World ratsnakes: a collection of papers. Bushmaster Publications, Berg. 432 pp.

Corsetti, L.; Romano, A. 2008. On the occurrence of the Italian aesculapian snake, Zamenis lineatus (Camerano, 1891), in Latium (central Italy). Acta Herpetologica 3(2): 179-183.

Corti, C.; Sindaco, R.; Romano, A. 2009. Zamenis lineatus. The IUCN Red List of Threatened Species 2009: e.T61595A12505636. http://dx.doi.org/10.2305/IUCN.UK.2009.RLTS.T61595A12505636.en.

Esposito, C.; Romano, A. 2011. Extension of known range area of Zamenis lineatus (Camerano, 1891). New northern and western limits. Herpetozoa 23(3-4): 86-87.

Lenk, P.; Wüster, W. 1999. A multivariate approach to the systematics of Italian Rat Snakes of the Elaphe longissima complex (Reptilia, Colubridae): revalidation of Camerano's Callopeltis longissimus var. lineata. Herpetological Journal 9: 153-162.

Zamenis longissimus

Acolat, L.; Moreau, R. 1956. Tendance a l'extension en Franche-comte de la couleuvre d'esculape (Coluber aesculapii Gesn. = Elaphe longissima Laurenti). La Terre et la Vie (Paris) 103(2): 109.

Acolat, L.; Moreau, R. 1957. La couleuvre d'Esculape (Coluber aesculapii Gesn. = Elaphe longissima Laurenti) en Franche-Comte. Bull. Soc. Hist. nat. Doubs 59: 99-101.

Aghasyan, A.; Avci, A.; Tuniyev, B.; Crnobrnja-Isailovic, J.; Lymberakis, P.; Andrén, C.; Cogalniceanu, D.; Wilkinson, J.; Ananjeva, N.B.; Üzüm, N.; Orlov, N.L.; Podloucky, R.; Tuniyev, S.; Kaya, U.; Böhme, W.; Ajtic, R.; Vogrin, M. & al. 2017. Zamenis longissimus. The IUCN Red List of Threatened Species 2017: e.T157266A49063773. http://dx.doi.org/10.2305/IUCN.UK.2017-2.RLTS.T157266A49063773.en.

Assmann, O.; Lorenz, B. 2016. [Search and location of native snakes]. (In German). Oeko-L 38(1): 33-35.

Barowicz, T. 1979. [The snake Elaphe longissima]. (In Polish). Wszechswiat 1979(3): 55.

Bea, A.; Pascual, X.; Vilella, J.F.; Gonzalez, D.; Andreu, C. 1978. Notas sobre reptiles ibericos: 3. Estudio preliminar sobre biometria y distribucion de Elaphe longissima (Laur. 1768) en la peninsula Iberica. (Reptilia, Colubridae). Instituto Municipal de Ciencias Naturales Miscelanea Zoologica 4(2): 191-204.

Beshkov, V.A. 1975. Untersuchungen über die Biologie und die Ökologie der Schlangen im Maleschewska-Gebirge (Südwest-Bulgarien). l. Über die Fortpflanzung der Äskulapnatter (Elaphe longissima Laur.). Ekologiya (Sofia) 1: 75-83.

Beshkov, V.A. 1976. Über die Biologie and Ökologie der Schlangen im Makschevskigebirge (Südwest-Bulgarien). 2. Nahrung und Ernährung der Äskulapnatter (Elaphe longissima longissima (Laur.). Ekologiya (Sofia) 2: 34-42.

Beshkov, V.A.; Nankinov, D.N. 1979. Materialen über die Ornithophagia der Schlangen im Gebirge 'Maleschevska' (Südwestbulgarien). Ekologiya (Sofia) 5: 55-63.

Bezman-Moseyko, O.S. 2010. New records of three rare snake species from the left bank of Dniester River (Pridnestrovie). Russian Journal of Herpetology 17(4): 280-284.

Bielawski, R.; Marecek, J.; Vozenilek, P. 1992. Do snake eggs with hard shell exist? Fauna Bohemiae Septentrionalis 17: 75-76.

Boettger, O. 1909. Die Äskulapschlange (Coluber longissimus Laur.) im Böhmerwald und die Zornnatter (Zamenis gemonensis Laur.) im Böhmerwald, in den Kleinen Karpathen, Süd-Steiermark und Kärnten. Zool. Beob. (Frankfurt am Main) 50: 341-342.

Böhme, W. 1993. Elaphe longissima - Äskulapnatter. pp. 331-372. In: Böhme, W. (ed.). Handbuch der Reptilien und Amphibien Europas. Band 3/I: Schlangen (Serpentes) 1 (Typhlopidae, Boidae, Colubridae 1: Colubrinae). Aula-Verlag, Wiesbaden. 480 pp.

Bonnet, X.; Naulleau, G.; Shine, R. 1999. The dangers of leaving home: dispersal and mortality in snakes. Biological Conservation 89(1): 39-50.

Bonnet, X.; Shine, R.; Naulleau, G.; Vacher Vallas, M. 1998. Sexual dimorphism in snakes: different reproductive roles favour different body plans. Proceedings of the Royal Society of London B Biological Sciences 265(1392): 179-183.

Budziszewski, A.; Zemanek, M. 1978. Notes on the protection of the serpent Elaphe longissima. Chronmy Przyrode Ojczysta 34(2): 63-66.

Cais, L. 1963. [New stands and some remarks on the biology of the snake Elaphe longissima Laur]. (In Polish, English summary). Przeglad Zoologiczny 7: 301-302.

Capizzi, D. 1997. Effects of phylogenesis, morphology and habitat preferences on predatory exposure in a Mediterranean small mammal community. Italian Journal of Zoology (Modena) 64(1): 61-67.

Capizzi, D.; Capula, M.; Evangelisti, F.; Filippi, E.; Luiselli, L.; Trujillo-Jesus, V. 1996. Breeding frequency, clutch size, reproductive status and correlated behaviours in sympatric females Elaphe quatuorlineata and Elaphe longissima (Reptilia: Colubridae). Revue d'Ecologie la Terre et la Vie 51(4): 297-311.

Capizzi, D.; Capula, M.; Rugiero, L.; Luiselli, L. 2008. Dietary patterns of two sympatric Mediterranean snakes (Hierophis viridiflavus and Zamenis longissimus) along a gradient of habitat alteration. Herpetological Journal 18(3): 141-146.

Capizzi, D.; Luiselli, L. 1996. Feeding relationships and competitive interactions between phylogenetically unrelated predators (owls and snakes). Acta Oecologica 17(4): 265-284.

Capizzi, D.; Luiselli, L.; Capula, M.; Rugiero, L. 1995. Feeding habits of a Mediterranean community of snakes in relation to prey availability. Revue d'Ecologie la Terre et la Vie 50(4): 353-363.

Capocaccia, L. 1964. Variabilita e sottospecie di Elaphe longissima (Laur.) in Italia. (Serpentes). Ann. Mus. civ. Stor. nat. Giacomo Doria 74 [1963-64]: 353-387.

Capocaccia, L. 1965. Intorno a Elaphe longissima (Laur.) della Sardegna (serpentes). Doriana 4(161): 1-4.

Capula, M.; Luiselli, L. 2002. Feeding strategies of Elaphe longissima from contrasting Mediterranean habitats in central Italy. Italian Journal of Zoology (Modena) 69(2): 153-156.

Cardenete, L.G. 2006. Culebra de esculapio, Zamenis longissimus en los picos de Europa (Asturias). Nueva cita en su limite suroccidental de distribucion. Boletin de la Asociacion Herpetologica Espanola 16(1-2): 2-3.

Cattaneo, A. 1975. Presenza di Elaphe longissima longissima (Laurenti, 1768) melanica a Castelfusano (Roma) (Reptilia, Squamata, Colubridae). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 116(3-4): 251-262.

Cattaneo, A.; Grano, M.; Cattaneo, C. 2013. Episodes of predation on adult specimens of edible dormouse, Glis glis (Linnaeus, 1766), by Zamenis longissimus (Laurenti, 1768). pp. 305-308. In: Schulz, K.D. (ed.). Old World ratsnakes: a collection of papers. Bushmaster Publications, Berg. 432 pp.

Ciach, M. 2015. [The record of Aesculapian Snake Zamenis longissimus in the Ustrzyki Dolne town]. (In Polish). Chronmy Przyrode Ojczysta 71(3): 239-240.

Cimatti, E. 2003. Sacred snakes of the Mediterranean. Reptilia (GB) 26: 52-58.

Couto, S.; Ucha Fernandez, J.L. 2004. Dos neuvas citas de Elaphe longissima (Culebra de Esculapio) en el Pirineo Central. Boletin de la Asociacion Herpetologica Espanola 15(1): 12.

Covaciu-Marcov, S.D.; Sas, I.; Lazar, V.; Szeibel, N.; Condure, N. 2008. The herpetofauna in the plain area from the western Satu Mare County, Romania. Oltenai Studii si Comunicari Stiintele Naturii 24: 161-166.

Drobny, M. 1993. Aspekte der Populationsökologie und der Fortpflanzungsbiologie der Äskulapnatter, Elaphe longissima (Laurenti 1768) in Ostbayern. Mertensiella 3: 135-155.

Dupres, F. 1991. The aesculapian snake. Herptile 16(4): 144-145.

Dykyi, I.V. 2000. [A find of the Aesculapian snake (Elaphe longissima) on the territory of Lvov Region]. (In Ukrainian). Vestnik Zoologii 34(4-5): 128.

Elosegui, R.; Garcia, J.M. 1974. Nueva cita de Elaphe longissima para Espana. Doñana Acta Vertebrata 1(1): 60.

Esterbauer, H. 2014. [The Aesculapian snake Zamenis longissimus (Laurenti, 1768). First record of a amelanististic colour mutation in Austria]. (In German). Oeko-L 36(2): 7-11.

Ferri, V.; Bettiga, M. 1992. Un caso di albinismo nel colubro di esculapio, Elaphe l. longissima (Laurenti, 1768). Naturalista Valtellinese 3: 91-96.

Filippi, E.; Luiselli, L. 2001. Use of microhabitat and substratum types by sympatric snakes in a Mediterranean area of central Italy. Ecologia Mediterranea 27(1): 141-153.

Froesch, P. 1983. Zucht und Aufzucht der Äskulapnatter, Elaphe longissima. Aquarium (Minden) 17(170): 438-441.

Garancsy, M. 1975. Why do we protect all of our reptiles? Buvar 30(5): 217-221.

Gebhart, J. 2011. [Switzerland - a large field terrarium? Reptiles of the canton of Saint Gallen]. (In German). Elaphe 19(4): 30-35.

Gezova, S.; Drugac, P.; Purkart, A.; Jablonski, D. 2018. Albinism in two snake species recorded from Slovakia. Russian Journal of Herpetology 25(1): 79-82.

Ghezzi, D. 2010. Presenza di Zamenis longissimus (Laurenti, 1768) (Reptilia, Colubridae) nei pressi del fiume Po (Stagno Lombardo, CR). Pianura 25: 162-164.

Ghira, I.; Stugren, B. 1988. Verteilung von Amphibien und Reptilien auf Höhenzonen im Retezat-Gebirge. Studia Universitatis Babes-Bolyai Biologia 33(2): 69-75.

Ghisolfi, M. 2013. [New report of Zamenis longissimus (Laurenti, 1768) (Reptilia, Colubridae) in the floodplain of the river Po near Spinadesco (Cr)]. (In Italian). Pianura 30: 119-121.

Ghiurca, D.; Gherghel, I.; Rosu, G. 2009. Contribution to knowledge of the distribution of herpetofauna in Tarcau Mountains (Romania). AES Bioflux 1(2): 73-79.

Glowacinski, Z.; Witkowski, Z. 1969. [New localities of the aesculapian snake, Elaphe longissima longissima (Laun) 1768, in Bieszczady Mts]. (In Polish, English summary). Przeglad Zoologiczny 13: 208-211.

Golder, F. 1985. Ein gemeitnsamer Massen-Eiablageplatz von Natrix natrix helvetica (Lacepede, 1789) und Elaphe longissima longissima (Laurenti, 1768) mit Daten über Eizeitigung und Schlupf (Serpentes: Colubridae). Salamandra 21(1): 10-16.

Gomille, A. 2002. Die Äskulapnatter Elaphe longissima. Verbreitung und Lebensweise in Mitteleuropa. Edition Chimaira, Frankfurt am Main. 158 pp.

Grinchyshyn, T.Y.; Fedonjuk, O.V. 2007. [New record of the aesculapian snake, Elaphe longissima (Reptilia, Colubridae), in Lviv region (Ukraine)]. (In Ukrainian). Vestnik Zoologii 41(3): 226.

Grodzinski, W. 1957. [Materials for the study of the vertebrate fauna of the western Bieszczads]. (In Polish, English summary). Zesz. Nauk Uniw. Jagiellonsk. No. 10, Seria Nauk Biol. Zoologia 1: 177-221.

Guiller, G.; Legentilhomme, J.; Lourdais, O. 2014. Phénologie d’activité d’une communauté de squamates au nord de la Loire-Atlantique. Bulletin de la Societe Herpetologique de France 150: 25-39.

Hales, J. 1969. The Aesculapian snake. Ochrana Fauny 3: 119-123.

Heimes, P. 1987. Das Naturschutzzentrum Hessen 3. Das Reptilienschutzprogramm. Verhandlungen der Gesellschaft für Ökologie 16: 480-481.

Heimes, P. 1991. Zum Vorkommen der Äskulapnatter im Rheingau-Taunus. Natur und Museum (Frankfurt am Main) 121(6): 171-181.

Heimes, P. 1993. Die Verbreitung der Reptilien im westlichen Taunus. Jahrbücher des Nassauischen Vereins für Naturkunde 114: 41-57.

Heimes, P. 1994. Morphologische Anomalien bei Äskulapnattern (Elaphe longissima) im Rheingau-Taunus. Salamandra 30(4): 268-271.

Heimes, P. 1995. Der Turniertanz der Äskulapnatter. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 48(5): 296-299.

Heimes, P.; Waitzmann, M. 1993. Die Äskulapnatter (Elaphe longissima (Laurenti, 1768)) in Deutschland. (Reptilia, Serpentes: Colubridae). Zoologische Abhandlungen (Dresden) 47(2): 157-192.

Hingley, K.J. 1987. Snakes of the genus Elaphe. Their care and breeding in captivity. Part 3. Snake Keeper 1(3): 6-10.

Hofer, U.; Misslin, S.; Camponovo, I. 2002. Monitoraggio delle popolazioni di Saettone (Elaphe longissima), di Biacco (Hierophis viridiflavus) e di Natrice dal collare (Natrix natrix helvetica) in localita Boschi, Stabio TI. Bollettino della Societa Ticinese di Scienze Naturali 90(1-2): 59-67.

IIftime, A.; Iftime, O. 2014. Thanatosis and autohaemorrhaging in the Aesculapian Snake Zamenis longissimus (Laurenti, 1768). Herpetozoa 26(3-4): 173-174.

Jablonski, D.; Musilova, R.; Zavadil, V. 2011. [A find of the Aesculapian Snake (Zamenis longissimus) in South Bohemia]. (In Czech). Sbornik Jihoceskeho Muzea v Ceskych Budejovicich Prirodni Vedy 51: 166-169.

Jaeschke, J. 1971. Zur Einbürgerung der Äskulapnatter in Oberhessen. Salamandra 7: 85.

Joger, U.; Amann, T.; Lenk, P.; Willand, U. 1998. Molekuläre merkmale und das Phylogenetische Artkonzept. Zoologische Abhandlungen (Dresden) 50(Supplement): 109-123.

Joger, U.; Fritz, U.; Guicking, D.; Kalyabina-Hauf, S.; Nagy, Z.T.; Wink, M. 2010. Relict populations and endemic clades in Palearctic reptiles: evolutionary history and implications for conservation. pp. 119-143. In: Habel, J.C. & Assmann, T. (eds.). Relict species: phylogeography and conservation biology. Springer, Heidelberg, Dordrecht, etc. 449 pp.

Jozefik, M. 1960. [Further details about the occurrence of Elaphe longissima longissima Laur. in Bieszczady]. (In Polish, English summary). Przeglad Zoologiczny 4: 212-214.

Kammel, W. 2008. Aktivität und Nahrungserwerb der Äskulapnatter, Zamenis longissimus longissimus (Laurenti, 1768) in Österreich (Squamata: Serpentes: Colubridae). Herpetozoa 20(3-4): 117-143.

Kammel, W. 2009. Jahres- und Tagesrhythmen in der Aktivität und Beobachtungshäufigkeit dreier mittelEuropäischer Schlangenarten (Squamata: Serpentes: Colubridae). Herpetozoa 22(1-2): 3-9.

Kammel, W. 2009. Äussere Morphologie und Geschlechterverhältnis bei der Äskulapnatter, Zamenis longissimus longissimus (Laurenti, 1768) in Österreich. Herpetozoa 21(3-4): 99-121.

Kammerer, P. 1909. Coluber longissimus im Böhmerwald, Zamenis gemonensis im Böhmerwald, Wienerwald, den kleinen Karpathen, Süd-Steiermark und Kärnten. Zoologische Jahrbücher (Abteilung für Systematik, Geographie und Biologie der Tiere) (Jena) 27: 647-660.

Kanuch, P.; Balaz, P. 2005. Bat as a prey of Elaphe longissima (Laurenti, 1768). Herpetozoa 18(1-2): 92-93.

Kazmierczak, T. 1965. [Distribution of the Aesculapian snake, Elaphe longissima longissima (Laur.), in Poland]. (In Polish, English summary). Przeglad Zoologiczny 9: 380-385.

Kazmierczak, T. 1965. The Aesculapian snake in Poland. Chronmy Przyrode Ojczysta 21(4): 20-30, 57.

Keller, E. 1958. Ein guter Fang. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 11: 247-248.

Klemmer, K. 1985. Status und Schutzproblematik der Äskulapnatter (Elaphe longissima). Natur und Landschaft 60(9): 351-353.

Kobel, H.R. 1967. Morphometrische Karyotypanalyse einiger Schlangenarten. Genetica (Gravenhage) 38: 1-31.

Konig, D. 1985. Langjährige Beobachtungen an der Äskulapnatter Elaphe longissima (Laurenti, 1768) Serpentes: Colubridae. Salamandra 21(1): 17-39.

Kovar, R.; Brabec, M.; Vita, R.; Bocek, R. 2014. Mortality rate and activity patterns of an Aesculapian Snake (Zamenis longissimus) population divided by a busy road. Journal of Herpetology 48(1): 24-33.

Kovar, R.; Brabec, M.; Vita, R.; Vodicka, R.; Bogdan, V. 2017. Habitat use of the Aesculapian Snake, Zamenis longissimus, at the northern extreme of its range in northwest Bohemia. Herpetological Bulletin 136: 1-9.

Kovar, R.; Brabec, M.; Vita, R.; Vodicka, R.; Bogdan, V. 2017. Nesting and over-wintering sites of Aesculapian snake, Zamenis longissimus, in an anthropogenic landscape in the northern extreme of its range. Herpetological Bulletin 136: 35-36.

Krofel, M. 2004. First record of albino Aesculapian Snake (Elaphe longissima) in Slovenia. Natura Sloveniae 6(2): 53-56.

Kurek, K.; Bury, S.; Bas, G.; Najberek, K.; Kaczmarski, M.; Sniezko, S. 2014. [Telemetry studies of the Aesculapian snake in the Bieszczady Mountains - preliminary results and evaluation of the methods used]. (In Polish). Chronmy Przyrode Ojczysta 70(6): 510-522.

Kurek, P.; Skowron, B. 2010. [The abundance of Aesculapian snake Elaphe longissima from Zloty Potok (Krakowsko-Czestochowska Uppland)]. (In Polish). Przeglad Zoologiczny 52-54(1-4): 123-126.

Kwet, A. 2007. Zamenis longissimus (Laurenti, 1768) Äskulapnatter. Reptilia (D) 12(67): 51-54.

Lelievre, H.; Blouin-Demers, G.; Bonnet, X.; Lourdais, O. 2010. Thermal benefits of artificial shelters in snakes: a radiotelemetric study of two sympatric colubrids. Journal of Thermal Biology 35(7): 324-331.

Lelievre, H.; Blouin-Demers, G.; Pinaud, D.; Lisse, H.; Bonnet, X.; Lourdais, O. 2011. Contrasted thermal preferences translate into divergences in habitat use and realized performance in two sympatric snakes. Journal of Zoology (London) 284(4): 265-275.

Lelievre, H.; Henanff, M. le; Blouin-Demers, G.; Naulleau, G.; Lourdais, O. 2010. Thermal strategies and energetics in two sympatric colubrid snakes with contrasted exposure. Journal of Comparative Physiology B Biochemical Systemic and Environmental Physiology 180(3): 415-425.

Lelievre, H.; Legagneux, P.; Blouin-Demers, G.; Bonnet, X.; Lourdais, O. 2012. Trophic niche overlap in two syntopic colubrid snakes (Hierophis viridiflavus and Zamenis longissimus) with contrasted lifestyles. Amphibia-Reptilia 33(1): 37-44.

Lelievre, H.; Moreau, C.; Blouin-Demers, G.; Bonnet, X.; Lourdais, O. 2012. Two syntopic colubrid snakes differ in their energetic requirements and in their use of space. Herpetologica 68(3): 358-364.

Lenk, P.; Wüster, W. 1999. A multivariate approach to the systematics of Italian Rat Snakes of the Elaphe longissima complex (Reptilia, Colubridae): revalidation of Camerano's Callopeltis longissimus var. lineata. Herpetological Journal 9: 153-162.

Leo, F. de; Donelli, O.; Scali, S. 2006. Use of natural and artificial shelters in a snake community of northern Italy. pp. 25-31. In: Zuffi, M.A.L. (ed.). Societas Herpetologica Italica. Atti del V Congresso Nazionale: Calci (PI), 29 settembre - 3 ottobre 2004. Firenze University Press, Firenze. 270 pp.

Leptien, R. 2011. Tiere unserer Heimat: Die Äskulapnatter. Reptilia (D) 16(89): 79-82.

Letacq, A.L. 1897. La couleuvre d'Esculape et ses stations dans le Departement de l'Orne. Bull. Soc. Rouen 32: 132-133.

Lotze, H.U. 1975. Zum Paarungsverhalten der Äskulapnatter, Elaphe longissima. Salamandra 11(2): 67-76.

Luiselli, L.; Rugiero, L. 1993. Food habits of the Aesculapian snake, Elaphe longissima, in central Italy: do arboreal snakes eat more birds than terrestrial ones? Journal of Herpetology 27(1): 116-117.

Mahr, K.; Riegler, G.; Hoi, H. 2014. Parental risk management in relation to offspring defence: bad news for kids. Proceedings of the Royal Society Biological Sciences Series B 282(1798): 20141670.

Maluquer, J. 1918. Notas Herpetologicas. Madrid Bol. Soc. Esp. Hist. Nat. 18: 402-406.

Mangili, E. 1947. Osservazioni su la riproduzione in cattivita di alcune specie di ofidii Italiani. Acta pont.acad. Scient. 11(20): 235-246.

Maretic, Z.; Russell, F.E. 1979. An unusual nonvenomous snake bite. Toxicon 17(4): 425-427.

Mariétan, I. 1928. Notes sur quelques vertebres du Valais. Bull. Murith. Sion 45: 23-30.

Mariétan, I. 1958. Les reptiles en Valais. Bulletin Murith. 75: 91-102.

Martinot, J.P. 1986. Repartition et statut de protection des amphibiens et reptiles du Parc National de la Vanoise et de sa zone peripherique. Travaux Scientifiques du Parc National de la Vanoise 15: 255-269.

Meinig, H.; Schlupmann, M. 1987. Herpetologische Eindrücke einer Iberienreise. Herpetofauna (Weinstadt) 9(49): 11-24.

Mertens, R. 1948. Neues über Vorkommen der Äskulapnatter in Deutschland. Natur und Volk 78(4-6): 78-80.

Mertens, R. 1969. Bemerkungen zu, Neue Ergebnisse über das Vorkommen der Äskulapnatter (Elaphe longissima [Laurenti]) in der Tschechoslowakei. Zoologische Garten (Leipzig) 37: 108-110.

Merwald, F. 1965. Die Reptilien und Amphibien der Steyregger Auen. Naturkundliches Jahrbuch Stadt Linz 1965: 307-318.

Mlynarski, M. 1987. [Problems of the protection of amphibians and reptiles in Poland]. (In Polish, English summary). Chronmy Przyrode Ojczysta 43(3): 18-26.

Molle, F. 1961. Von der Äskulapnatter in Schlangenbad. Natur und Volk 91: 409-411.

Müller, K. 1952. Über eine Nachzucht von Elaphe longissima longissima (Laurenti). DATZ (Die Aquarien- und Terrarien-Zeitschrift) 5: 272-273.

Müller, K. 1952. Über eine Nachtzucht von Elaphe longissima longissima (Laurenti). DATZ (Die Aquarien- und Terrarien-Zeitschrift) 5: 272-273.

Müller, R. 1952. Der Äskulapstab. Eine neue sinnvolle Deutung des ärztlichen Zunftzeichens. Forschungen und Fortschritte 1942 25: 246.

Musilova, R.; Vavadil, V.; Kotlik, P. 2008. Isolated populations of Zamenis longissimus (Reptilia: Squamata) above the northern limit of the continuous range in Europe: origin and conservation status. Acta Societatis Zoologicae Bohemicae 71(3-4): 197-208.

Musilova, R.; Zavadil, V.; Markova, S.; Kotlik, P. 2010. Relics of the Europe's warm past: phylogeography of the Aesculapian snake. Molecular Phylogenetics and Evolution 57(3): 1245-1252.

Najbar, B. 1986. Does Aesculapian snake (Elaphe longissima longissima Laur. 1768) have to go extinct in Poland? Przeglad Zoologiczny 30(4): 425-429.

Najbar, B. 1999. [Breeding biology of the Aesculapian snake Elaphe longissima longissima (Laurenti) in the Bieszczady Zachodnie Mountains (SE Poland)]. (In Polish, English resume). Chronmy Przyrode Ojczysta 55(2): 5-20.

Najbar, B. 1999. Sklad pokarmu weza Eskulapa Elaphe longissima longissima (Laurenti) w Bieszczadach Zachodnich. [The diet of the Aesculapian snake Elaphe longissima longissima (Laurenti) in the Bieszczady Zachodnie Mountains (SE Poland)]. (In Polish, English summary). Chronmy Przyrode Ojczysta 55(2): 21-33.

Najbar, B. 2007. Food habits of Zamenis longissimus (Laurenti, 1768) (Reptilia: Serpentes: Colubridae) in Bieszczady (south-eastern Poland). Vertebrate Zoology 57(1): 73-77.

Naulleau, G. 1987. Use of biotelemetry in the study of free ranging snakes: example of Elaphe longissima. pp. 289-292. In: van Gelder, J.J., Strijbosch, H. & Bergers, P.J.M. (eds.). Proceedings of the 4th Ordinary General Meeting of the Societas Europaea Herpetologica. Societas Europaea Herpetologica, Faculty of Sciences, Nijmegen. 473 pp.

Naulleau, G. 1989. Elaphe longissima. pp. 154-155. In: Castanet, J. & Guyetant, R. Atlas de repartition des amphibiens et reptiles de France. Societe Herpetologique de France, Paris. 191 pp.

Naulleau, G. 1989. Etude biotelemetrique des deplacements et de la temperature chez la couleuvre d'esculape Elaphe longissima (Squamata, Colubridae) en zone forestiere. Bulletin de la Societe Herpetologique de France 52: 45-53.

Naulleau, G. 1992. Reproduction de la couleuvre d'esculape Elaphe longissima Laurenti (Reptilia, Colubridae) dans le centre ouest de la France. Bulletin de la Societe Herpetologique de France 62: 9-17.

Naulleau, G. 1994. Activite et temperature corporelle automnales et hivernales chez la couleuvre d'esculape Elaphe longissima (Squamata, Colubridae) dans le centre ouest de la France. Bulletin de la Societe Herpetologique de France 64 [1992]: 21-35.

Naulleau, G.; Bonnet, X. 1995. Reproductive ecology, body fat reserves and foraging mode in females of two contrasted snake species: Vipera aspis (terrestrial, viviparous) and Elaphe longissima (semi-arboreal, oviparous). Amphibia-Reptilia 16(1): 37-46.

Niebergall, P. 2003. Häutung einer adulten Äskulapnatter (Elaphe longissima) einschliesslich der Haut der Hemipenes. Elaphe 11(2): 62-63.

Nilson, G.; Andrén, C. 1984. A taxonomic account of the Iranian ratsnakes of the Elaphe longissima species-group. Amphibia-Reptilia 5(2): 157-171.

Olimpiev, N.K. 1973. An encounter between buzzard and Esculapian snake. Priroda (Moscow) 1973(2): 122.

Orth, K. 2013. Die Vierstreifennatter Elaphe quatuorlineata - Wissenswertes über die "europäische Riesenschlange" mit Berücksichtigung verwandter Arten Östliche Vierstreifennatter (Elaphe sauromates) und Äskulapnatter (Zamenis longissimus). Kleintierverlag Thorsten Geier, Biebertal. 67 pp.

Pilyavsky, B.R. 1983. A new finding of Aesculapian snake in the Ternopol region. Vestnik Zoologii 1983(2): 83-84.

Pizzuti Piccoli, A.; Cattaneo, A. 2006. Un episodio di insolita presa alimentare in Zamenis longissimus (Laurenti, 1768). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano 147(2): 340-342.

Popgeorgiev, G.S.; Tzankov, N.D.; Kornilev, Y.V.; Naumov, B.; Stojanov, A. 2010. Species diversity of amphibians and reptiles in the special protected area "Besaparski Ridove", southern Bulgaria. Biotechnology & Biotechnological Equipment 24(2) (Special Issue): 661-666.

Popgeorgiev, G.S.; Tzankov, N.D.; Kornilev, Y.V.; Naumov, B.Y.; Stoyanov, A.Y. 2014. Amphibians and reptiles in Ponor Special Protection Area (Natura 2000), western Bulgaria: species diversity, distribution and conservation. Acta Zoologica Bulgarica Suppl. 5: 85-96.

Pot, M. 1976. La couleuvre d'esculape, hôte inhabituel d'un nichoir. Nos Oiseaux 33(6): 282.

Puschnig, R. 1930. Die Verbreitung der Äskulapnatter (Coluber longissimus Lau.) in Kärnten. Carinthia II 119-120: 60-62.

Radovanovic, M. 1957. Einige Beobachtungen an Amphibien und Reptilien. Zoologischer Anzeiger 159: 130-137.

Ratnikov, V.Y. 2004. Identification of some eurasian species of Elaphe (Colubridae, Serpentes) on the basis of vertebrae. Russian Journal of Herpetology 11(2): 91-98.

Rausch, H. 1979. Zur Fortpflanzung der Äskulapnatter. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 32(6): 209-210.

Royans, D. 2009. Aesculapina ratsnake conservation. Herptile 34(2): 46-51.

Rugiero, L.; Capizzi, D.; Luiselli, L. 2002. Interactions between sympatric snakes, Coluber viridiflavus and Elaphe longissima: are there significant inter-annual differences in coexistence patterns? Ecologia Mediterranea 28(2): 75-91.

Rugiero, L.; Capula, M.; Capizzi, D.; Amori, G.; Milana, G.; Lai, M.; Luiselli, L.M. 2018. Long-term observations on the number of roadkilled Zamenis longissimus (Laurenti, 1768) in a hilly area of Central Italy. Herpetozoa 30(3-4): 212-217.

Rugiero, L.; Capula, M.; Capizzi, D.; Amori, G.; Milana, G.; Lai, M.; Luiselli, L.M. 2018. Long-term observations on the number of roadkilled Zamenis longissimus (Laurenti, 1768) in a hilly area of Central Italy. Herpetozoa 30(3-4): 212-217.

Rugiero, L.; Luiselli, L. 2004. Ecological notes on two colubrid snakes (Coluber viridiflavus and Elaphe longissima) in a surburban habitat (Rome, central Italy). Herpetological Bulletin 87: 8-12.

Rutschke, J. 1994. Haltung und Zucht der Äskulapnatter Elaphe longissima (Laurenti, 1768). Herpetofauna (Weinstadt) 16(89): 14-24.

Salvador, A. 1974. Die Äskulapnatter (Elaphe longissima) in Spanien. Salamandra 10(1): 42.

Sarauw, G.F.L. 1894. Haslingen, Æskulapslangen og Grønøglen. Naturen og Mennesk. (Copenhagen) 12: 258-274.

Scali, S.; Montonati, S. 2000. Analisi multivariata del dimorfismo sessuale in due specie di colubrini italiani (Coluber viridiflavus ed Elaphe longissima: Reptilia, Serpentes, Colubridae) su basi biometriche. pp. 429-434. In: Giacoma, C. (ed.). Atti I Congresso Nazionale della Societas Herpetologica Italica (Torino, 2-6 Ottobre 1996). [Proceedings of the first National Congress of the Societas Herpetologica Italica (Turin, 2-6 October 1996)]. Museo Regionale di Scienze Naturali, Torino. 821 pp.

Scaravelli, D. 2006. Caso di predazione di Elaphe longissima su Muscicapa striata nella Romagna meridionale. Quaderno di Studi e Notizie di Storia Naturale della Romagna 22: 141-142.

Schneppat, U.; Schmocker, H. 1983. Die Verbreitung der Reptilien im Kanton Graubünden. Ein Bericht über das Reptilienprojekt Graubünden für den Untersuchungszeitraum 1978-1982. Jahresbericht der Naturforschenden Gesellschaft Graubünden N.F. 100: 47-133.

Schweiger, M. 1994. Erstnachweis von Elaphe longissima (Laurenti, 1768) für die zentrale Osttürkei (Squamata: Serpentes: Colubridae). Herpetozoa 7(3-4): 149-151.

Seglie, D.; Sindaco, R. 2013. [Faunistic records in Piedmont and Aosta Valley (NW Italy), VI]. (In Italian). Rivista Piemontese di Storia Naturale 34: 439-452.

Semenzato, M.; Richard, J.; Amato, S. 1996. Boschi e risorgive planiziari: ambienti importanti per il mantenimento della continuita distributiva del popolamento erpetologico tra l'area montana e quella di pianura del Veneto. Studi Trentini di Scienze Naturali Acta Biologica 71: 33-40.

Skowron, B. 2003. Uwagi o wystepowaniu weza Eskulapa Elaphe longissima w Zlotym Potoku na Jurze Krakowsko-Czestochowskiej [Remarks on the occurrence of Elaphe longissima in Zloty Potok in the Krakow-Czestochowa Jura]. (In Polish). Chronmy Przyrode Ojczysta 59(3): 88-89.

Stevens, K. 1995. The European ratsnakes of the genus Elaphe. British Herpetological Society Bulletin 54: 10-20.

Strodicke, M.; Gerisch, B. 1999. Morphologische Merkmalsvariabilität bei Elaphe longissima (Laurenti, 1768), unter besonderer Berücksichtigung zweier isolierter Populationen an der Nordgrenze des Artareals (Squamata: Serpentes: Colubridae). Herpetozoa 11(3-4): 121-139.

Strugariu, A.; Gherghel, I. 2008. A preliminary report on the composition and distribution of the herpetofauna in the Lower Prut River Basin (Romania). North-Western Journal of Zoology 4 (Supplement 1): S49-S69.

Strugariu, A.; Sahlean, T.C.; Volosciuc-Hutuleac, M.V.; Puscasu, C.M. 2006. Preliminary data regarding the distribution of reptilian fauna in Suceava County (Romania). North-Western Journal of Zoology 2(1): 39-43.

Stugren, B.; Agadakos, N. 1987. Herpetologische Verhältnisse im Südosten von Peloponnes. Studia Universitatis Babes-Bolyai Biologia 32(2): 27-28.

Szabo, I. 1956. Contributions a l'herpeto-faune de la montagne Dunazug entre Szentendre, Visegrad et Esztergom. Allattani Kozlemanyck 45(3-4): 123-131.

Szyndlor, Z. 1984. Post and present distribution of Aesculapian snake Elaphe longissima (Laurenti) at the territory of Poland. Przeglad Zoologiczny 28(4): 513-523.

Taborski, A. 1959. [Remarks upon the Aesculapius snake, Elaphe longissima longissima Laur. 1768, and its distribution in Poland]. (In Polish, English summary). Przeglad Zoologiczny 3: 188-189.

Taborski, A.; Gruszka, S. 1964. [The colony of Elaphe longissima longissima Laur. 1768 in Bieszczady]. (In Polish, English summary). Przeglad Zoologiczny 8: 268-270.

Theou, P. 2016. Putative predation of Miniopterus schreibersii (Vespertilionidae, Chiroptera) by Zamenis longissimus (Colubridae, Reptilia) in the Albanian National Park of Prespa Lakes. Mammalia 80(5): 571-572.

Tiser, J. 1973. Herpetofauna ve zbytcich pevnosti v Komárne. Ochrana Fauny 6(1): 29-30.

Vaccaro, A.; Turrisi, G.F. 2007. Ritrovamento di Zamenis longissimus (Laurenti, 1768) (Reptilia, Colubridae) sull'Isola d'Elba (Toscana, Italia). Acta Herpetologica 2(1): 59-63.

Vlugt, R. van der 1994. Sacred snakes that heal. Litteratura Serpentium (English Edition) 14(4): 116.

Vogel, Z. 1968. Neue Ergebnisse über das Vorkommen der Äskulapnatter (Elaphe longissima) in der Tschechoslowakei. Zoologische Garten (Leipzig) 35: 166-178.

Vozenilek, P. 1973. Herpetologické poznámky z Vihorlatu. (In Slovak). Ochrana Fauny 5(1): 18-19.

Waitzmann, M. 1991. Zur Morphologie einiger Reptilien des südlichen Odenwaldes (Nordbaden, Südhessen). Salamandra 27(4): 266-281.

Waitzmann, M. 1993. Zur Situation der Äskulapnatter Elaphe longissima (Laurenti, 1768) in der Bundesrepublik Deutschland. Mertensiella 3: 115-133.

Waitzmann, M. 1997. Gefährdung und Schutzmöglichkeiten der Äskulapnatter in der Bundesrepublik Deutschland. Reptilia (D) 2(7): 25-28.

Werb, K. 1982. Brief notes on breeding the snake Elaphe longissima in captivity. Herptile 7(4): 12.

Wilson, M.; Stille, B.; Stille, M. 2014. Herpetofauna of Paxos, Ionian Islands, Greece, including two species new to the island. Herpetozoa 27(1-2): 108-112.

Wirth, M. 2009. Königinnen der Europäischen Schlangen: Vierstreifennater (Elaphe quatuorlineata), Äskulap-natter (Zamenis longissimus) und Leopardnatter (Zamenis situla). Draco 10(3) (39): 61-74.

Wutschert, R. 1984. Neues über die Reptilienfauna der Insel Korfu. Salamandra 20(4): 221-228.

Zagar, A.; Carretero, M.A.; Krofel, M.; Luznik, M.; Podnar, M.; Tvrtkovic, N. 2014. Reptile survey in Dinara Mountain (Croatia) revealed the southernmost known population of Horvath's Rock Lizard (Iberolacerta horvathi). Natura Croatica 23(1): 235-240.

Zagorodniuk, I.V. 2003. [Aesculapian snake (Elaphe longissima) on the northern macroslope of the Ukrainian Carpathians]. (In Ukrainian). Vestnik Zoologii 37(3): 22.

Zamenis persicus

Bennemann, R. 2007. Zamenis persicus (Werner 1913) in der Terrarienhaltung. Ophidia 1(1): 28-31.

Mamet, S.; Kudryavtsev, S.V. 1997. Notes on the reproductive biology of rare Asian ratsnake Elaphe persica. Russian Journal of Herpetology 4(2): 200-202.

Mamet, S.; Kudryavtsev, S.V. 1998. Notes on the reproductive biology of rare Asian ratsnake (Elaphe persica). [continuation]. Russian Journal of Herpetology 5(1): 17-19.

Nilson, G.; Andrén, C. 1984. A taxonomic account of the Iranian ratsnakes of the Elaphe longissima species-group. Amphibia-Reptilia 5(2): 157-171.

Schätti, B.; Tillack, F.; Schulz, K.D. 2010. History, status, and validity of the racer Zamenis persicus Jan, 1863, and the scientific name of the ratsnake Z. persicus (Werner, 1913) (Squamata, Colubridae). Zoosystematics and Evolution 86(2): 275-281.

Szczerbak, N.N.; Eremchenko, V.K. 1987. [New findings of reptiles on the territory of the USSR]. (In Russian). Vestnik Zoologii 1987(5): 85.

Tuniyev, B.; Ananjeva, N.B.; Aghasyan, A.; Orlov, N.L.; Tuniyev, S.; Anderson, S.C. 2009. Zamenis persica. The IUCN Red List of Threatened Species 2009: e.T164577A5909409. http://dx.doi.org/10.2305/IUCN.UK.2009.RLTS.T164577A5909409.en.

Werner, F. 1914. Neue oder seltene Reptilien und Frösche des Naturhistorischen Museums in Hamburg. Hamburg Jahrb. wiss. Anst. 30(2): 1-51.

Zamenis scalaris

Anonymous. 2005. Rhinechis scalaris ladder snake (Schinz, 1822). Reptilia (GB) 40: 43-46.

Anonymous. 2006. Scientific name: Rhinechis scalaris (formerly Elaphe scalaris) common name: the European laddersnake. Herptile 31(2): 63-69.

Alarcos, G.; Flechoso, F.; Rodriguez-Pereira, A.; Lizana, M. 2009. [Predation by the ladder snake Rhinechis scalaris on a litter of the common vole (Microtus arvalis)]. Boletin de la Asociacion Herpetologica Espanola 20: 61-62.

Amo, L.; Lopez, P.; Martin, J. 2004. Chemosensory recognition and behavioral responses of wall lizards, Podarcis muralis, to scents of snakes that pose different risks of predation. Copeia 2004(3): 691-696.

Belmonte, J. 1993. Estudio comparativo sobre la influencia del laboreo en las poblaciones de vertebrados en la campina de Jerez. Boletin de Sanidad Vegetal Plagas 19(2): 211-220.

Bonny, K. 1998. Elaphe scalaris (Schinz, 1822) - Haltung und Vermehrung der Treppennatter im Terrarium. Elaphe 6(4): 13-16.

Bueno-Enciso, J.; Ferrer, E.S.; Barrientos, R.; Serrano-Davies, E.; Sanz, J.J. 2016. Habitat fragmentation influences nestling growth in Mediterranean blue and great tits. Acta Oecologica 70: 129-137.

Butler, R. 1991. Notes on the keeping and breeding of the ladder snake, Elaphe scalaris. Herptile 16(1): 28-34.

Butler, R. 2004. Retention of juvenile markings into adulthood in a female ladder snake Rhinechis scalaris (=Elaphe scalaris). Herptile 29(4): 157-161.

Calmonte, T.; Ferri, V. 1987. Un serpent nuovo per la fauna italiana: il colubro scalare, Elaphe scalaris (Schinz, 1822) (Reptilia, Colubridae). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 128(3-4): 314-316.

Chen, X.; Lemmon, A.R.; Lemmon, E.M.; Pyron, R.A.; Burbrink, F.T. 2017. Using phylogenomics to understand the link between biogeographic origins and regional diversification in ratsnakes. Molecular Phylogenetics and Evolution 111: 206-218.

Cheylan, M. 1986. Mise en evidence d'une activite nocturne chez le serpent mediterraneen Elaphe scalaris (Ophidia, Colubridae). Amphibia-Reptilia 7(2): 181-186.

Cheylan, M.; Guillaume, C.P. 1993. Elaphe scalaris - Treppennatter. pp. 397-429. In: Böhme, W. (ed.). Handbuch der Reptilien und Amphibien Europas. Band 3/I: Schlangen (Serpentes) 1 (Typhlopidae, Boidae, Colubridae 1: Colubrinae). Aula-Verlag, Wiesbaden. 480 pp.

Coffinet, K. 1950. Coluber quaterradiatus a Bourron? Bull. Ass. Nat. Loing 26(10): 107-108.

Deas, J. 1981. More Spanish herptiles. Herptile 6(4): 25-27.

Dupres, F. 1991. The ladder snake, Elaphe scalaris. Herptile 16(4): 142-143.

Ewald, P. 1989. Elaphe scalaris. pp. 156-157. In: Castanet, J. & Guyetant, R. Atlas de repartition des amphibiens et reptiles de France. Societe Herpetologique de France, Paris. 191 pp.

Feriche, M.; Pleguezuelos, J.M.; Cerro, A. 1993. Sexual dimorphism and sexing of Mediterranean colubrids based on external characteristics. Journal of Herpetology 27(4): 357-362.

Font, E.; Roca, V.; Lluch, J.; Navarro, P.; Desfilis, E. 2002. Primer caso de somatodicotomia en una serpiente de escalera, Elaphe scalaris. Boletin de la Asociacion Herpetologica Espanola 13(1-2): 39-41.

Gimenez, J.C.M. 2003. Bicefalia en Elaphe scalaris (Schinz, 1822) (Ophidia, Colubridae). Butlletí de la Societat Catalana d’Herpetologia 16: 114-141.

Helfenberger, N.; Schulz, K.D. 2013. A checklist of the extinct taxa allied to the Old World ratsnakes, with remarks on the phylogeny and radiation of Elaphe, Rhinechis and Zamenis. pp. 423-427. In: Schulz, K.D. (ed.). Old World ratsnakes: a collection of papers. Bushmaster Publications, Berg. 432 pp.

Hingley, K.J. 1987. Snakes of the genus Elaphe. Their care and breeding in captivity. Part 3. Snake Keeper 1(3): 6-10.

Hingley, K.J. 1996. Keeping and breeding the ladder snake, Elaphe scalaris. Litteratura Serpentium (English Edition) 16(1): 18-20.

Knoepffler, L.P. 1960. Additifs à la faune herpétologique des Îles d'Hyères. Vie et Milieu 11: 331.

Laferrere, M. 1970. Observations erpetologiques. Riviera Scient. 1970: 89-90.

Lesparre, D. 2001. Un caso de albinismo en la culebra de escalera (Elaphe scalaris). Boletin de la Asociacion Herpetologica Espanola 12(1): 17.

Manjon, N. 2011. [A case of total albinism in Rhinechis scalaris]. (In Spanish). Boletin de la Asociacion Herpetologica Espanola 22: 78-79.

Mouchet, M. 1950. Sur un specimen de Coluber (Elaphis) quaterradiatus Gmel.; tue a Arbonne en 1943. Y aurait-il identite entre Coluber quaterradiatus et C. aesculapii forma romanus? Ovoviviparite de Coronella austriaca. Sur la frequence des ophidiens dans le Massif de Fontainebleau. Bull. Ass. Nat. Loing 26(10): 107-108.

Nicolau, S.M. 1956. Anfibios y reptiles de la Pobla de Segur. Pinneos 12(39-42): 45-55.

Niederhauser, H. 1981. Die Zucht der Treppennatter Elaphe scalaris. Aquarium (Minden) 15(144): 314-318.

Nulchis, V.; Biaggini, M.; Carretero, M.A.; Harris, D.J. 2008. Unexpectedly low mitochondrial DNA variation within the ladder snake Rhinechis scalaris. North-Western Journal of Zoology 4(1): 119-124.

Pedrajas, L.; Ceacero, F.; García-Muñoz, E. 2011. Natural history notes: Rinechis [rhinechis] scalaris (Ladder Snake). Maximum size. Herpetological Review 42(2): 303.

Pleguezuelos, J.M.; Feriche, M. 2006. Reproductive ecology of a Mediterranean ratsnake, the ladder snake Rhinechis scalaris (Schinz, 1822). Herpetological Journal 16(2): 177-182.

Pleguezuelos, J.M.; Feriche, M.; Garcia-Pena, H. 1990. Caracteres externes et coloration chez Elaphe scalaris (Schinz, 1822) (Squamata, Colubridae) de la Peninsule Iberique. Bulletin de la Societe Herpetologique de France 55: 24-39.

Pleguezuelos, J.M.; Feriche, M.; Reguera, S.; Santos, X. 2010. Patterns of tail breakage in the ladder snake (Rhinechis scalaris) reflect differential predation pressure according to body size. Zoology (Jena) 113(5): 269-274.

Pleguezuelos, J.M.; Feriche, M.; Santos, X. 2013. Tail-breakage effects on snake-body condition. Zoologischer Anzeiger 252(2): 243-245.

Pleguezuelos, J.M.; Fernandez-Cardenete, J.R.; Honrubia, S.; Feriche, M. 2007. Correlates between morphology, diet and foraging mode in the ladder snake Rhinechis scalaris (Schinz, 1822). Contributions to Zoology 76(3): 179-186.

Pleguezuelos, J.M.; Sá-Sousa, P.; Pérez-Mellado, V.; Marquez, R.; Cheylan, M.; Corti, C.; Martínez-Solano, I. 2009. Rhinechis scalaris. The IUCN Red List of Threatened Species 2009: e.T61565A12516902. http://dx.doi.org/10.2305/IUCN.UK.2009.RLTS.T61565A12516902.en.

Rage, J.C. 1970. Elaphe scalaris (la couleuvre à échelons): son extension en Provence. Bull. mens. Soc. linn. Lyon 39: 206-208.

Rijst, H. van der 1988. Successful breeding with Elaphe scalaris. Litteratura Serpentium (English Edition) 8(2): 62-66.

Silva-Rocha, I.; Salvi, D.; Sillero, N.; Mateo, J.A.; Carretero, M.A. 2015. Snakes on the Balearic Islands: an invasion tale with implications for native biodiversity conservation. PLoS ONE 10(4): e0121026.

Stevens, K. 1995. The European ratsnakes of the genus Elaphe. British Herpetological Society Bulletin 54: 10-20.

Zamenis situla

Anonymous. 1980. Umfrage zur Terrarienhaltung von Elaphe situla. Herpetofauna (Weinstadt) 2(6): 31-32.

Bergman, J. 1995. Neues zur Herpetofauna der Insel Skiathos, nördliche Sporaden, Griechenland. Herpetofauna (Weinstadt) 17(98): 26-28.

Böhme, W.; Lymberakis, P.; Ajtic, R.; Tok, C.V.; Ugurtas, I.H.; Sevinç, M.; Crochet, P.A.; Corti, C.; Haxhiu, I.; Sindaco, R.; Avci, A.; Crnobrnja-Isailovic, J.; Kumlutas, Y. 2009. Zamenis situla. The IUCN Red List of Threatened Species 2009: e.T61444A12485786. http://dx.doi.org/10.2305/IUCN.UK.2009.RLTS.T61444A12485786.en.

Bolle, S. 1952. Elaphe leopardina - ein heikler Pflegling? DATZ (Die Aquarien- und Terrarien-Zeitschrift) 5(3): 78-80.

Boost, E. 1981. Pflege und Zucht der Leopardnatter (Elaphe situla). DATZ (Die Aquarien- und Terrarien-Zeitschrift) 34(1): 27-30.

Broggi, M.F. 1978. Herpetologische Beobachtungen auf der Insel Lesbos (Griechenland). Salamandra 14(4): 161-171.

Broggi, M.F. 1994. Feldherpetologische Beobachtungen und Bemerkungen zu schutzenswerten Biotopen auf griechischen Inseln (Amphibia; Reptilia; Griechenland). Herpetozoa 7(1-2): 29-34.

Broggi, M.F. 1999. Notizen zur Herpetofauna der Ägäisinsel Gokceada. Herpetozoa 12(1-2): 73-78.

Broggi, M.F. 2010. The herpetofauna of Alonissos (northern Sporades, Greece) (Amphibia, Reptilia). Herpetozoa 23(1-2): 71-78.

Broggi, M.F. 2014. The herpetofauna of Kimolos (Milos Archipelago, Greece). Herpetozoa 27(1-2): 102-103.

Bruno, S. 1969. Morfologia, distribuzione e biologia di Elaphe situla (Linnaeus) 1758. Atti Accad. Gioenia Sci. nat. Catania (Ser. 7) 1: 3-42.

Buchholz, K.F.; Schultze-Westrum, T. 1964. Zur Kenntnis der Schlangenfauna der nördlichen Sporaden. Zoologischer Anzeiger 173: 127-136.

Carlino, P.; Pauwels, O.S.G. 2013. Natural history notes: Zamenis situla (Leopard Snake). Diet and feeding behavior. Herpetological Review 44(3): 527.

Cattaneo, A. 1985. Reperti erpetologici nelle Cicladi occidentali: Testudo marginata a Milos, Hemidactylus turcicus ed Elaphe situla a Sifnos (Reptilia). Natura (Milano) 75(1-4) [1984]: 75-78.

Cattaneo, A. 1989. Note espetologiche sulle isole greche di Serifos, Sifnos e Milos (Cicladi occidentali). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 130(2): 57-76.

Cattaneo, A. 1997. L'erpetofauna dell'isola greca di Skiathos (Sporadi settentrionali). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 136(2): 145-156.

Cattaneo, A. 2010. Note eco-morfologiche su alcune species ofidiche egee, con particolare riferimento alle popolazioni delle cicladi centro-orientali (Reptilia). Naturalista Siciliano 34(3-4): 319-350.

Cimatti, E. 2003. Sacred snakes of the Mediterranean. Reptilia (GB) 26: 52-58.

Dimitropoulos, A. 1988. A white striped form of leopard snake Elaphe situla (Linnaeus, 1738) from N.E. Greece (Serpentes, Colubridae). Herptile 13(3): 131-132.

Duerr, P. 2004. Vergleichende Längenmessung bei Schlangen und deren abgestreiften Häuten. Elaphe 12(3): 59.

Dusej, G. 1986. Zum Vorkommen der Leopardnatter Elaphe situla (Linnaeus, 1758) auf Kea (NW Cycladen) (Serpentes: Colubridae). Salamandra 22(2-3): 213-214.

Farkas, B.; Toth, T. 1999. Geographic distribution: Elaphe situla (Leopard Snake). Herpetological Review 30(2): 113.

Grillitsch, H.; Tiedemann, F. 1984. Zur Herpetofauna der griechischen Inseln Kea, Spanopoula, Kithnos, Sifnos, Kitriani (Cycladen), Alonissos und Piperi (nördliche Sporaden). Annalen des Naturhistorischen Museums in Wien Serie B Botanik und Zoologie 86: 7-28.

Henle, K. 1989. Herpetologische Beobachtungen in Griechenland. Herpetofauna (Weinstadt) 11(59): 6-10.

Hingley, K.J. 1987. Snakes of the genus Elaphe. Their care and breeding in captivity. Part 3. Snake Keeper 1(3): 6-10.

Hingley, K.J. 1992. Keeping and breeding the Leopard Snake, Elaphe situla, in captivity. Herptile 17(3): 120-125.

Hingley, K.J. 1993. A second successful breeding of the Leopard Snake, Elaphe situla. Herptile 18(1): 40-41.

Hingley, K.J. 1995. Captive husbandry and reproduction of the Leopard Snake Elaphe situla. Litteratura Serpentium (English Edition) 15(5): 123-127.

Hribal, J. 2006. Elaphe situla. The leopard snake. Reptilia (GB) 44: 70-73.

Karmyshev, Y.V. 2001. [New data on distribution of the leopard snake (Elaphe situla) in Crimea]. (In Russian). Vestnik Zoologii 35(1): 58.

Klesius, T. 2015. Die Leopardnatter – Zamenis situla. Haltung und Nachzucht der „schönsten Schlange Europas“. Kleintierverlag Thorsten Geier, Biebertal. 71 pp.

Kukushkin, O.V.; Tsvelykh, A.N. 2004. [Distribution and ecological-morphological peculiarities of the leopard snake, Elaphe situla (Serpentes, Colubridae), in the Crimea]. (In Russian, English summary). Zoologicheskii Zhurnal 83(4): 439-448.

Kyriazi, P.; Kornilios, P.; Nagy, Z.T.; Poulakakis, N.; Kumlutas, Y.; Ilgaz, C.; Avci, A.; Göcmen, B.; Lymberakis, P. 2013. Comparative phylogeography reveals distinct colonization patterns of Cretan snakes. Journal of Biogeography 40(6): 1143-1155.

Lederer, G. 1934. Ein Beitrag zur Ökologie der Leopardnatter (Elaphe situla L. = Coluber leopardinus Bon.). Blätter für Aquarien- und Terrarienkunde 1934: 341-343.

Lindholm, W.A. 1929. Zur Nomenklatur einiger Eidechsen und Schlangen Europas. Zoologischer Anzeiger 81: 73-82, 96.

Mangili, E. 1947. Osservazioni su la riproduzione in cattivita di alcune specie di ofidii Italiani. Acta pont.acad. Scient. 11(20): 235-246.

Marchetti, C. 1968. Die Leopardnatter, Elaphe situla (Linnaeus, 1758), ihre Eier und Aufzucht. Aqua Terra 5: 22-24.

Mertens, R.; Müller, L. 1929. Zur Nomenklatur einiger Eidechsen und Schlangen Europas. Zoologischer Anzeiger 84: 295-302.

Moravec, J.; Böhme, W. 2005. Morphological variation and sex ratio in the leopard snake (Zamenis situla) from Sozopol (Bulgaria). Russian Journal of Herpetology 12: 74-76. (Supplement).

Naumov, B.; Tomovic, L.M. 2007. A review of distribution and conservation status of Zamenis situla (Linnaeus, 1758) (Reptilia: Colubridae) in Bulgaria. Acta Herpetologica 2(1): 7-10.

Obst, F.J. 1981. Bemerkenswerte Schlangen aus Bulgarien in unserer Sammlung: Coluber rubriceps Venzmer und Elaphe situla (L.). (Reptilia, Ophidia, Colubridae). Faunistische Abhandlungen (Dresden) 8(17): 177-180.

Obst, F.J.; Szczerbak, N.N.; Böhme, W. 1993. Elaphe situla - Leopardnatter. pp. 431-453. In: Böhme, W. (ed.). Handbuch der Reptilien und Amphibien Europas. Band 3/I: Schlangen (Serpentes) 1 (Typhlopidae, Boidae, Colubridae 1: Colubrinae). Aula-Verlag, Wiesbaden. 480 pp.

Paynot, J.P.; Drucker, B. 1990. Elevage et reproduction d'Elaphe situla (L.) (Serpentes, Colubridae) en terrarium interieur. Bulletin de la Societe Herpetologique de France 54: 74-78.

Pozio, E. 1983. The biology of freeliving and captive Elaphe situla. Litteratura Serpentium 3(2-3): 50-60.

Ratnikov, V.Y. 2004. Identification of some eurasian species of Elaphe (Colubridae, Serpentes) on the basis of vertebrae. Russian Journal of Herpetology 11(2): 91-98.

Rugiero, L.; Capizzi, D.; Luiselli, L. 1998. Aspects of the ecology of the Leopard Snake, Elaphe situla, in southeastern Italy. Journal of Herpetology 32(4): 626-630.

Schreiber, R. 2011. Haltung und Zucht der Leopardnatter - Zamenis situla (Linnaeus, 1758). Sauria (Berlin) 33(2): 3-8.

Sigg, H. 1988. Demanding beauty - the requirements of Elaphe situla in its natural habitat and in the vivarium. Snake Keeper 2(1): 4-11.

Sonntag, H. 1965. Meine ersten Erfahrungen mit Leopardnattern. DATZ (Die Aquarien- und Terrarien-Zeitschrift) 18: 54-56.

Stevens, K. 1995. The European ratsnakes of the genus Elaphe. British Herpetological Society Bulletin 54: 10-20.

Tkachev, D. 1993. Distribution and biology of Elaphe situla in the Crimean Peninsula, notes on keeping in the terrarium. Litteratura Serpentium (English Edition) 13(4): 128-131.

Werb, K. 1980. Notes on the leopard snake, Elaphe situla with comments on the maintenance of this species in captivity. British Herpetological Society Bulletin 2: 29-30.

Werb, K. 1998. Additional notes on breeding the Leopard Snake Elaphe situla in captivity, including the believed first record of double clutching. Herptile 23(4): 179-181.

Werb, K. 1998. Breeding the Leopard Snake, Elaphe situla in captivity. Herptile 23(2): 68-71.

Wirth, M. 2009. Königinnen der Europäischen Schlangen: Vierstreifennater (Elaphe quatuorlineata), Äskulap-natter (Zamenis longissimus) und Leopardnatter (Zamenis situla). Draco 10(3) (39): 61-74.

Zuffi, M.A.L.; Carlino, P. 2004. The observations on the biometry and the reproductive biology of the Leopard snake, Zamenis situlus, in Italy. Italian Journal of Zoology (Modena) 71: 113-116.