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ę Rune Midtgaard

Taxonomy of the family Trionychidae
Bibliography of the genus Pelodiscus
Biodiversity of the family Trionychidae


East Asian Softshell Turtles

Fern÷stliche Weichschildkr÷ten

ěstasiatiske Bl°dskildpadder

1844 Pelodiscus Gray (type species: Trionyx sinensis Wiegmann 1835)
1869 Landemania Gray (type species: Landemania irrorata Gray 1869; syn. Fritz & Havas 2006)
1880 Ceramopelta Heude (type species: Ceramopelta latirostris Heude 1880; syn. Fritz & Havas 2006)
1880 Cinctisternum Heude (type species: Cinctisternum bicinctum Heude 1880; syn. Fritz & Havas 2006)
1880 Coelognathus Heude [not Coelognathus Fitzinger 1843 (Serpentes); not Coelognathus Hessling 1852 (Acarina)] (type species: Coelognathus novem-costatus Heude 1880; syn. Fritz & Havas 2006)
1880 Coptopelta Heude (type species: Coptopelta septem-costata Heude 1880; syn. Fritz & Havas 2006)
1880 Gomphopelta Heude (type species: Gomphopelta officinae Heude 1880; syn. Fritz & Havas 2006)
1880 Psilognathus Heude (type species: Psilognathus laevis Heude 1880; syn. Fritz & Havas 2006)
1880 Temnognathus Heude (type species: Temnognathus mordax Heude 1880; syn. Fritz & Havas 2006)
1880 Tortisternum Heude (type species: Tortisternum novem-costatum Heude 1880; syn. Fritz & Havas 2006)

Contents: 7 species, of which 3 (42.9%) is endemic.
Endemism: 0% 100%
Remarks: Previously included in Trionyx (e.g., Wermuth & Mertens 1977). Fritz, Gong, Auer, Kuchling, Schneeweiss & Hundsd÷rfer (2010) found seven distinct genetic lineages within Pelodiscus, but the taxonomic and nomenclatural status of these is still unresolved, apart from axenaria, maackii, parviformis, and sinensis. The validity of these four species was supported by Stuckas & Fritz (2011).
Distribution: E. Asia.
Reported from: China (Anhui, Beijing, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Heilongjiang, Henan, Hong Kong, Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Jilin, Liaoning, Macau, Ningxia, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan, Zhejiang), Japan (Honshu, Kyushu), Laos, North Korea, Russia, South Korea, Taiwan, Vietnam.

Pelodiscus axenaria

Hunan Softshell Turtle



1991 Trionyx axenaria Zhou, Zhang & Fang
Pelodiscus axenaria David 1994

Remarks: Validity confirmed by Chen, Liu & Zhang (2005), Chen, Liu, Li & Zhang (2006), and Gong, Fritz, Vamberger, Gao & Farkas (2022). Previously included the populations now assigned to huangshanensis and shipian.
Distribution: China (Guangdong, Guangxi, Hunan, Jiangxi).

Pelodiscus huangshanensis

Huangshan Softshell Turtle



2021 Pelodiscus huangshanensis Gong, Peng, Huang, Lin, Huang, Xu, Yang & Nie

Remarks: Previously regarded as a population of axenaria.
Distribution: China (Anhui).

Pelodiscus maackii

Amur Softshell Turtle, Northern Chinese Softshell Turtle



1858 Trionyx maackii Brandt
Amyda maackii Stejneger 1907
Pelodiscus maackii David 1994

Remarks: Validity as a separate species confirmed by Fritz, Gong, Auer, Kuchling, Schneeweiss & Hundsd÷rfer (2010).
Distribution: China (Heilongjiang, Inner Mongolia, Jilin, Liaoning), Japan (Honshu, Kyushu), North Korea, Russia, South Korea.

Pelodiscus parviformis

Lesser Chinese Softshell Turtle


Kinesisk DvŠrgbl°dskildpadde

1997 Pelodiscus parviformis Tang

Remarks: Validity of parviformis was supported by the molecular and morphological studies of Yang, Tang, Ding, Guo & Wang (2011).
Distribution: China (Guangxi, Hunan).

Pelodiscus shipian

Chinese Stone Slab Soft-shelled Turtle



2022 Pelodiscus shipian Gong, Fritz, Vamberger, Gao & Farkas

Remarks: Previously regarded as a population of axenaria (Gong, Fritz, Vamberger, Gao & Farkas 2022).
Distribution: China (Hunan, Jiangxi).

Pelodiscus sinensis

Chinese Softshell Turtle

Chinesische Weichschildkr÷te

Kinesisk Bl°dskildpadde

1787 Testudo rostrata Thunberg [suppressed] (ICZN 1991: Opinion 1659; Fritz & Havas 2006)
Emydes rostrata Brongniart 1805
1804 Testudo semimembranacea Hermann [rejected] (ICZN 1963: Opinion 660)
1835 Trionyx sinensis Wiegmann 1835
Pelodiscus sinensis Fitzinger 1843
Amyda sinensis Stejneger 1907
Pelodiscus sinensis sinensis Ferri 2002
1835 Trionyx stellatus var. japonica Temminck & Schlegel in Siebold (Fritz & Havas 2006)
Trionyx japonicus Temminck & Schlegel in Siebold 1835
Amyda japonica Stejneger 1907
Pelodiscus sinensis japonicus Joseph-Ouni 2004
1842 Trionyx tuberculatus Cantor (Fritz & Havas 2006)
Potamochelys tuberculatus Gray 1864
Amyda tuberculata Schmidt 1927
Trionyx sinensis tuberculatus Smith 1931
Pelodiscus sinensis tuberculatus Ferri 2002
1844 Tyrse perocellata Gray (Fritz & Havas 2006)
Trionyx perocellatus Gray 1856
Potamochelys perocellatus Gray 1864
Landemania perocellata Gray 1869
Gymnopus perocellatus David 1872
1857 Trionyx schlegelii Brandt (Fritz & Havas 2006)
Amyda schlegelii Stejneger 1907
1869 Landemania irrorata Gray (Fritz & Havas 2006)
1880 Ceramopelta latirostris Heude (Fritz & Havas 2006)
1880 Cinctisternum bicinctum Heude (Fritz & Havas 2006)
1880 Coelognathus novem-costatus Heude (Fritz & Havas 2006)
Tortisternum novem-costatum Heude 1880
1880 Coptopelta septem-costata Heude (Fritz & Havas 2006)
1880 Gomphopelta officinae Heude (Fritz & Havas 2006)
1880 Psilognathus laevis Heude (Fritz & Havas 2006)
1880 Temnognathus mordax Heude (Fritz & Havas 2006)
1880 Tortisternum novem-costatum Heude (Fritz & Havas 2006)
1887 Trionyx cartilagineus var. newtoni Bethencourt-Ferreira (Fritz & Havas 2006)
Trionyx sinensis newtoni Bethencourt-Ferreira 1897
1932 Amyda schlegelii haseri Pavlov (Fritz & Havas 2006)
1932 Amyda schlegelii licenti Pavlov (Fritz & Havas 2006)
1933 Amyda schlegelii var. laoshanica Pavlov (Fritz & Havas 2006)

Remarks: Records from Hainan refer to parviformis or variegatus (Farkas, Ziegler, Pham, Ong & Fritz 2019). It is uncertain whether the species is indigenous to Laos, where it has recently been recorded (TeyniÚ & David 2010).
Introduced to: Bosnia-Herzegovina, Brazil (Para), Croatia, East Timor, Guam, Indonesia (Kalimantan, Sumatra, West Timor), Iran, Japan (Honshu, Kyushu, Shikoku, Bonin Islands, Ryukyu Islands [incl. Okinawa Islands, Sakishima Islands, Satsunan Islands]), Laos, Latvia, Malaysia (Sarawak, West Malaysia), Northern Mariana Islands, Philippines (Bohol, Cebu, Leyte, Luzon, Mindanao, Mindoro, Panay), Singapore, Slovenia, South Africa (Gauteng), South Korea, Spain, Thailand, USA (Hawaii).
Distribution: China (Anhui, Beijing, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, Hong Kong, Hubei, Hunan, Inner Mongolia, Jiangsu, Jiangxi, Macau, Ningxia, Shaanxi, Shandong, Shanxi, Sichuan, Tianjin, Yunnan, Zhejiang), Taiwan, Vietnam.

Pelodiscus sinensis
© Rune Midtgaard

Pelodiscus variegatus

Large-spotted Softshell Turtle


Storplettet Bl°dskildpadde

2019 Pelodiscus variegatus Farkas, Ziegler, Pham, Ong & Fritz

Distribution: China (Hainan), Vietnam.