Reptiles of Laos    Bibliography of the reptiles of Laos Taxonomic history of the reptiles of Laos Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus siamensis Siamese Freshwater Crocodile Bezuijen, Cox, Thorbjarnarson, Phothitay, Hedemark & Rasphone 2013 Bezuijen, Simpson, Behler, Daltry & Tempsiripong 2012 Mateus 2001 Nguyen, Ho & Nguyen 2009 Souvannasy, Luu, Soudthichak, Wayakone, Le, Nguyen & Ziegler 2018 Souvannasy, Luu, Wayakone, Sounivong, Soudthichak, Nguyen, Le & Ziegler 2021 Stuart 1999 Teynie & David 2010 Ziegler, Luu, Soudthichak & Nguyen 2015

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthosaura (Pricklenapes)   Acanthosaura coronata Crowned Pricklenape Manthey & Manthey 2017 Manthey 2008 [as "cf. coronata"]   Acanthosaura lepidogaster Brown Pricklenape David & Ineich in Bourret 2009 Manthey 2008 [as "cf. lepidogaster"] Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004   Acanthosaura nataliae Gia Lai Pricklenape Ananjeva, Orlov & Kalyabina-Hauf 2008 David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Acanthosaura prasina Lesser Green Pricklenape Brakels, Phiapalath & Nguyen 2021 Genus Calotes (South Asian Forest Agamas)   Calotes bachae Eastern Blue Crested Agama Manthey & Manthey 2017   Calotes emma Forest Crested Agama David & Ineich in Bourret 2009 Manthey & Manthey 2017 Manthey 2008 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie & David 2014   Calotes goetzi Cambodian Blue Crested Agama David & Ineich in Bourret 2009 [as C. mystaceus] Manthey 2008 [as C. mystaceus] Nguyen, Ho & Nguyen 2009 [as C. mystaceus] Stuart 1999 [as C. mystaceus] Teynie & David 2010 [as C. mystaceus] Wagner, Ihlow, Hartmann, Flecks, Schmitz & Boehme 2021   Calotes versicolor South Indian Crested Agama David & Ineich in Bourret 2009 Manthey & Manthey 2017 [as "cf. versicolor"] Manthey 2008 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Genus Draco (Gliding Dragons)   Draco blanfordii Tavoy Gliding Dragon Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Draco maculatus Black-spotted Gliding Dragon David & Ineich in Bourret 2009 Manthey & Manthey 2017 Manthey 2008 [as "cf. maculatus"] Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus grandis Greater Anglehead Agama David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Genus Leiolepis (Butterfly Agamas)   Leiolepis belliana Common Butterfly Agama David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010   Leiolepis rubritaeniata Red-banded Butterfly Agama David & Ineich in Bourret 2009 Hartmann, Sothanin, Handschuh & Boehme 2012 Manthey & Manthey 2017 Manthey 2010 Nguyen, Ho & Nguyen 2009 Peters 1971 Stuart 1999 Teynie & David 2010 Genus Mantheyus (Phu Wua Rock Agama)   Mantheyus phuwuanensis Phu Wua Rock Agama David & Ineich in Bourret 2009 Manthey & Manthey 2017 Manthey 2010 Stuart 1999 Teynie & David 2010 Genus Physignathus (Indochinese Water Dragon)   Physignathus cocincinus Indochinese Water Dragon David & Ineich in Bourret 2009 Manthey & Manthey 2017 Manthey 2010 Nguyen, Ho & Nguyen 2009 Streicher 2014 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Wermuth 1967 Genus Pseudocalotes (Long-headed Agamas)   Pseudocalotes microlepis Small-scaled Long-headed Agama David & Ineich in Bourret 2009 Hallermann & Boehme 2000 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010   * Pseudocalotes poilani Laotian Long-headed Agama David & Ineich in Bourret 2009 Hallermann & Boehme 2000 Manthey 2010 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Wermuth 1967   Pseudocalotes ziegleri Vietnamese Long-headed Agama Brakels, Maury & Nguyen 2021  Family Anguidae (Slowworms, etc.) Genus Dopasia (Oriental Glass Lizards)   Dopasia gracilis Burmese Glass Lizard Brakels, Souvannakhoummane & Nguyen 2021 Brygoo 1987 David & Ineich in Bourret 2009 Nguyen, Boehme, Nguyen, Le, Pahl, Haus & Ziegler 2011 Nguyen, Ho & Nguyen 2009   Dopasia harti South Chinese Glass Lizard David & Ineich in Bourret 2009 Nguyen, Boehme, Nguyen, Le, Pahl, Haus & Ziegler 2011 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Dopasia sokolovi Buon Luoi Glass Lizard Brakels, Phiapalath & Nguyen 2021  Family Gekkonidae (Typical Geckos) Genus Cnemaspis (Asian Rock and Forest Geckos)   * Cnemaspis laoensis Laotian Rock Gecko Grismer 2010 Grismer, Wood, Anuar, Riyanto, Ahmad, Muin, Sumontha, Grismer, Chan, Quah & Pauwels 2014 Teynie & David 2010 Genus Cyrtodactylus (Australasian Bent-toed Geckos)   * Cyrtodactylus bansocensis Ban Soc Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Luu, Nguyen, Le, Bonkowski & Ziegler 2016 Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020   Cyrtodactylus buchardi Champasak Bent-toed Gecko David, Teynie & Ohler 2004 Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Teynie & David 2010   * Cyrtodactylus calamei Tham Nok Aen Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016   Cyrtodactylus cryptus U Bo Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016   * Cyrtodactylus darevskii Boulapha Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov 2014   * Cyrtodactylus hinnamnoensis Hin Nam No Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Luu, Nguyen, Calame, Hoang, Southichack, Bonkowski & Ziegler 2013 [as phongnhakebangensis]   * Cyrtodactylus jaegeri Khammouane Brown-headed Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Luu, Calame, Bonkowski, Nguyen & Ziegler 2014 Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020   Cyrtodactylus jarujini Phu Wua Bent-toed Gecko David & Ineich in Bourret 2009 Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Stuart 1999 Teynie & David 2010 Youmans & Grismer 2006   * Cyrtodactylus khammouanensis Khammouane Yellow-headed Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov 2014   * Cyrtodactylus lomyenensis Lomyen Cave Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Luu, Calame, Nguyen, Le, Bonkowski & Ziegler 2016 [as rufford] Ngo & Pauwels 2010 Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020 Teynie & David 2014   * Cyrtodactylus muangfuangensis Vientiane Bent-toed Gecko Sitthivong, Luu, Ha, Nguyen, Le & Ziegler 2019   * Cyrtodactylus multiporus Many-pored Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov 2014   Cyrtodactylus ngati Dien Bien Bent-toed Gecko Grismer, Rujirawan, Yodthong, Stuart, Le, Le, Chuaynkern, Wood & Aowphol 2022   * Cyrtodactylus ngoiensis Ngoi Bent-toed Gecko Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020   * Cyrtodactylus pageli Vang Vieng Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020 Schneider, Nguyen, Schmitz, Kingsada, Auer & Ziegler 2011 Teynie & David 2014   Cyrtodactylus pseudoquadrivirgatus Thua Thien-Hue Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Luu, Nguyen, Calame, Hoang, Southichack, Bonkowski & Ziegler 2013   * Cyrtodactylus sommerladi Hang Toi Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016   * Cyrtodactylus soudthichaki Khun Don Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Luu, Calame, Nguyen, Bonkowski & Ziegler 2015 Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020   * Cyrtodactylus spelaeus Khuang Lang Cave Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov 2014   * Cyrtodactylus teyniei Borikhamxay Bent-toed Gecko David, Nguyen, Schneider & Ziegler 2011 Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020 Teynie & David 2014   * Cyrtodactylus thathomensis Thathom Bent-toed Gecko Nazarov, Pauwels, Konstantinov, Chulisov, Orlov & Poyarkov 2018   * Cyrtodactylus vilaphongi Luang Prabang Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Schneider, Nguyen, Le, Nophaseud, Bonkowski & Ziegler 2014   Cyrtodactylus wayakonei Vieng Phoukha Bent-toed Gecko Luu, Bonkowski, Nguyen, Le, Schneider, Ngo & Ziegler 2016 Nguyen, Kingsada, Roesler, Auer & Ziegler 2010 Schneider, Luu, Sitthivong, Teynie, Le, Nguyen & Ziegler 2020 Genus Dixonius (Southeast Asian Leaf-toed Geckos)   * Dixonius lao Khammouane Leaf-toed Gecko Nguyen, Sitthivong, Ngo, Luu, Nguyen, Le & Ziegler 2020   Dixonius siamensis Siamese Leaf-toed Gecko David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   * Dixonius somchanhae Nasaithong Leaf-toed Gecko Nguyen, Luu, Sitthivong, Ngo, Nguyen, Le & Ziegler 2021 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko David & Ineich in Bourret 2009 Stuart 1999 Teynie & David 2010 Genus Gekko (Tokays)   * Gekko boehmei Bualapha Grey-headed Karst Gecko Luu, Calame, Nguyen, Le & Ziegler 2015   * Gekko bonkowskii Lesser Thakhek Karst Gecko Luu, Calame, Nguyen, Le & Ziegler 2015   Gekko gecko Tokay David & Ineich in Bourret 2009 Manthey & Manthey 2017 Nguyen, Ho & Nguyen 2009 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004   * Gekko kabkaebin Laotian Gliding Gecko Grismer, Wood, Grismer, Quah, Neang, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza & Brown 2019 [as Ptychozoon kabkaebin]   * Gekko khunkhamensis Khunkham Gecko Sitthivong, Lo, Nguyen, Ngo, Khotpathoom, Le, Ziegler & Luu 2021   Gekko lionotum Smooth-backed Gliding Gecko Grismer, Wood, Grismer, Quah, Neang, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza & Brown 2019 [as Ptychozoon lionotum] Manthey & Manthey 2017 [as Ptychozoon lionotum] Teynie & David 2010 [as Ptychozoon lionotum] Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 [as Ptychozoon lionotum]   * Gekko nadenensis Naden Karst Gecko Luu, Nguyen, Le, Bonkowski & Ziegler 2017   Gekko petricolus Ubon Sandstone Gecko David & Ineich in Bourret 2009 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Stuart 1999 Teynie & David 2010   * Gekko sengchanthavongi Bualapha Brown-headed Karst Gecko Luu, Calame, Nguyen, Le & Ziegler 2015   * Gekko thakhekensis Greater Thakhek Karst Gecko Luu, Calame, Nguyen, Le, Bonkowski & Ziegler 2014   Gekko trinotaterra Three-banded Gliding Gecko Grismer, Wood, Grismer, Quah, Neang, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza & Brown 2019 [as Ptychozoon trinotaterra] Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 [as Ptychozoon trinotaterra]   * Gekko aaronbaueri Lom Yen Karst Gecko Ngo, Pham, Phimvohan, David & Teynie 2015 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 [as scientiadventura] Teynie & David 2010 [as scientiadventura] Teynie & David 2014 [as scientiadventura] Genus Hemidactylus (Half-toed Geckos)   Hemidactylus bowringii Hong Kong House Gecko David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Hemidactylus frenatus Asian House Gecko David & Ineich in Bourret 2009 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004   Hemidactylus garnotii Fox Gecko David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Hemidactylus platyurus Flat-tailed Gecko David & Ineich in Bourret 2009 Manthey & Manthey 2017 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Genus Hemiphyllodactylus (Slender Geckos)   * Hemiphyllodactylus indosobrinus Champasak Slender Gecko Eliades, Phimmachak, Sivongxay, Siler & Stuart 2019   * Hemiphyllodactylus kiziriani Laotian Slender Gecko Nguyen, Botov, Le, Nophaseud, Zug, Bonkowski & Ziegler 2014   * Hemiphyllodactylus serpispecus Houaphanh Slender Gecko Eliades, Phimmachak, Sivongxay, Siler & Stuart 2019  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   Takydromus sexlineatus Six-lined Grass Lizard Bischoff 1992 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004  Family Scincidae (Skinks) Genus Dasia (Indo-Malayan Tree Skinks)   Dasia olivacea Olive Tree Skink Greer 1970 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Genus Eutropis (South Asian Sun Skinks)   Eutropis longicaudata Long-tailed Sun Skink David & Ineich in Bourret 2009 Manthey & Manthey 2017 Nguyen, Ho & Nguyen 2009 Smith 1935 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004   Eutropis macularia Speckled Forest Skink David & Ineich in Bourret 2009 Manthey & Manthey 2017 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004   Eutropis multifasciata Many-lined Sun Skink David & Ineich in Bourret 2009 Manthey & Manthey 2017 Nguyen, Ho & Nguyen 2009 O'Shea, Kusuma & Kaiser 2018 Stuart 1999 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia microcerca Laotian Striped Skink David & Ineich in Bourret 2009 [as vittigera] Nguyen, Ho & Nguyen 2009 [as vittigera] Poyarkov, Geissler, Gorin, Dunayev, Hartmann & Suwannapoom 2019 Stuart 1999 [as vittigera] Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 [as vittigera] Genus Lygosoma (Southeast Asian Writhing Skinks)   Lygosoma corpulentum Annam Writhing Skink David & Ineich in Bourret 2009 Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Lygosoma siamense Siamese Writhing Skink David & Ineich in Bourret 2009 [as L. quadrupes] Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011 [as L. quadrupes] Nguyen, Ho & Nguyen 2009 [as L. quadrupes] Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer 2018 Teynie & David 2010 [as L. quadrupes] Teynie, David, Ohler & Luanglath 2004 [as L. quadrupes] Welch, Cooke & Wright 1990 [as L. quadrupes] Genus Plestiodon (Blue-tailed Skinks)   Plestiodon quadrilineatus Four-striped Long Skink Brakels, Xayyasith & Nguyen 2021 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Riopa (Asian Gracile Skinks)   Riopa angeli Dong Nai Gracile Skink Cota, Geissler, Chan-ard & Makchai 2011 David & Ineich in Bourret 2009 Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Riopa haroldyoungi Banded Gracile Skink Chuaynkern, Duengkae, Chuaynkern, Tanomtong & Patawang 2013 Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011 Moravec & Boehme 2008 Orlov, Streltzov & Peney 2010 Teynie & David 2010 Genus Scincella (South Asian and American Ground Skinks)   Scincella melanosticta Black-spotted Ground Skink David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Scincella ochracea Tawny Ground Skink David & Ineich in Bourret 2009 Teynie & David 2010   Scincella reevesii Speckled Ground Skink Stuart 1999 Teynie & David 2010   Scincella rufocaudata Red-tailed Forest Skink David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Scincella rupicola Rock-dwelling Ground Skink David & Ineich in Bourret 2009 Nguyen, Nguyen, Boehme & Ziegler 2010 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Teynie, David, Ohler & Luang-Lath 2004 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus indicus Common Indian Forest Skink David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010   Sphenomorphus lineopunctulatus Southeast Asian Spot-lined Forest Skink Hartmann, Nguyen, Ohler, Chippich, Handschuh & Boehme 2010 Teynie & David 2010   Sphenomorphus maculatus Spotted Forest Skink David & Ineich in Bourret 2009 Manthey & Manthey 2017 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Sphenomorphus tridigitus Three-toed Forest Skink David & Ineich in Bourret 2009 Greer, David & Teynie 2006 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Genus Subdoluseps (Asian Agile Skinks)   Subdoluseps bowringii Short-legged Agile Skink David & Ineich in Bourret 2009 Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Welch, Cooke & Wright 1990 Genus Tropidophorus (Stream Skinks)   Tropidophorus baviensis Bavi Stream Skink Okabe, Sanamxay, Nguyen & Motokawa 2020 Teynie & David 2010   Tropidophorus berdmorei Burmese Stream Skink Cox, Dijk, Nabhitabhata & Thirakhupt 1998 Teynie & David 2010   Tropidophorus cocincinensis Indochinese Stream Skink Chuaynkern, Nabhitabhata, Inthara, Kamsook & Somsri 2005 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Tropidophorus laotus Laotian Stream Skink David & Ineich in Bourret 2009 Smith 1935 Stuart 1999 Teynie & David 2010   Tropidophorus latiscutatus Broad-scaled Stream Skink Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Tropidophorus microlepis Small-scaled Stream Skink Chuaynkern, Nabhitabhata, Inthara, Kamsook & Somsri 2005 David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus nebulosus Clouded Monitor Auffenberg 1994 [as V. bengalensis] Nguyen, Ho & Nguyen 2009 Streicher 2014 [as V. bengalensis] Stuart 1999 Teynie & David 2010   Varanus salvator Asian Water Monitor David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Streicher 2014 Stuart 1999 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla nasuta Long-nosed Green Whip Snake Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Wallach, Williams & Boundy 2014   Ahaetulla prasina Short-nosed Whip Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Wallach, Williams & Boundy 2014 Genus Amphiesma (South Asian Keelbacks)   Amphiesma stolatum Buff-striped Keelback Deuve 1970 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga cyanea Green Cat Snake Deuve 1970 Maury, David & Sudavanh 2022 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Orlov & Ryabov 2002 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie & David 2014   Boiga multomaculata Marbled Cat Snake Deuve 1970 Farkas & Fritz 1999 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Orlov & Ryabov 2002 Stuart 1999 Teynie & David 2010   Boiga siamensis Grey Cat Snake Deuve 1970 [as cynodon] Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie & David 2014 Genus Calamaria (Oriental Reed Snakes)   * Calamaria nebulosa Clouded Reed Snake Lee 2021 Stuart & Heatwole 2008 [as C. yunnanensis] Teynie & David 2010 [as C. yunnanensis]   Calamaria pavimentata Brown Reed Snake Deuve 1970 Inger & Marx 1965 Nguyen, Ho & Nguyen 2009 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Deuve 1970 Manthey & Manthey 2017 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus radiatus Copper-headed Rat Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Schulz 1996 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis cyanochloris Green Bronzeback Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 [as "D. cf. cyanochloris"] Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Dendrelaphis ngansonensis Nganson Bronzeback Deuve 1970 Nguyen, Ho & Nguyen 2009 Nicodemo & Bain 2007 Teynie & David 2010   Dendrelaphis pictus Southern Painted Bronzeback Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Vogel & Rooijen 2011   Dendrelaphis proarchos Northern Painted Bronzeback Biakzuala, Lalremsanga, Tariang, Vabeiryureilai, Muansanga, Hrima, Kumar, Kundu, Purkayastha & Vogel 2022 Vogel & Rooijen 2011   Dendrelaphis subocularis Burmese Mountain Bronzeback Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe taeniura Stripe-tailed Rat Snake Brakels, Nanthavong & Nguyen 2021 Deuve 1970 Nguyen, Ho & Nguyen 2009 Schulz 1996 Stuart 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Euprepiophis (East Asian Rat Snakes)   Euprepiophis mandarinus Mandarin Rat Snake Brakels, Phimmachak & Nguyen 2021 Deuve 1970 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Zhao & Adler 1993 Genus Fowlea (Fishing Keelbacks)   Fowlea flavipunctata Yellow-spotted Keelback Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel 2022 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Vogel & David 2006 Vogel & David 2012   Fowlea piscator Checkered Keelback Deuve 1970 Farkas & Fritz 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Vogel & David 2006 Vogel & David 2012 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma coeruleum Blue-eyed Bush Rat Snake David, Campbell, Deuti, Hauser, Luu, Nguyen, Orlov, Pauwels, Scheinberg, Sethy, Smits, Teynie & Vogel 2022 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 [as prasinum] Nguyen, Ho & Nguyen 2009 [as prasinum] Stuart & Heatwole 2008 [as prasinum] Stuart 1999 [as prasinum] Teynie & David 2010 [as prasinum] Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 [as prasinum] Zhao & Adler 1993 [as prasinum]   Gonyosoma oxycephalum Red-tailed Green Rat Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Schulz 1996 Stuart 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius andreae Quang Binh Keelback Ziegler, Luu, Nguyen, Ha, Ngo, Le & Nguyen 2019   Hebius annamensis Annam Keelback David, Pauwels, Nguyen & Vogel 2015 David, Vogel, Nguyen, Orlov, Pauwels, Teynie & Ziegler 2021 Nguyen, Ho & Nguyen 2009 Stuart & Chuaynkern 2007 Stuart 2006 Teynie & David 2010   Hebius atemporalis Tonkin Keelback Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Wallach, Williams & Boundy 2014   Hebius bitaeniatus Two-striped Keelback Lottier & David 2021   Hebius boulengeri White-browed Keelback Brakels, Maury & Nguyen 2021 David, Vogel & Rooijen 2013 Purkayastha & David 2019 Teynie & David 2010 Wallach, Williams & Boundy 2014   Hebius chapaensis Chapa Flat-nosed Snake David, Pauwels, Nguyen & Vogel 2015 David, Vogel, Nguyen, Orlov, Pauwels, Teynie & Ziegler 2021 Maury, David & Sudavanh 2022 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Ren, Wang, Nguyen, Hoang, Zhong, Jiang, Guo & Li 2018 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Hebius khasiensis Khasi Hills Keelback David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 David, Vogel & Rooijen 2013 Deuve 1970 Maury, David & Sudavanh 2022 Nguyen, Ho & Nguyen 2009 Purkayastha & David 2019 Teynie & David 2010 Wallach, Williams & Boundy 2014   Hebius leucomystax White-lipped Keelback Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Stuart & Heatwole 2008 Teynie & David 2010 Wallach, Williams & Boundy 2014   Hebius sauteri Kosempo Keelback Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Genus Liopeltis (Ringneck Snakes)   Liopeltis frenata Common Ringneck Snake Brakels, Nanthavong & Nguyen 2021 Deuve 1970 Hauser 2018 Nguyen, Ho & Nguyen 2009 Smith 1943 Stuart 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Zhao & Adler 1993   Liopeltis stoliczkae Northeast Indian Ringneck Snake Stuart & Heatwole 2008 Stuart 1999 Teynie & David 2010 Genus Lycodon (Oriental Wolf Snakes)   * Lycodon banksi Khammouane Wolf Snake Luu, Bonkowski, Nguyen, Le, Calame & Ziegler 2018 Wang, Qi, Lyu, Zeng & Wang 2020   Lycodon capucinus Island Wolf Snake Iskandar & Colijn 2001 Luu, Bonkowski, Nguyen, Le, Calame & Ziegler 2018 Maury, David & Sudavanh 2022 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   * Lycodon davidi Vientiane Wolf Snake Luu, Bonkowski, Nguyen, Le, Calame & Ziegler 2018 Teynie & David 2014 Vogel, Nguyen, Kingsada & Ziegler 2012   Lycodon davisonii Burmese Bridle Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010   Lycodon fasciatus Yellow-banded Wolf Snake Deuve 1970 Luu, Bonkowski, Nguyen, Le, Calame & Ziegler 2018 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Smith 1943 Teynie & David 2010 Vogel & David 2010 Vogel & David 2019 Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009   Lycodon futsingensis Fujian Wolf Snake Luu, Bonkowski, Nguyen, Le, Calame & Ziegler 2018 Luu, Nguyen, Calame, Hoang, Southichack, Bonkowski & Ziegler 2013 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020   Lycodon laoensis Laotian Wolf Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Smith 1943 Stuart 1999 Taylor 1965 Teynie & David 2010 Wang, Qi, Lyu, Zeng & Wang 2020 Zhao & Adler 1993   Lycodon meridionalis Vietnamese Big-toothed Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Orlov & Ryabov 2004 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Lycodon ruhstrati Taiwanese Wolf Snake Brakels, Maury & Nguyen 2021 Luu, Bonkowski, Nguyen, Le, Calame & Ziegler 2018 Luu, Nguyen, Calame, Hoang, Southichack, Bonkowski & Ziegler 2013   Lycodon septentrionalis Northern Big-toothed Snake Das & Vasudevan 2015 Nguyen, Ho & Nguyen 2009 Orlov & Ryabov 2004 Smith 1943 Taylor 1965 Teynie & David 2010 Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009   Lycodon subcinctus Malayan Banded Wolf Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Taylor 1965 Teynie & David 2010 Genus Oligodon (Kukri Snakes)   Oligodon barroni Chonburi Kukri Snake David, Vogel & Rooijen 2008 Nguyen, Ho & Nguyen 2009 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Oligodon catenatus Chain-banded Kukri Snake Teynie & David 2010   Oligodon cinereus Black-barred Kukri Snake David, Das & Vogel 2011 David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Deuve 1970 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Zhao & Adler 1993   Oligodon deuvei Orange-striped Kukri Snake David, Vogel & Rooijen 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972 [as mouhoti] Teynie & David 2010   Oligodon eberhardti Tam Dao Kukri Snake Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Zhao & Adler 1993   Oligodon fasciolatus Small-banded Kukri Snake David, Cox, Pauwels, Chanhome & Thirakhupt 2004 David, Das & Vogel 2011 David, Vogel & Pauwels 2008 Nguyen, Ho & Nguyen 2009 Stuart 1999 [as cyclurus] Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Oligodon inornatus Southeast Asian Plain Kukri Snake Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Oligodon joynsoni Grey Kukri Snake David, Das & Vogel 2011 David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Teynie & David 2010   Oligodon nagao Lang Son Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012   Oligodon ocellatus Ocellated Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 David, Vogel & Pauwels 2008 Nguyen, Ho & Nguyen 2009 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Oligodon taeniatus Striped Kukri Snake David, Vogel & Rooijen 2008 Deuve 1970 Nguyen, Ho & Nguyen 2009 Teynie & David 2007 Teynie & David 2010 Zhao & Adler 1993   * Oligodon teyniei Xepian Kukri Snake David, Hauser & Vogel 2022 Genus Opisthotropis (Mountain Stream Keelbacks)   * Opisthotropis durandi Luang Prabang Mountain Stream Keelback Brakels, Maury & Nguyen 2021 David, Pauwels, Nguyen & Vogel 2015 Teynie, Lottier, David, Nguyen & Vogel 2014 Genus Oreocryptophis (Red Mountain Rat Snake)   Oreocryptophis porphyraceus Red Mountain Rat Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Schulz & Helfenberger 1998 Schulz 1996 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Zhao & Adler 1993 Genus Plagiopholis (Blotch-necked Snakes)   Plagiopholis delacouri Laotian Blotch-necked Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Smith 1943 Teynie & David 2010 Genus Pseudoxenodon (Mock Cobras)   Pseudoxenodon bambusicola Bamboo Mock Cobra Nguyen, Ho & Nguyen 2009 Stuart & Heatwole 2008 Stuart 1999 Teynie & David 2010   Pseudoxenodon macrops Common Large-eyed Mock Cobra Farkas & Fritz 1999 Nanhoe & Ouboter 1987 Nguyen, Ho & Nguyen 2009 Stuart & Heatwole 2008 Stuart 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas carinata Malayan Keeled Rat Snake Maury, David & Sudavanh 2022 Nguyen, Fournier, Brakels & Poyarkov 2020   Ptyas dhumnades Black-striped Rat Snake Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Ptyas korros Indochinese Rat Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Streicher 2014 Stuart 1999 Taylor 1965 Teynie & David 2007 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004   Ptyas major Greater Green Snake Deuve 1970 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Ptyas mucosa Common Oriental Rat Snake Deuve 1970 Maury, David & Sudavanh 2022 Nguyen, Ho & Nguyen 2009 Sindaco, Venchi & Grieco 2013 Streicher 2014 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004   Ptyas multicincta Many-banded Green Snake Hauser 2018 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Smith 1943 Teynie & David 2010 Teynie & David 2014 Zhao & Adler 1993   Ptyas nigromarginata Oriental Green Rat Snake Brakels & Nguyen 2020 Lottier & David 2021 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis chrysargos Speckled-bellied Keelback Deuve 1970 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010   Rhabdophis helleri Northern Red-necked Keelback Deuve 1970 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Teynie & David 2010 Zhao & Adler 1993   Rhabdophis leonardi Burmese Mountain Keelback Lottier & David 2021   Rhabdophis nigrocinctus Black-banded Keelback Nguyen, Ho & Nguyen 2009 Stuart & Heatwole 2008 Stuart 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Rhabdophis nuchalis Collared Keelback Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Rhabdophis siamensis Siamese Red-necked Keelback David & Vogel 2021 Deuve 1970 [as s. subminiatus] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as s. subminiatus] Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis collaris Collared Black-headed Snake Brakels, Nanthavong, Khotpathoom & Nguyen 2021 Deuve 1970 Nguyen, Ho & Nguyen 2009 Smith 1943 Stuart 1999 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Zhao & Adler 1993 Genus Trimerodytes (Asian Water Snakes)   Trimerodytes aequifasciatus Equal-banded Water Snake Stuart & Heatwole 2008 Stuart 1999 Teynie & David 2010   Trimerodytes percarinatus Chinese Keelback Mallik, Chowdhury, Bhatt & Captain 2019 Nguyen, Ho & Nguyen 2009 Stuart & Heatwole 2008 Stuart 1999 Teynie & David 2010   Trimerodytes praemaxillaris Brown Stream Snake Deuve 1970 Smith 1943 Stuart 1999 Teynie & David 2010 Zhao & Adler 1993   Trimerodytes yapingi Yaping Keelback Dinh, Phimpasone, Ha, Nguyen, Ziegler & Luu 2022 Genus Xenochrophis (Oriental Painted Keelbacks)   Xenochrophis trianguligerus South Asian Triangle Keelback Nguyen, Ho & Nguyen 2009 Teynie & David 2010  Family Cylindrophiidae (Asian Pipe Snakes) Genus Cylindrophis (Oriental Pipe Snakes)   Cylindrophis ruffus Red-tailed Pipe Snake Deuve 1970 McDiarmid, Campbell & Toure 1999 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus candidus Malayan Krait Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Deuve 1970 Maury, David & Sudavanh 2022 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010   Bungarus fasciatus Banded Krait Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Deuve 1970 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010   Bungarus multicinctus Many-banded Krait Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Stuart 1999   Bungarus slowinskii Red River Krait Brakels, Phiapalath & Nguyen 2021 Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018   Bungarus wanghaotingi Yunnan Krait Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Zhao & Adler 1993 Genus Calliophis (South Asian Coral Snakes)   Calliophis maculiceps Small-spotted Coral Snake Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Deuve 1970 Fukuyama, Hibino, Nishikawa & Sanamxay 2020 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as Maticora maculiceps michaelis] Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Naja (Typical Cobras)   Naja siamensis Siamese Spitting Cobra Chan-ard, Stuart & Wuster 2000 Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Deuve 1970 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Sinomicrurus (East Asian Coral Snakes)   Sinomicrurus annularis Hong Kong Coral Snake Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021   Sinomicrurus kelloggi Indochinese Coral Snake Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Nguyen, Ho & Nguyen 2009 Nguyen, Nanthavong, Coudrat, Brakels & Poyarkov 2020 [as houi] Teynie & David 2010   Sinomicrurus macclellandii Southeast Asian Coral Snake Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Deuve 1970 Nguyen, Ho & Nguyen 2009 Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014  Family Homalopsidae (Australasian Mud Snakes) Genus Enhydris (Asian Mud Snakes)   Enhydris enhydris Striped Mud Snake Deuve 1970 Gyi 1970 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Enhydris subtaeniata Mekong Mud Snake Murphy & Voris 2014 Murphy 2007 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Voris, Murphy, Karns, Kremer & O'Connell 2012 Genus Homalopsis (Masked Water Snakes)   Homalopsis nigroventralis Mekong Masked Water Snake Deuve 1970 Murphy & Voris 2014 Murphy 2007 Murphy, Voris, Murthy, Traub & Cumberbatch 2012 Nguyen, Ho & Nguyen 2009 [as H. buccata] Stuart 1999 [as H. buccata] Teynie & David 2007 [as H. buccata] Teynie & David 2010 Genus Hypsiscopus (Orange-bellied Mud Snake and Matanna Mud Snake)   Hypsiscopus murphyi Northern Orange-bellied Mud Snake Bernstein, Voris, Stuart, Phimmachak, Seateun, Sivongxay, Neang, Karns, Andrews, Osterhage, Phipps & Ruane 2022 Deuve 1970 [as plumbea] Farkas & Fritz 1999 [as plumbea] Gyi 1970 [as plumbea] Manthey & Manthey 2017 [as plumbea] Murphy & Voris 2014 [as plumbea] Murphy 2007 [as plumbea] Nguyen, Ho & Nguyen 2009 [as plumbea] Stuart 1999 [as plumbea] Teynie & David 2007 [as plumbea] Teynie & David 2010 [as plumbea] Teynie & David 2014 [as plumbea] Teynie, David, Ohler & Luanglath 2004 [as plumbea] Genus Subsessor (Southeast Asian Mud Snake)   Subsessor bocourti Southeast Asian Mud Snake Deuve 1970 Murphy 2007 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Deuve 1970 Maury, David & Sudavanh 2022 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Zhao & Adler 1993 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis indochinensis Indochinese Sand Snake Hartmann, Handschuh & Boehme 2011 Nguyen, Ho & Nguyen 2009 Stuart & Heatwole 2008 [as P. condanarus] Stuart 1999 [as P. condanarus] Teynie & David 2007 Teynie & David 2010  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas berdmorei Mon Slug Eater Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022   Pareas formosensis Taiwanese Red-eyed Slug Eater Maury, David & Sudavanh 2022   Pareas geminatus Jiangcheng Slug Eater Ding, Chen, Suwannapoom, Nguyen, Pogarkov & Vogel 2020   Pareas macularius Black-spotted Slug Eater Deuve 1970 Hauser 2017 Maury, David & Sudavanh 2022 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Smith 1943 Teynie & David 2010 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020 Zhao & Adler 1993   Pareas margaritophorus White-spotted Slug Eater Deuve 1970 Hauser 2017 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Deuve 1970 McDiarmid, Campbell & Toure 1999 Nguyen, Ho & Nguyen 2009 Stimson 1969 Stuart 1999 Teynie & David 2010 Genus Python (Typical Pythons)   Python bivittatus Burmese Python Deuve 1970 Nguyen, Ho & Nguyen 2009 Streicher 2014 [as molurus] Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004  Family Typhlopidae (Typical Blind Snakes) Genus Argyrophis (Southeast Asian Blind Snakes)   Argyrophis diardii Greater Blind Snake Deuve 1970 Maury, David & Sudavanh 2022 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Argyrophis muelleri Malayan Bicoloured Blind Snake Iskandar & Colijn 2001 Teynie & David 2010 Genus Ramphotyphlops (Western Pacific Blind Snakes)   Ramphotyphlops lineatus Striped Blind Snake Teynie & David 2010 Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Deuve 1970 McDiarmid, Campbell & Toure 1999 McDowell 1974 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Wallach 2009 Wallach 2020  Family Viperidae (Vipers) Genus Calloselasma (Malayan Pitviper)   Calloselasma rhodostoma Malayan Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Deuve 1970 Gloyd & Conant 1990 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Nguyen, Ho & Nguyen 2009 Stuart 1999 Teynie & David 2010 Genus Ovophis (Asian Mountain Pitvipers)   Ovophis convictus Indo-Malayan Mountain Pitviper Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Ovophis makazayazaya Eastern Mountain Pitviper Maury, David & Sudavanh 2022 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Popeia (South Asian Bamboo Pitvipers)   Popeia popeiorum Northern Bamboo Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Maury, David & Sudavanh 2022 Ngo, Nguyen, Nguyen, Luu, Phimphasone, Ha, Ziegler & Le 2022 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Regenass & Kramer 1981 Stuart 1999 Teynie & David 2010 Vogel, David & Pauwels 2004 Genus Protobothrops (Habus and allies)   Protobothrops cornutus Horned Tree Pitviper Teynie & David 2010   Protobothrops mucrosquamatus Brown-spotted Pitviper Brakels, Maury, Phimmachak & Nguyen 2021 Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Stuart & Heatwole 2008 Stuart 1999 Teynie & David 2010   Protobothrops sieversorum Three-horned Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Nguyen, Ho & Nguyen 2009 Teynie & David 2010 Teynie & David 2014 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014 Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus albolabris Southeast Asian White-lipped Tree Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Ngo, Nguyen, Nguyen, Luu, Phimphasone, Ha, Ziegler & Le 2022 Nguyen, Ho & Nguyen 2009 Regenass & Kramer 1981 Streicher 2014 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie & David 2014   Trimeresurus guoi Jiangcheng Green Pitviper Brakels, Maury & Nguyen 2021   Trimeresurus macrops Big-eyed Green Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Malhotra, Thorpe, Mrinalini & Stuart 2011 Maury, David & Sudavanh 2022 Ngo, Nguyen, Nguyen, Luu, Phimphasone, Ha, Ziegler & Le 2022 Teynie & David 2010 Teynie & David 2014 Teynie, David, Ohler & Luanglath 2004 Genus Viridovipera (Green Tree Pitvipers)   Viridovipera gumprechti Phu Luang Green Tree Pitviper Ao, David, Bordoloi & Ohler 2004 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Ngo, Nguyen, Nguyen, Luu, Phimphasone, Ha, Ziegler & Le 2022 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Teynie & David 2010   Viridovipera stejnegeri Chinese Green Tree Pitviper Ngo, Nguyen, Nguyen, Luu, Phimphasone, Ha, Ziegler & Le 2022 Teynie, Nguyen, Lorvelec, Piquet, Lottier & David 2014   Viridovipera truongsonensis Truong Son Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Teynie & David 2014   Viridovipera vogeli Khao Yai Green Tree Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Ngo, Nguyen, Nguyen, Luu, Phimphasone, Ha, Ziegler & Le 2022 Nguyen, Ho & Nguyen 2009 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004  Family Xenodermidae (Asian Litter Snakes) Genus Fimbrios (Southern Bearded Snakes)   Fimbrios klossi Lesser Bearded Snake Stuart & Heatwole 2008 Teynie & David 2010 Genus Parafimbrios (Northern Bearded Snakes)   Parafimbrios lao Laotian Bearded Snake Nguyen, Pham, Nguyen, Le & Ziegler 2015 Teynie & Hauser 2017 Teynie, David, Lottier, Le, Vidal & Nguyen 2015  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis unicolor Common Asian Sunbeam Snake Deuve 1970 Maury, David & Sudavanh 2022 McDiarmid, Campbell & Toure 1999 Nguyen, Ho & Nguyen 2009 Orlov 2000 Stuart 1999 Teynie & David 2007 Teynie & David 2010 Teynie, David, Ohler & Luanglath 2004

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schoppe & Das 2011 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010   Cuora bourreti Central Indochinese Box Turtle Lehr, Fritz & Obst 1998 McCord & Joseph-Ouni 2002 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Hallam, Sayavong, Nanthavong, Sayaleng, Vongsa & Robichaud 2011 Teynie & David 2010   Cuora cyclornata Vietnamese Three-striped Box Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Cuora galbinifrons Northern Indochinese Box Turtle Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Streicher 2014 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010   Cuora mouhotii Keeled Box Turtle Das 1987 Das, McCormack, Dijk, Hoang & Struijk 2016 King & Burke 1989 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010   Cuora picturata Southern Indochinese Box Turtle Nguyen, Ho & Nguyen 2009 Genus Cyclemys (Asian Leaf Turtles)   Cyclemys oldhamii Dark-throated Leaf Turtle Fritz & Havas 2006 [as shanensis tcheponensis] Fritz, Guicking, Auer, Sommer, Wink & Hundsdoerfer 2008 King & Burke 1989 [as dentata] Nguyen, Ho & Nguyen 2009 [as dentata] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Streicher 2014 [as tcheponensis] Stuart 1999 [as dentata] Teynie & David 2010 [as tcheponensis] Genus Geoemyda (Black-breasted Leaf Turtles)   Geoemyda spengleri Common Black-breasted Leaf Turtle Dawson, Gaillard, Gong, Lin, McCormack, Nanthavong, Nguyen, Nguyen, Pham & Shi 2022 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Hallam, Sayavong, Nanthavong, Sayaleng, Vongsa & Robichaud 2011 Genus Heosemys (Forest Turtles)   Heosemys annandalii Yellow-headed Temple Turtle Fritz & Havas 2006 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010   Heosemys grandis Giant Asian Pond Turtle Fritz & Havas 2006 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Streicher 2014 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010 Genus Malayemys (Southeast Asian Snail-eating Turtles)   Malayemys khoratensis Khorat Snail-eating Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sumontha, Brophy, Kunya, Wiboonatthapol & Pauwels 2016 [as isan]   Malayemys subtrijuga Mekong Snail-eating Turtle Brophy 2005 Dawson, Ihlow & Platt 2020 Fritz & Havas 2006 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Streicher 2014 Stuart & Timmins 2000 Stuart 1999 Teynie & David 2010 Genus Sacalia (Eastern Eyed Turtles)   Sacalia quadriocellata Four-eyed Turtle Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010 Genus Siebenrockiella (Black Marsh Turtle and Philippine Pond Turtle)   Siebenrockiella crassicollis Black Marsh Turtle Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Teynie & David 2010  Family Platysternidae (Big-headed Turtle) Genus Platysternon (Big-headed Turtle)   Platysternon megacephalum Big-headed Turtle King & Burke 1989 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010  Family Testudinidae (Tortoises) Genus Indotestudo (Asian Tortoises)   Indotestudo elongata Elongated Tortoise Fritz & Havas 2006 Ihlow, Dawson, Hartmann & Som 2016 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Streicher 2014 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010 Genus Manouria (Indochinese Tortoises)   Manouria impressa Impressed Tortoise Calame, Gray, Hurley, Timmins & Thongsamouth 2013 King & Burke 1989 Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom & Poyarkov 2020 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Streicher 2014 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010  Family Trionychidae (Softshell Turtles) Genus Amyda (Southeast Asian Softshell Turtles)   Amyda ornata Black-rayed Softshell Turtle Auliya, Dijk, Moll & Meylan 2016 [as A. cartilaginea] Fritz, Gemel, Kehlmaier, Vamberger & Praschag 2014 King & Burke 1989 [as A. cartilaginea] Nguyen, Ho & Nguyen 2009 [as A. cartilaginea] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart & Timmins 2000 [as A. cartilaginea] Stuart 1999 [as A. cartilaginea] Stuart, Dijk & Hendrie 2001 [as A. cartilaginea] Teynie & David 2010 [as A. cartilaginea] Teynie, David, Ohler & Luanglath 2004 [as A. cartilaginea] Genus Palea (Wattle-necked Softshell Turtle)   Palea steindachneri Wattle-necked Softshell Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Das 2008 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart & Timmins 2000 Stuart 1999 Stuart, Dijk & Hendrie 2001 Teynie & David 2010 Wermuth & Mertens 1977 [as bibroni]