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Taxonomy of the genus Chironius Biodiversity of the family Colubridae

Bibliography of the genus
Chironius (Sipos)

(Reptilia: Serpentes: Colubridae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Chironius in general

Bailey, J.R. 1955. The snakes of the genus Chironius in southeastern South America. Occasional Papers of the Museum of Zoology University of Michigan 571: 1-21.

Cunha, O.R. da; Nascimento, F.P. do 1982. Ofidios da Amazonia. 18. O genero Chironius Fitzinger, na Amazonia oriental (Ophidia: Colubridae). Boletim do Museu Paraense Emilio Goeldi Nova Serie Zoologia 119: 1-17.

Cunha, O.R. da; Nascimento, F.P. do 1983. Ofidios da Amazonia. 15. As especies de Chironius da Amazonia oriental (Para, Amapa e Maranhao). (Ophidia: Colubridae). Memorias do Instituto Butantan (Sao Paulo) 46 [1982]: 139-172.

Dixon, J.R.; Wiest, J.A.; Cei, J.M. 1993. Revision of the Neotropical snake genus Chironius Fitzinger (Serpentes, Colubridae). Museo Regionale di Scienze Naturali Monografie (Turin) 13: 1-280.

Donoso-Barros, R. 1969. Contribution al conocimiento del genero Chironius (Serpentes: Colubridae). Boletin de la Sociedad de Biologia de Concepcion 41: 189-197.

Goldberg, S.R. 2007. Note on reproduction of whipsnakes, genus Chironius (Serpentes: Colubridae), from Costa Rica. Bulletin of the Chicago Herpetological Society 42(9): 148-149.

Hamdan, B.; Pereira, A.G.; Loss-Oliveira, L.; Rödder, D.; Schrago, C.G. 2017. Evolutionary analysis of Chironius snakes unveils cryptic diversity and provides clues to diversification in the Neotropics. Molecular Phylogenetics & Evolution 116: 108-119

Henderson, R.W. 2004. Lesser Antillean snake faunas: distribution, ecology, and conservation concerns. Oryx 38(3): 311-320.

Hollis, J.L. 2006. Phylogenetics of the genus Chironius Fitzinger, 1826 (Serpentes, Colubridae) based on morphology. Herpetologica 62(4): 435-453.

Klaczko, J.; Montingelli, G.G.; Zaher, H. 2014. A combined morphological and molecular phylogeny of the genus Chironius Fitzinger, 1826 (Serpentes: Colubridae). Zoological Journal of the Linnean Society 171(3): 656-667.

Müller, P. 1969. Zur Verbreitung der Gattung Chironius auf den südbrasilianischen Inseln. (Serpentes, Colubridae). Senckenbergiana Biologica 50: 133-141.

Peters, W.C.H. 1868. Über neue Säugethiere (Colobus, Rhinolophus, Vesperus) und neue oder weniger bekannte Amphibien (Hemidactylus, Herpetodryas, Spilotes, Elaphis, Lamprophis, Erythrolamprus). Monatsberichte der Königlich Akademie der Wissenschaften zu Berlin 1868(December): 637-642.

Torres-Carvajal, O.; Echevarría, L.Y.; Lobos, S.E.; Venegas, P.J.; Kok, P.J.R. 2018. Phylogeny, diversity and biogeography of Neotropical Sipo Snakes (Serpentes: Colubrinae: Chironius). Molecular Phylogenetics and Evolution 130 [2019]: 315-329.

Wiest, J.A. 1979. Revision of the Neotropical snake genus Chironius Fitzinger (Serpentes, Colubridae). Dissertation Abstracts International (B) 39(10): 4773.

Chironius bicarinatus

Almeida, L.D.S.; Almeida, L.H. de; Pedrozo, M.; Toledo Moroti, M. de 2018. Natural history notes: Chironius bicarinatus (Two-headed Sipo). Diet. Herpetological Review 49(4): 750.

Almeida-Santos, S.M.; Marques, O.A.V. 2002. Male-male ritual combat in the colubrid snake Chironius bicarinatus from the Atlantic Forest, southeastern Brazil. Amphibia-Reptilia 23(4): 528-533.

Bovo, R.P.; Sueiro, L.R. 2012. Records of predation on Itapotihyla langsdorffi (Anura: Hylidae) by Chironius bicarinatus (Serpentes: Colubridae) with notes on foraging substrate. Herpetology Notes 5: 291-292.

Carreira, S.; Lombardo, I. 2007. Geographic distribution: Chironius bicarinatus (Two-keeled Whipsnake). Herpetological Review 38(1): 102.

Carvalho-Silva, S.P. de; Fernandes, R. 1994. Natural history notes: Chironius bicarinatus (NCN). Foraging behavior. Herpetological Review 25(1): 27.

Guimaraes, M.; Gaiarsa, M.P.; Cavalheri, H.B. 2014. Morphological adaptations to arboreal habitats and heart position in species of the Neotropical Whipsnakes genus Chironius. Acta Zoologica (Copenhagen) 95(3): 341-346.

Marques, O.A.V.; Almeida-Santos, S.M.; Rodrigues, M.; Camargo, R. 2009. Mating and reproductive cycle in the Neotropical colubrid snake Chironius bicarinatus. South American Journal of Herpetology 4(1): 76-80.

Marques, O.A.V.; Falcetti, C.A.; Ribeiro, R.A.K.; Pereira, D.N. 2004. Natural history notes: Bothrops Jararacussu (Jararacussu). Ophiophagy. Herpetological Review 35(1): 58.

Oliveira, S.V. de 2008. Observaçao do comportamento predatorio de Chironius bicarinatus (Serpentes, Colubridae) em Hypsiboas pulchellus (Anura, Hylidae), serra do sudeste, Rio Grande do Sul, Brasil. Biodiversidade Pampeana 6(2): 1-3.

Tonini, J.F.R.; Moraes Carao, L. de; Souza Pinto, I. de; Gasparini, J.L.; Reis Leite, Y.L.; Costa, L.P. 2010. Non-volant tetrapods from Reserva Biologica de Duas Bocas, State of Espirito Santo, southeastern Brazil. Biota Neotropica 10(3): 339-351.

Zocche, J.J.; Ceron, K.; Schmidt Gonzaga, G.; Plucenio, G.P.; Carvalho, F. 2018. Predation on Chironius bicarinatus (Wied, 1820) (Serpentes: Colubridae) by a Roadside Hawk Rupornis magnirostris (Gmelin, 1788) (Aves: Accipitridae). Herpetology Notes 11: 129-131.

Chironius brazili

Hamdan, B.; Fernandes, D.S. 2015. Taxonomic revision of Chironius flavolineatus (Jan, 1863) with description of a new species (Serpentes: Colubridae). Zootaxa 4012(1): 97–119.

Chironius carinatus

Alemán, C. 1952. Apuntes sobre reptiles y anfibios de la region Baruta-El Hatillo. Memoria de la Sociedad de Ciencias Naturales La Salle 12(31): 11-30.

Alemán, C. 1953. Contribution al estudio de los reptiles y batracios de la Sierra de Perija. Memoria de la Sociedad de Ciencias Naturales La Salle 13(35): Unpaginated.

Dunn, E.R. 1947. Snakes of the Lérida Farm (Chiriqui Volcano, western Panamá). Copeia 1947: 153-157.

Goeldi, E.A. 1897. Die Eier von 13 brasilianischen Reptilien, nebst Bemerkungen über Lebens- und Fortpflanzungsweise letzterer. Zoologische Jahrbücher (Syst.) 10: 640-676.

Goldberg, S.R. 2007. Note on reproduction of whipsnakes, genus Chironius (Serpentes: Colubridae), from Costa Rica. Bulletin of the Chicago Herpetological Society 42(9): 148-149.

Henderson, R.W.; Nickerson, M.A.; Ketcham, S. 1976. Short term movements of the snakes Chironius carinatus, Helicops angulatus and Bothrops atrox in Amazonian Peru. Herpetologica 32(3): 304-310.

Prado, A.; Hoge, A.R. 1947. Observações sobre serpentes do Peru, com a descrição de uma nova especie. Ciencia (Mexico) 8(6-9): 180.

Schmidt, K.P.; Walker, W.F. 1943. Peruvian snakes from the University of Arequipa. Field Museum of Natural History Zoological Series 24: 279-296.

Shreve, B. 1947. On Venezuelan reptiles and amphibians collected by Dr. G. Kugler. Bulletin of the Museum of Comparative Zoology at Harvard 99(5): 517-537.

Sousa, J.C.; Santos, F.P.; Sanches, P.R.; Costa-Campos, C.E.; Nunes, A.L.S. 2018. Natural history notes: Chironius carinatus (Amazon Whipsnake). Predation. Herpetological Review 49(2): 336.

Werner, F. 1910. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. Hamburg Jahrb. wiss. Anst. 26 [1908-1909]: 205-247.

Chironius challenger

Kok, P.J.R. 2010. A new species of Chironius Fitzinger, 1826 (Squamata: Colubridae) from the Pantepui region, northeastern South America. Zootaxa 2611: 31-44.

Chironius diamantina

Fernandes, D.S.; Hamdan, B. 2014. A new species of Chironius Fitzinger, 1826 from the state of Bahia, Northeastern Brazil (Serpentes: Colubridae). Zootaxa 3881(6): 563-575.

Chironius exoletus

Frota, J.G. da; Yuki, R.N. 2002. Geographic distribution: Chironius exoletus (Green Keeled Racer). Herpetological Review 33(1): 67.

Guimaraes, M.; Gaiarsa, M.P.; Cavalheri, H.B. 2014. Morphological adaptations to arboreal habitats and heart position in species of the Neotropical Whipsnakes genus Chironius. Acta Zoologica (Copenhagen) 95(3): 341-346.

Gusmao, A.C.; Casagrande, L.P.; Silva, A.M. da; Crispin, M.A.; Fereira-García, J.R.; Souza, M.R. de; Suzeck, E.; Bernarde, P.S. 2011. Natural history notes: Chironius exoletus (Vine Snake). Reproductive behavior. Herpetological Review 42(2): 284.

Köhler, J. 2010. Chironius exoletus (Linnaeus, 1758) (Fortsetzung von Seite 3). Elaphe 18(2): 74.

Lema, T. de; Ibarra Vieria, M.; Leitao de Araujo, M. 1984. Fauna reptiliana do norte da Grande Porto Alegre, Rio Grande do Sul, Brasil. Revista Brasileira de Zoologia 2(4): 203-227.

Mendes Castanho, L. 1996. Natural history notes: Phyllomedusa distincta (Leaf-frog). Predation. Herpetological Review 27(3): 141.

Rodrigues, M.G. 2008. Chironius exoletus (common whipsnake): prey and possible diet convergence. Herpetological Bulletin 105: 41-42.

Rojas-Runjaic, F.J.M.; Infante-Rivero, E.E. 2006. Reptilia, Squamata, Colubridae, Chironius exoletus: distribution extension, new state record. Check List 2(3): 82-83.

Starace, F. 1998. Contribution to the study of the snakes of French Guyana. III. The presence of Chironius exoletus (Linnaeus, 1758) in French Guyana. Litteratura Serpentium (English Edition) 18(1): 13-16.

Sudré, V.; Curcio, F.F.; Sales-Nunes, P.M.; Pellegrino, K.C.M.; Rodrigues, M.T. 2017. Who is the Red-bearded Snake, anyway? Clarifying the taxonomic status of Chironius pyrrhopogon (Wied, 1824) (Serpentes: Colubridae). Zootaxa 4319(1): 143–156.

Chironius flavolineatus

Albuquerque, S. de; Freitas, M.A. de 2011. Geographic distribution: Chironius flavolineatus (Liana Snake). Herpetological Review 42(4): 570-571.

Aximoff, I.; Cintra, H.; Pontes, J. 2017. Natural history notes: Chironius flavolineatus (Boettger's Sipo). Diet and habitat use. Herpetological Review 48(1): 205.

Borges-Nojosa, D.M.; Cassiano Lima, D. 2009. Geographic distribution: Chironius flavolineatus (cipo snake). Herpetological Review 40(2): 237.

Colli, G.R.; Peres, A.K. 1997. Geographic distribution: Chironius flavolineatus. Herpetological Review 28(3): 158.

Colli, G.R.; Peres, A.K. 1998. Geographic distribution: Chironius flavolineatus (Cobra-cipo, Boettger's Sipo). Herpetological Review 29(1): 53.

Espinoza, N.C. de 1977. New records of colubrid snakes from Peru, with notes on their ecology and distribution. Herpetological Review 8(3): 3.

Feio, R.N.; Santos, P.S.; Fernandes, R.; Freitas, T.S. de 1999. Natural history notes: Chironius flavolineatus (NCN). Courtship. Herpetological Review 30(2): 99.

Hamdan, B.; Fernandes, D.S. 2015. Taxonomic revision of Chironius flavolineatus (Jan, 1863) with description of a new species (Serpentes: Colubridae). Zootaxa 4012(1): 97–119.

Hamdan, B.; Scali, S.; Fernandes, D.S. 2014. On the identity of Chironius flavolineatus (Serpentes: Colubridae). Zootaxa 3794(1): 134-142.

Pinto, R.R.; Fernandes, R.; Marques, O.A.V. 2008. Morphology and diet of two sympatric colubrid snakes, Chironius flavolineatus and Chironius quadricarinatus (Serpentes: Colubridae). Amphibia-Reptilia 29(2): 149-160.

Pinto, R.R.; Marques, O.A.V.; Fernandes, R. 2010. Reproductive biology of two sympatric colubrid snakes, Chironius flavolineatus and Chironius quadricarinatus, from the Brazilian cerrado domain. Amphibia-Reptilia 31(4): 463-473.

Piratelli, A.J.; Pereira, M.R.; Cordeiro Siqueira, M.A. 1996. Predation of Chironius flavolineatus (Squamata: Colubridae) by Nystalus maculatus (Piciformes: Bucconidae). Ararajuba 4(2): 113.

Rufray, V.; Thonnel, A.; Cobigo, M. 2015. Découverte d’une nouvelle espèce de serpent en Guyane française: Chironius flavolineatus Jan, 1863. Bulletin de la Société Herpétologique de France 154: 71-73.

Sawaya, R.J.; Spina, F.; Monteiro, C.S.; Oliveira, J.L.; Brasileiro, C.A. 2003. Natural history notes: Chironius flavolineatus (NCN). Predation. Herpetological Review 34(4): 371-372.

Chironius flavopictus

Amaral, A. do 1929. Valor systematico de varias formas de ophidios neotropicos. Memorias do Instituto Butantan (Sao Paulo) 4: 3-68.

Saenger, M. von; Ray, J.M. 2016. Geographic distribution: Chironius flavopictus (Sipo Keelback). Herpetological Review 47(1): 83.

Werner, F. 1910. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. Hamburg Jahrb. wiss. Anst. 26 [1908-1909]: 205-247.

Chironius foveatus

Guimaraes, M.; Gaiarsa, M.P.; Cavalheri, H.B. 2014. Morphological adaptations to arboreal habitats and heart position in species of the Neotropical Whipsnakes genus Chironius. Acta Zoologica (Copenhagen) 95(3): 341-346.

Mallet-Rodrigues, F.; Noronha, M.L.M. de 2014. Natural history notes: Chironius foveatus (South American Sipo). Diet. Herpetological Review 45(1): 141-142.

Muscat, E.; Entiauspe-Neto, O.M. 2016. Natural history notes: Chironius foveatus (Atlantic Forest Sipo Snake). Defensive behavior. Herpetological Review 47(3): 475.

Rocha, C.F.D.; Vrcibradic, D.; Sluys, M. van 1999. Natural history notes: Chironius multiventris foveatus (NCN). Prey. Herpetological Review 30(2): 99-100.

Chironius fuscus

Amaral, A. do 1929. Valor systematico de varias formas de ophidios neotropicos. Memorias do Instituto Butantan (Sao Paulo) 4: 3-68.

Bringsøe, H. 1999. Herpetologi i Fransk Guyana. 1. Harmonika-positur hos snoge. [Herpetology in French Guiana. 1. The accordion posture of snakes]. (In Danish, English summary). Nordisk Herpetologisk Forening 42(5): 130-135.

Dunn, E.R. 1947. Snakes of the Lérida Farm (Chiriqui Volcano, western Panamá). Copeia 1947: 153-157.

Guimaraes, M.; Gaiarsa, M.P.; Cavalheri, H.B. 2014. Morphological adaptations to arboreal habitats and heart position in species of the Neotropical Whipsnakes genus Chironius. Acta Zoologica (Copenhagen) 95(3): 341-346.

Nascimento, L.P.; Siqueira, D.M.; Santos-Costa, M.C. dos 2013. Diet, reproduction, and sexual dimorphism in the Vine Snake, Chironius fuscus (Serpentes: Colubridae), from Brazilian Amazonia. South American Journal of Herpetology 8(3): 168-174.

Silva-Ferreira, D.S.; Sousa, J.C.; Costa-Campos, C.E. 2017. Natural history notes: Chironius fuscus (Brown Sipo). Diet. Herpetological Review 48(1): 206.

Souza Filho, G.A. de; Moura-Leite, J.C. de; Grochoski, M.E.; Abrahão-Morato, S.A. 2012. Chironius fuscus (Linnaeus, 1758) (Serpentes: Colubridae): distribution extension, new state record and variation in southern Brazil. Check List 8(6): 1315-1318.

Chironius grandisquamis

Amaral, A. do 1929. Valor systematico de varias formas de ophidios neotropicos. Memorias do Instituto Butantan (Sao Paulo) 4: 3-68.

Boulenger, G.A. 1915. Descriptions of a new Amphisbæna and a new snake discovered by Dr. H.G.F. Spurrell in southern Colombia. Proceedings of the Zoological Society of London 1915(4): 659-661.

Goldberg, S.R. 2007. Note on reproduction of whipsnakes, genus Chironius (Serpentes: Colubridae), from Costa Rica. Bulletin of the Chicago Herpetological Society 42(9): 148-149.

López Guevara, H.J.; Ubeda-Olivas, M.F.; Cruz, J.; Sunyer, J. 2015. Distribution notes: Chironius grandisquamis (Peters, 1868). Mesoamerican Herpetology 2(4): 543-544.

Moss, W.; Fitzgerald, K. 1987. Geographic distribution: Chironius grandisquamis. Herpetological Review 18(1): 20.

Chironius laevicollis

Costa, A.C.O.R. da; Hauzman, E.; Scartozzoni, R.R. 2005. Chironius laevicollis (pale-necked whipsnake): reproduction. Herpetological Bulletin 92: 26-27.

Espinoza, N.C. de 1977. New records of colubrid snakes from Peru, with notes on their ecology and distribution. Herpetological Review 8(3): 3.

Guimaraes, M.; Gaiarsa, M.P.; Cavalheri, H.B. 2014. Morphological adaptations to arboreal habitats and heart position in species of the Neotropical Whipsnakes genus Chironius. Acta Zoologica (Copenhagen) 95(3): 341-346.

Marques, O.A.V.; Puorto, G. 1996. Geographic distribution: Chironius laevicollis (Brazilian Cipo). Herpetological Review 27(4): 212.

Marques, O.A.V.; Sazima, I. 2003. Ontogenetic color changes may strengthen suggestion about systematic affinities between two species of Chironius (Serpentes, Colubridae). Phyllomedusa 2(1): 65-67.

Chironius maculoventris

Marra Santos, F.J.; Repenning, M.; Beier, C.; Funk Pontes, G.M. 2015. First record of Chironius maculoventris Dixon, Wiest & Cei, 1993 (Squamata: Serpentes: Colubridae) in Brazil. Herpetology Notes 8: 169-171.

Chironius monticola

Espinoza, N.C. de 1977. New records of colubrid snakes from Peru, with notes on their ecology and distribution. Herpetological Review 8(3): 3.

Maddock, S.T.; Tolhurst, B.; Brown, M.; Peck, M.; Villacis Pérez, E.; Noe Morales, J. 2011. Body bending behaviour: more widespread than previously thought? New reports from two snake species of Northwest Ecuador. Herpetology Notes 4: 79-81.

Chironius multiventris

Embert, D. 2002. Geographic distribution: Chironius multiventris multiventris (Cobra Cipo, Sacaiboia). Herpetological Review 33(3): 226.

Hoge, A.R. 1969. A new species of Chironius (Serpentes - Colubridae). Memorias do Instituto Butantan (Sao Paulo) 34: 93-95.

Klaczko, J.; Andrade Machado, F. de; Scrocchi, G.J.; Zaher, H. 2010. Taxonomic status of Chironius multiventris and Chironius cochranae (Serpentes). Herpetologica 66(4): 476-484.

Schmidt, K.P.; Walker, W.F. 1943. Peruvian snakes from the University of Arequipa. Field Museum of Natural History Zoological Series 24: 279-296.

Chironius quadricarinatus

Argôlo, A.J.S. 1998. Geographic distribution: Chironius quadricarinatus quadricarinatus. Herpetological Review 29(3): 175.

Embert, D. 2002. Geographic distribution: Chironius quadricarinatus maculoventris (Cobra Cipo, Sacaiboia). Herpetological Review 33(3): 226-227.

Freitas, M.A. de; Silva, T.F.S.; Rodrigues, M.T. 2007. Geographic distribution: Chironius quadricarinatus. Brazil: Bahia. Herpetological Review 38(3): 354.

Pinto, R.R.; Fernandes, R.; Marques, O.A.V. 2008. Morphology and diet of two sympatric colubrid snakes, Chironius flavolineatus and Chironius quadricarinatus (Serpentes: Colubridae). Amphibia-Reptilia 29(2): 149-160.

Pinto, R.R.; Marques, O.A.V.; Fernandes, R. 2010. Reproductive biology of two sympatric colubrid snakes, Chironius flavolineatus and Chironius quadricarinatus, from the Brazilian cerrado domain. Amphibia-Reptilia 31(4): 463-473.

Chironius scurrulus

Fuenmayor, G.R.; Marca, E. la 2001. Geographic distribution: Chironius scurrulus (Machete). Herpetological Review 32(1): 58-59.

Hoge, A.R. 1969. Chironius scurulus (Wagler) recorded from Venezuela. Memorias do Instituto Butantan (Sao Paulo) 34: 85.

Hoge, A.R.; Nina, A.C. de M. 1964. Serpentes coietadas pelo Instituto Nacional de Pesquisas da Amazonia. Memorias do Instituto Butantan (Sao Paulo) 30 [1960-1962]: 71-96.

Hoge, A.R.; Santos, N.P.; Heitor, S.; Lopes, L.A.; Souza, I.M. de 1973. Serpentes coletadas pelo Projeto Rondon VII em Iauareté, Brasil. Memorias do Instituto Butantan (Sao Paulo) 36 [1973]: 221-232.

Marques, O.A.V.; Sazima, I. 2003. Ontogenetic color changes may strengthen suggestion about systematic affinities between two species of Chironius (Serpentes, Colubridae). Phyllomedusa 2(1): 65-67.

Santos, R.M. dos 2003. Contribuiçao ao conhecimento morfologico de Chironius scurrulus (Wagler, 1824) (Serpentes: Colubridae). Boletim do Museu Paraense Emilio Goeldi (Zoologia) 18(1): 35-42.

Chironius septentrionalis

Grundler, M.C.; Fitzpatrick, S.W. 2018. Natural history notes: Chironius septentrionalis (South American Sipo). Diet. Herpetological Review 49(3): 544.

Chironius vincenti

Henderson, R.W.; Haas, G.T. 1993. Status of the West Indian snake Chironius vincenti. Oryx 27(3): 181-184.

Henderson, R.W.; Powell, R. 1996. Chironius vincenti. Catalogue of American Amphibians and Reptiles 635: 1-2.

Henderson, R.W.; Sajdak, R.A.; Henderson, R.M. 1988. The rediscovery of the West Indian colubrid snake Chironius vincenti. Amphibia-Reptilia 9(4): 415-416.