Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk). Latest update: December 31st, 2022.
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Genus
Hemorrhois
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Palearctic Whip Snakes
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Westpaläarktische Zornnattern
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Vestpalæarktiske Pilsnoge
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1826 | |
Hemorrhois Boie (type species: Coluber hippocrepis Linnaeus 1758; syn. Williams & Wallach 1989) |
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Haemorrhois Boie 1827 [unjustified emendation] |
1830 | |
Periops Wagler (type species: Coluber hippocrepis Linnaeus 1758) |
1843 | |
Chilolepis Fitzinger (type species: Coluber cliffordi Schlegel 1837) |
Contents:
4 species, none of which are endemic.
Endemism: 0% 100%
Remarks:
Previously regarded as a synonym of
Coluber (e.g., Mertens & Wermuth 1960). Often misspelled Haemorrhois.
Distribution:
N. Africa, S. Europe, Middle East, SW. and C. Asia.
Reported from:
Afghanistan,
Algeria,
Armenia,
Azerbaijan,
China
(Xinjiang),
Cyprus,
Egypt
(incl.
Sinai),
Georgia,
Gibraltar,
Greece
(Aegean Islands
[incl.
Dodecanese Islands
(incl.
Rhodes)]),
Iran,
Iraq,
Israel,
Italy
(Pantelleria),
Jordan,
Kazakhstan,
Kyrgyzstan,
Lebanon,
Libya,
Mauritania,
Morocco,
Pakistan,
Portugal,
Russia,
Spain,
Syria,
Tajikistan,
Tunisia,
Turkey,
Turkmenistan,
Uzbekistan,
Western Sahara.
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Hemorrhois algirus
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Algerian Whip Snake, Algerian Grass Snake
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Algerische Zornnatter
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Algiersk Pilsnog
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1863 | |
Periops algira Jan |
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Zamenis algirus Boulenger 1891 |
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Coluber algirus Villiers 1950 |
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Coluber florulentus algirus Kramer & Schnurrenberger 1959 |
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Haemorrhois algirus [sic] Welch 1980 |
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Eremiophis algirus Welch 1983 |
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Hemorrhois algirus Schätti & Utiger 2001 |
1929 | |
Coluber algirus intermedius Werner |
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Coluber florulentus intermedius Pasteur & Bons 1960 |
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Coluber hippocrepis intermedius Bons 1962 |
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Hemorrhois algirus intermedius Baha el Din 2006 |
1962 | |
Coluber algirus villiersi Bons (Schätti 1986) |
Remarks:
Chippaux & Jackson (2019) included Niger in the distribution, but no localities in the country were shown in their map.
No other references have been found to support the presence of the species in niger.
Introduced to:
Malta.
Distribution:
Algeria,
Egypt,
Libya,
Mauritania,
Morocco,
Tunisia,
Western Sahara.
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Hemorrhois hippocrepis
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Horseshoe Whip Snake, Horseshoe Snake
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Hufeisen-Natter
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Hesteskosnog
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1758 | |
Coluber hippocrepis Linnaeus |
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Periops hippocrepis Wagler 1830 |
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Zamenis hippocrepis Günther 1858 |
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Tyria hippocrepis Cope 1862 |
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Haemorrhois hippocrepis [sic] Welch 1980 |
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Hemorrhois hippocrepis Welch 1983 |
1766 | |
Coluber domesticus Linnaeus (Böhme (ed.) 1993) |
1824 | |
Natrix bahiensis Wagler (Böhme (ed.) 1993) |
1834 | |
Coluber diadema Bonelli (Böhme (ed.) 1993) |
1985 | |
Coluber hippocrepis nigrescens Cattaneo (Böhme (ed.) 1993) |
Remarks:
Faraone, Melfi, Nicola, Giacalone & Valvo (2020) showed that the Italian populations (Pantelleria, Sardinia) are most closely related to Tunisian and East Algerian populations, and their studies
supported the hypothesis of a recent origin of the Italian populations, probably determined by human-mediated dispersal from North Africa.
Introduced to:
Italy
(Pantelleria,
Sardinia),
Spain
(Balearic Islands
[Mallorca,
Pityusic Islands
(Ibiza)]).
Distribution:
Algeria,
Gibraltar,
Morocco,
Portugal,
Spain,
Tunisia.
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Hemorrhois hippocrepis
© Henrik Bringsøe
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Hemorrhois nummifer
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Coin-marked Whip Snake, Coin-marked Snake, Coin Snake, Leaden-coloured Racer, Lead Wood Snake, (Asian Racer, Reuss' Whip Snake)
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Münzennatter
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Mønt-pilsnog
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1758 | |
Coluber tyria Linnaeus [suppressed] (ICZN (2011): Opinion 2282) |
1834 | |
Coluber nummifer Reuss |
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Zamenis ravergieri nummifer Boettger 1880 |
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Elaphe nummifera Schmidt 1939 |
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Coluber ravergieri nummifer Werner 1971 |
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Hemorrhois nummifer Schätti & Utiger 2001 |
1858 | |
Zamenis caudaelineatus Günther [part] (Schätti & Agasian 1985) |
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Elaphe caudaelineata Schmidt 1939 |
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Haemorrhois caudaelineata [sic] Welch 1983 |
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Eremiophis caudaelineata Welch 1983 |
1863 | |
Periops neglectus Jan (Schätti & Agasian 1985) |
1917 | |
Zamenis ater Venzmer [not Zamenis ater Günther 1872] (Schätti & Agasian 1985) |
1926 | |
Elaphe dione plumbea Cernov (Schätti & Agasian 1985) |
1929 | |
Zamenis venzmeri Werner [substitute name for Zamenis ater Venzmer 1917] (Schätti & Agasian 1985) |
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Coluber venzmeri Bodenheimer 1944 |
1936 | |
Coluber nummifer pallaryi Angel (Schätti & Agasian 1985) |
1944 | |
Elaphe quatuorlineata sauromates bahtiyari Bodenheimer (Schätti & Agasian 1985; Böhme (ed.) 1993) |
Remarks:
Coluber tyria Linnaeus 1758, a name often used previously for Spalerosophis diadema, is most likely a senior subjective synonym of nummifer (Schätti, Tillack & Kucharzewski 2009). It has now been suppressed (see above).
Distribution:
Armenia,
Azerbaijan,
Cyprus,
Egypt
(incl.
Sinai),
Greece
(Aegean Islands
[incl.
Dodecanese Islands
(incl.
Rhodes)]),
Iran,
Iraq,
Israel,
Jordan,
Kazakhstan,
Kyrgyzstan,
Lebanon,
Syria,
Tajikistan,
Turkey,
Turkmenistan,
Uzbekistan.
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Hemorrhois nummifer
© Henrik Bringsøe
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Hemorrhois ravergieri
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Mountain Whip Snake, Variegated Whip Snake, Versicolored Wood Snake, (Spotted Whip Snake, Ravergier's Whip Snake)
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Bunte Zornnatter
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Bjergpilsnog
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1832 | |
Coluber ravergieri Menetries |
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Zamenis ravergieri Boulenger 1893 |
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Haemorrhois ravergieri [sic] Welch 1983 |
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Eremiophis ravergieri Welch 1983 |
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Hemorrhois ravergieri Schätti & Utiger 2001 |
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Platyceps ravergieri Bosch & Bischoff 2004 |
1832 | |
Coluber maculatus Dwigubskij (Böhme (ed.) 1993) |
1838 | |
Coluber cliffordi Martin [not Coluber cliffordi Schlegel 1837] (Böhme (ed.) 1993) |
1858 | |
Zamenis caudaelineatus Günther [part] (Schätti & Agasian 1985) |
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Elaphe caudaelineata Schmidt 1939 |
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Haemorrhois caudaelineata [sic] Welch 1983 |
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Eremiophis caudaelineata Welch 1983 |
1865 | |
Periops caudolineatus nera Jan (Böhme (ed.) 1993) |
1873 | |
Zamenis fedtschenkoi Strauch (Böhme (ed.) 1993) |
1885 | |
Zamenis kaufmanni Grum-Grzimailo (Böhme (ed.) 1993) |
1896 | |
Zamenis glazunovi Nikolski (Böhme (ed.) 1993) |
1952 | |
Coluber ravergieri cernovi Mertens [substitute name for Elaphe dione plumbea Cernov 1926] (Schätti & Agasian 1985) |
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Haemorrhois ravergieri cernovi [sic] Welch 1983 |
Other common names:
cernovi: Turkish Whip Snake
Remarks:
Previous reports from Iraq (e.g., Khalaf 1959, Leviton & Anderson 1970; repeated by Venchi & Sindaco 2007) are based on confusion with nummifer (Schätti & Agasian 1985). Records from Egypt (e.g., Flower 1933; Schmidt & Marx 1956) refer to nummifer (Baha el Din 2006). Reported from India by some sources, however, it is not considered an Indian species (Whitaker & Captain 2004).
Distribution:
Afghanistan,
Armenia,
Azerbaijan,
China
(Xinjiang),
Georgia,
Iran,
Iraq,
Israel,
Jordan,
Kazakhstan,
Lebanon,
Pakistan,
Russia,
Syria,
Tajikistan,
Turkey,
Turkmenistan,
Uzbekistan.
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Hemorrhois ravergieri
© Henrik Bringsøe
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