Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus Recent updates
Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update: February 11th, 2024.


Taxonomy of the family Colubridae
Bibliography of the genus Fowlea
Biodiversity of the family Colubridae








Genus
Fowlea

Fishing Keelbacks, (Oriental Painted Keelbacks, Striped Water Snakes)

Fischnattern

Fiskesnoge

1868 Fowlea Theobald (type species: Fowlea peguensis Theobald 1868)
1893 Diplophallus Cope (type species: Hydrus piscator Schneider 1799)
Contents: 10 species, of which 5 (50.0%) are endemic.
Endemism: 0% 100%
Remarks: Previously included in Natrix (e.g., Smith 1943) and, more recently, Xenochrophis (e.g., Williams & Wallach 1989). Revalidated as a separate genus by Purkayastha, Kalita, Brahma, Doley & Das (2018). According to Cheng, Ren, Zhu & Li (2021), the gender of Fowlea is masculine, however, The Reptile Database (see Fowlea punctata, viewed January 1st, 2023) pointed out that in accordance with Article 30.2.4 of the Code (ICZN 1999), it is to be treated as feminine.
Distribution: S. and E. Asia.
Reported from: Afghanistan, Bangladesh, Bhutan, Cambodia, China (Anhui, Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangsu, Jiangxi, Tibet, Yunnan, Zhejiang), India (Andaman & Nicobar Islands [Andaman Islands], Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Telangana, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Indonesia (Bali, Java, Sumatra), Laos, Malaysia (West Malaysia), Myanmar, Nepal, Pakistan, Sri Lanka, Taiwan, Thailand, Vietnam (incl. Cat Ba Island).

Fowlea asperrima

Sri Lankan Painted Keelback

Srilankanische Fischnatter

Srilankansk Fiskesnog

1891 Tropidonotus asperrimus Boulenger
Natrix piscator asperrimus Smith 1943
Xenochrophis asperrimus Welch 1988
Fowlea asperrimus Purkayastha, Kalita, Brahma, Doley & Das 2018

Remarks: Validity as a distinct species confirmed by Vogel & David (2006).
Distribution: Sri Lanka.


Fowlea flavipunctata

Yellow-spotted Keelback, Common Keelback

Gelbfleckiger Fischnatter

Gulplettet Fiskesnog

1860 Amphiesma flavipunctatum Hallowell
Natrix piscator flavipunctata Smith 1943
Natrix flavipunctata Taylor 1965
Xenochrophis flavipunctatus Welch 1988
Fowlea flavipunctatus Purkayastha, Kalita, Brahma, Doley & Das 2018
1907 Tropidonotus piscator ornata Wall (Vogel & David 2012)

Remarks: Previously regarded as a synonym of piscator. Taylor (1965) separated the two species, based on sympatry in Thailand. Validity confirmed by Vogel & David (2006). It is still not clear, however, whether one or both species are present in parts of the range of piscator (in the strict sense) (see for instance Zhao & Adler 1993). A record from Assam (India) (Smith 1943) refer to schnurrenbergeri (Vogel & David 2006). Listed for India by Aengals, Kumar, Palot & Ganesh (2018), but without discussion or references. Das & Yaakob (2007) included Sarawak in the distribution of flavipunctata, but the species was not regarded as a Bornean species by Stuebing, Inger & Lardner (2014), which is followed herein. A record of piscator from Shaanxi (Song 2002), presumably referring to flavipunctata, needs confirmation.
Introduced to: Singapore.
Distribution: Cambodia, China (Anhui, Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangsu, Jiangxi, Tibet, Yunnan, Zhejiang), Laos, Malaysia (West Malaysia), Myanmar, Taiwan, Thailand, Vietnam (incl. Cat Ba Island).


Fowlea flavipunctata
© Rune Midtgaard

Fowlea melanzosta

Southern Keelback, Javan Keelback

Südliche Fischnatter

Sydlig Fiskesnog

1807 Coluber Melanzostus Gravenhorst
Tropidonotus melanozostus Boie 1827 [unjustified emendation]
Tropidonotus quincunciatus melanozostus Müller 1890
Tropidonotus piscator melanozosta Boettger 1892
Natrix piscator melanzostus Smith 1943
Xenochrophis piscator melanzostus Bosch 1985
Xenochrophis flavipunctatus melanozostus Manthey & Grossmann 1997
Xenochrophis melanzostus Das 1999
Fowlea melanzostus Purkayastha, Kalita, Brahma, Doley & Das 2018
1834 Coluber lippus Reuss (Vogel & David 2012)

Remarks: Validity as a distinct species confirmed by Vogel & David (2006). Reports from the Andaman Islands (India) refer to tytleri (Vogel & David 2006). Presence in Sulawesi needs confirmation, since only a few voucher specimens are available from the island (Lang & Vogel 2005). Reports from Borneo are erroneous (Vogel & David 2006).
Distribution: Indonesia (Bali, Java, Sumatra).


Fowlea piscator

Indian Checkered Keelback, Common Pond Snake, Asian Fishing Snake, Fish Snake, (Asian Water Snake)

Indische Schachbrett-Fischnatter

Indisk Fiskesnog

1799 Hydrus piscator Schneider
Tropidonotus piscator Boie 1827
Nerodia piscator Wall 1923
Natrix piscator Smith 1930
Fowlea piscator Malnate 1960
Xenochrophis piscator Malnate & Minton 1965
1799 Hydrus palustris Schneider (Vogel & David 2012)
1803 Coluber anostomosatus Daudin (Vogel & David 2012)
1803 Coluber braminus Daudin (Vogel & David 2012)
1803 Coluber dora Daudin (Vogel & David 2012)
1803 Coluber mortuarius Daudin (Vogel & David 2012)
1803 Coluber umbratus Daudin (Vogel & David 2012)
1834 Coluber bengalensis Gray (Vogel & David 2012)
1834 Coluber rectangulus Gray (Vogel & David 2012)
1837 Tropidonotus quincunciatus Schlegel (Vogel & David 2012)
1868 Tropidonotus striolatus Blyth (Vogel & David 2012)
1907 Tropidonotus piscator lateralis Wall (Vogel & David 2012)
1907 Tropidonotus piscator punctatus Wall (Vogel & David 2012)

Remarks: Previously included asperrima, flavipunctata, melanzosta, sanctijohannis, tytleri, and unicolor. Records of piscator from Sri Lanka refer to unicolor. Records from Cambodia, Singapore, Vietnam, West Malaysia, some records from Laos, and most Chinese records refer to flavipunctata. Records from Indonesia refer to melanzosta. Records from the Andaman Islands refer to tytleri. A record from Shaanxi (Song 2002), presumably referring to flavipunctata, needs confirmation.
Introduced to: Singapore.
Distribution: Afghanistan, Bangladesh, Bhutan, China (Guangxi, Yunnan), India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Telangana, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Laos, Myanmar, Nepal, Pakistan, Thailand.


Fowlea piscator
© Henrik Bringsøe

Fowlea punctulata

Burmese Spotted Keelback, (Spotted Keelback)

Burmesische Gefleckte Fischnatter

Burmesisk Plettet Fiskesnog

1858 Tropidonotus punctulatus Günther
Nerodia punctulata Wall 1923
Natrix punctulata Smith 1943
Fowlea punctulata Malnate 1960
Xenochrophis punctulatus Welch 1988
1868 Fowlea peguensis Theobald (Smith 1943)

Remarks: Records from India (e.g., Borang, Bhatt, Chaudhury, Borkotoki & Bhutia 2005; Sanyal & Gayen 2006) appear to be undocumented.
Distribution: Myanmar, Thailand.


Fowlea sanctijohannis

Kashmir Keelback, (St. John's Keelback, Olive Water Snake)

Sankt-Johannis-Fischnatter

Kashmir-fiskesnog

1890 Tropidonotus sancti-johannis Boulenger
Nerodia piscator sancti-johannis Bourret 1935
Natrix piscator sancti-johannis Bourret 1936
Natrix sancti-johannis Khan 1984
Xenochrophis piscator sanctijohannis Kramer 1977
Xenochrophis sanctijohannis Welch 1988
Fowlea sanctijohannis Purkayastha, Kalita, Brahma, Doley & Das 2018
1907 Tropidonotus piscator obscurus Wall (Vogel & David 2012)

Remarks: Previously regarded as a synonym of piscator (e.g., Wall 1923; Pope 1923; Smith 1943), revalidated as a separate species by Khan (1984), but subsequently considered a montane colour morph of, and placed in the synonymy of, piscator (Vogel & David 2012). Nevertheless, several authors (e.g.; Das & Das 2017; Purkayastha, Kalita, Brahma, Doley & Das 2018; Ghosh & Das 2021; Wangyal & Das 2021) continue to list sanctijohannis as a valid species. Pending further studies, it is retained herein as a separate species.
Distribution: Afghanistan, India (Arunachal Pradesh, Himachal Pradesh, Jammu & Kashmir, Uttar Pradesh, West Bengal [incl. Darjeeling]), Myanmar, Nepal, Pakistan.





Fowlea schnurrenbergeri

Bar-necked Keelback, (Kramer's Keelback)

Nackenband-Fischnatter

Nordvestlig Fiskesnog

1977 Xenochrophis flavipunctatus schnurrenbergeri Kramer
Xenochrophis schnurrenbergeri Vogel & David 2006
Fowlea schnurrenbergeri Purkayastha, Kalita, Brahma, Doley & Das 2018

Distribution: India (Assam, Bihar, Odisha, Uttar Pradesh, West Bengal), Nepal, Pakistan.


Fowlea tytleri

Andaman Keelback

Andamanische Fischnatter

Andamansk Fiskesnog

1863 Tropidonotus tytleri Blyth
Xenochrophis tytleri Vogel & David 2006
Fowlea tytleri Purkayastha, Kalita, Brahma, Doley & Das 2018

Remarks: Köhler, Than & Thammachoti (2021) synonymized tytleri with piscator, however, this was rejected by Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel (2022). Although reported from the Nicobar Islands (as piscator or tytleri) (e.g., Biswas 1984), Vogel & David (2012) regarded the presence in these islands as doubtful.
Distribution: India (Andaman & Nicobar Islands: Andaman Islands)


Fowlea unicolor

Sri Lankan Checkered Keelback

Srilankanische Schachbrett-Fischnatter

Srilankansk Fiskesnog

1887 Tropidonotus quincunciatus var. unicolor Müller
Fowlea unicolor Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel 2022

Remarks: Previously regarded as a synonym of piscator or sanctijohannis (Vogel & David 2012). Revalidated as a separate species by Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel (2022). Records of piscator from Sri Lanka refer to unicolor. The species is probably also present in S. India, but this needs verification (Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel 2022).
Distribution: Sri Lanka.





Fowlea yunnanensis

Yunnan Olive Keelback

Chinesische Fischnatter, Chinesische Olivfarbige Kielnatter

Kinesisk Fiskesnog, Kinesisk Olivenkølsnog

1879 Atretium schistosum yunnanensis Anderson
Helicops schistosus yunnanensis Annandale 1911
Helicops yunnanensis Pope 1935
Atretium yunnanensis Smith 1943
Fowlea yunnanensis Cheng, Ren, Zhu & Li 2021
1909 Helicops schistosus andersonii Wall (Zhao & Adler 1993)

Remarks: Previously assigned to the genus Atretium. Transferred to Fowlea by Cheng, Ren, Zhu & Li (2021) and Deepak, Cooper, Poyarkov, Kraus, Burin, Das, Narayanan, Streicher, Smith & Gower (2022).
Distribution: China (Yunnan).