Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update: February 11th, 2024.
 
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Genus
Fowlea
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Fishing Keelbacks, (Oriental Painted Keelbacks, Striped Water Snakes)
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Fischnattern
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Fiskesnoge
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1868 |  |
Fowlea Theobald (type species: Fowlea peguensis Theobald 1868) |
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1893 | |
Diplophallus Cope (type species: Hydrus piscator Schneider 1799) |
Contents:
10 species, of which 5 (50.0%) are endemic.
Endemism: 0%  100%
Remarks:
Previously included in
Natrix
(e.g., Smith 1943) and, more recently,
Xenochrophis
(e.g., Williams & Wallach 1989).
Revalidated as a separate genus by Purkayastha, Kalita, Brahma, Doley & Das (2018).
According to Cheng, Ren, Zhu & Li (2021), the gender of Fowlea is masculine, however, The Reptile Database (see Fowlea punctata, viewed January 1st, 2023) pointed out that
in accordance with Article 30.2.4 of the Code (ICZN 1999), it is to be treated as feminine.
Distribution:
S. and E. Asia.
Reported from:
Afghanistan,
Bangladesh,
Bhutan,
Cambodia,
China
(Anhui,
Fujian,
Guangdong
[incl.
Nan Ao Island],
Guangxi,
Guizhou,
Hainan,
Hong Kong,
Hunan,
Jiangsu,
Jiangxi,
Tibet,
Yunnan,
Zhejiang),
India
(Andaman & Nicobar Islands
[Andaman Islands],
Andhra Pradesh,
Arunachal Pradesh,
Assam,
Bihar,
Chhattisgarh,
Delhi,
Goa,
Gujarat,
Himachal Pradesh,
Jammu & Kashmir,
Jharkhand,
Karnataka,
Kerala,
Madhya Pradesh,
Maharashtra,
Manipur,
Meghalaya,
Mizoram,
Nagaland,
Odisha,
Punjab,
Rajasthan,
Sikkim,
Tamil Nadu,
Telangana,
Tripura,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Indonesia
(Bali,
Java,
Sumatra),
Laos,
Malaysia
(West Malaysia),
Myanmar,
Nepal,
Pakistan,
Sri Lanka,
Taiwan,
Thailand,
Vietnam
(incl.
Cat Ba Island).
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Fowlea asperrima
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Sri Lankan Painted Keelback
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Srilankanische Fischnatter
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Srilankansk Fiskesnog
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1891 | |
Tropidonotus asperrimus Boulenger |
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Natrix piscator asperrimus Smith 1943 |
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Xenochrophis asperrimus Welch 1988 |
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Fowlea asperrimus Purkayastha, Kalita, Brahma, Doley & Das 2018 |
Remarks:
Validity as a distinct species confirmed by Vogel & David (2006).
Distribution:
Sri Lanka.
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Fowlea flavipunctata
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Yellow-spotted Keelback, Common Keelback
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Gelbfleckiger Fischnatter
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Gulplettet Fiskesnog
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1860 | |
Amphiesma flavipunctatum Hallowell |
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Natrix piscator flavipunctata Smith 1943 |
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Natrix flavipunctata Taylor 1965 |
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Xenochrophis flavipunctatus Welch 1988 |
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Fowlea flavipunctatus Purkayastha, Kalita, Brahma, Doley & Das 2018 |
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1907 | |
Tropidonotus piscator ornata Wall (Vogel & David 2012) |
Remarks:
Previously regarded as a synonym of piscator.
Taylor (1965) separated the two species, based on sympatry in Thailand.
Validity confirmed by Vogel & David (2006).
It is still not clear, however, whether one or both species are present in parts of the range of piscator (in the strict sense)
(see for instance Zhao & Adler 1993).
A record from Assam (India) (Smith 1943) refer to schnurrenbergeri (Vogel & David 2006).
Listed for India by Aengals, Kumar, Palot & Ganesh (2018), but without discussion or references.
Das & Yaakob (2007) included Sarawak in the distribution of flavipunctata, but the species was not regarded as a Bornean species by
Stuebing, Inger & Lardner (2014), which is followed herein.
A record of piscator from Shaanxi (Song 2002), presumably referring to flavipunctata, needs confirmation.
Introduced to:
Singapore.
Distribution:
Cambodia,
China
(Anhui,
Fujian,
Guangdong
[incl.
Nan Ao Island],
Guangxi,
Guizhou,
Hainan,
Hong Kong,
Hunan,
Jiangsu,
Jiangxi,
Tibet,
Yunnan,
Zhejiang),
Laos,
Malaysia
(West Malaysia),
Myanmar,
Taiwan,
Thailand,
Vietnam
(incl.
Cat Ba Island).
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Fowlea flavipunctata
© Rune Midtgaard
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Fowlea melanzosta
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Southern Keelback, Javan Keelback
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Südliche Fischnatter
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Sydlig Fiskesnog
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1807 | |
Coluber Melanzostus Gravenhorst |
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Tropidonotus melanozostus Boie 1827 [unjustified emendation] |
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Tropidonotus quincunciatus melanozostus Müller 1890 |
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Tropidonotus piscator melanozosta Boettger 1892 |
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Natrix piscator melanzostus Smith 1943 |
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Xenochrophis piscator melanzostus Bosch 1985 |
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Xenochrophis flavipunctatus melanozostus Manthey & Grossmann 1997 |
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Xenochrophis melanzostus Das 1999 |
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Fowlea melanzostus Purkayastha, Kalita, Brahma, Doley & Das 2018 |
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1834 | |
Coluber lippus Reuss (Vogel & David 2012) |
Remarks:
Validity as a distinct species confirmed by Vogel & David (2006).
Reports from the Andaman Islands (India) refer to tytleri (Vogel & David 2006).
Presence in Sulawesi needs confirmation, since only a few voucher specimens are available from the island (Lang & Vogel 2005).
Reports from Borneo are erroneous (Vogel & David 2006).
Distribution:
Indonesia
(Bali,
Java,
Sumatra).
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Fowlea piscator
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Indian Checkered Keelback, Common Pond Snake, Asian Fishing Snake, Fish Snake, (Asian Water Snake)
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Indische Schachbrett-Fischnatter
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Indisk Fiskesnog
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1799 | |
Hydrus piscator Schneider |
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Tropidonotus piscator Boie 1827 |
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Nerodia piscator Wall 1923 |
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Natrix piscator Smith 1930 |
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Fowlea piscator Malnate 1960 |
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Xenochrophis piscator Malnate & Minton 1965 |
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1799 | |
Hydrus palustris Schneider (Vogel & David 2012) |
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1803 | |
Coluber anostomosatus Daudin (Vogel & David 2012) |
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1803 | |
Coluber braminus Daudin (Vogel & David 2012) |
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1803 | |
Coluber dora Daudin (Vogel & David 2012) |
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1803 | |
Coluber mortuarius Daudin (Vogel & David 2012) |
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1803 | |
Coluber umbratus Daudin (Vogel & David 2012) |
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1834 | |
Coluber bengalensis Gray (Vogel & David 2012) |
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1834 | |
Coluber rectangulus Gray (Vogel & David 2012) |
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1837 | |
Tropidonotus quincunciatus Schlegel (Vogel & David 2012) |
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1868 | |
Tropidonotus striolatus Blyth (Vogel & David 2012) |
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1907 | |
Tropidonotus piscator lateralis Wall (Vogel & David 2012) |
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1907 | |
Tropidonotus piscator punctatus Wall (Vogel & David 2012) |
Remarks:
Previously included asperrima, flavipunctata, melanzosta, sanctijohannis, tytleri, and unicolor.
Records of piscator from Sri Lanka refer to unicolor.
Records from Cambodia, Singapore, Vietnam, West Malaysia, some records from Laos, and most Chinese records refer to flavipunctata.
Records from Indonesia refer to melanzosta.
Records from the Andaman Islands refer to tytleri.
A record from Shaanxi (Song 2002), presumably referring to flavipunctata, needs confirmation.
Introduced to:
Singapore.
Distribution:
Afghanistan,
Bangladesh,
Bhutan,
China
(Guangxi,
Yunnan),
India
(Andhra Pradesh,
Arunachal Pradesh,
Assam,
Bihar,
Chhattisgarh,
Delhi,
Goa,
Gujarat,
Himachal Pradesh,
Jammu & Kashmir,
Jharkhand,
Karnataka,
Kerala,
Madhya Pradesh,
Maharashtra,
Manipur,
Meghalaya,
Mizoram,
Nagaland,
Odisha,
Punjab,
Rajasthan,
Sikkim,
Tamil Nadu,
Telangana,
Tripura,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Laos,
Myanmar,
Nepal,
Pakistan,
Thailand.
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Fowlea piscator
© Henrik Bringsøe
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Fowlea punctulata
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Burmese Spotted Keelback, (Spotted Keelback)
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Burmesische Gefleckte Fischnatter
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Burmesisk Plettet Fiskesnog
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1858 | |
Tropidonotus punctulatus Günther |
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Nerodia punctulata Wall 1923 |
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Natrix punctulata Smith 1943 |
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Fowlea punctulata Malnate 1960 |
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Xenochrophis punctulatus Welch 1988 |
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1868 | |
Fowlea peguensis Theobald (Smith 1943) |
Remarks:
Records from India (e.g., Borang, Bhatt, Chaudhury, Borkotoki & Bhutia 2005; Sanyal & Gayen 2006) appear to be undocumented.
Distribution:
Myanmar,
Thailand.
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Fowlea sanctijohannis
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Kashmir Keelback, (St. John's Keelback, Olive Water Snake)
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Sankt-Johannis-Fischnatter
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Kashmir-fiskesnog
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1890 | |
Tropidonotus sancti-johannis Boulenger |
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Nerodia piscator sancti-johannis Bourret 1935 |
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Natrix piscator sancti-johannis Bourret 1936 |
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Natrix sancti-johannis Khan 1984 |
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Xenochrophis piscator sanctijohannis Kramer 1977 |
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Xenochrophis sanctijohannis Welch 1988 |
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Fowlea sanctijohannis Purkayastha, Kalita, Brahma, Doley & Das 2018 |
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1907 | |
Tropidonotus piscator obscurus Wall (Vogel & David 2012) |
Remarks:
Previously regarded as a synonym of piscator (e.g., Wall 1923; Pope 1923; Smith 1943), revalidated as a separate species by
Khan (1984), but subsequently considered a montane colour morph of, and placed in the synonymy of, piscator (Vogel & David 2012).
Nevertheless, several authors (e.g.; Das & Das 2017; Purkayastha, Kalita, Brahma, Doley & Das 2018; Ghosh & Das 2021; Wangyal & Das 2021) continue to list sanctijohannis as a valid species.
Pending further studies, it is retained herein as a separate species.
Distribution:
Afghanistan,
India
(Arunachal Pradesh,
Himachal Pradesh,
Jammu & Kashmir,
Uttar Pradesh,
West Bengal
[incl.
Darjeeling]),
Myanmar,
Nepal,
Pakistan.
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Fowlea schnurrenbergeri
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Bar-necked Keelback, (Kramer's Keelback)
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Nackenband-Fischnatter
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Nordvestlig Fiskesnog
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1977 | |
Xenochrophis flavipunctatus schnurrenbergeri Kramer |
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Xenochrophis schnurrenbergeri Vogel & David 2006 |
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Fowlea schnurrenbergeri Purkayastha, Kalita, Brahma, Doley & Das 2018 |
Distribution:
India
(Assam,
Bihar,
Odisha,
Uttar Pradesh,
West Bengal),
Nepal,
Pakistan.
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Fowlea tytleri
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Andaman Keelback
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Andamanische Fischnatter
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Andamansk Fiskesnog
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1863 | |
Tropidonotus tytleri Blyth |
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Xenochrophis tytleri Vogel & David 2006 |
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Fowlea tytleri Purkayastha, Kalita, Brahma, Doley & Das 2018 |
Remarks:
Köhler, Than & Thammachoti (2021) synonymized tytleri with piscator, however, this was rejected by Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel (2022).
Although reported from the Nicobar Islands (as piscator or tytleri) (e.g., Biswas 1984), Vogel & David (2012) regarded the presence in these islands as doubtful.
Distribution:
India
(Andaman & Nicobar Islands:
Andaman Islands)
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Fowlea unicolor
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Sri Lankan Checkered Keelback
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Srilankanische Schachbrett-Fischnatter
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Srilankansk Fiskesnog
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1887 | |
Tropidonotus quincunciatus var. unicolor Müller |
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Fowlea unicolor Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel 2022 |
Remarks:
Previously regarded as a synonym of piscator or sanctijohannis (Vogel & David 2012).
Revalidated as a separate species by Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel (2022).
Records of piscator from Sri Lanka refer to unicolor.
The species is probably also present in S. India, but this needs verification (Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel 2022).
Distribution:
Sri Lanka.
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Fowlea yunnanensis
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Yunnan Olive Keelback
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Chinesische Fischnatter, Chinesische Olivfarbige Kielnatter
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Kinesisk Fiskesnog, Kinesisk Olivenkølsnog
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1879 | |
Atretium schistosum yunnanensis Anderson |
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Helicops schistosus yunnanensis Annandale 1911 |
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Helicops yunnanensis Pope 1935 |
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Atretium yunnanensis Smith 1943 |
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Fowlea yunnanensis Cheng, Ren, Zhu & Li 2021 |
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1909 | |
Helicops schistosus andersonii Wall (Zhao & Adler 1993) |
Remarks:
Previously assigned to the genus
Atretium.
Transferred to Fowlea by Cheng, Ren, Zhu & Li (2021) and Deepak, Cooper, Poyarkov, Kraus, Burin, Das, Narayanan, Streicher, Smith & Gower (2022).
Distribution:
China
(Yunnan).
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