Reptiles of Cambodia    Bibliography of the reptiles of Cambodia Taxonomic history of the reptiles of Cambodia Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus porosus Saltwater Crocodile Daltry & Dany 2000 King & Burke 1989 Platt, Holloway, Evans, Paudyal, Piron & Rainwater 2006 Thouk & Tana 1994   Crocodylus siamensis Siamese Freshwater Crocodile Bezuijen, Simpson, Behler, Daltry & Tempsiripong 2012 Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Han, Leng, Ratanapich, Piseth, Sovannara, Simpson, Starr. Brook, Frechette & Daltry 2015 Platt, Sovannara, Kheng, Stuart & Walston 2006

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthosaura (Pricklenapes)   Acanthosaura capra Greater Green Pricklenape Ananjeva, Orlov & Kalyabina-Hauf 2008 Daltry & Dany 2000 David & Ineich in Bourret 2009 Manthey 2008 Nguyen, Ho & Nguyen 2009 Stuart, Sok & Neang 2006   Acanthosaura cardamomensis Cardamom Mountains Pricklenape Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Wood, Grismer, Grismer, Neang, Chav & Holden 2010   Acanthosaura coronata Crowned Pricklenape Ananjeva, Orlov & Kalyabina-Hauf 2008 David & Ineich in Bourret 2009 Manthey 2008 Nguyen, Ho & Nguyen 2009 Stuart, Rowley, Neang, Emmett & Som 2010 Stuart, Sok & Neang 2006   Acanthosaura crucigera Northern Masked Pricklenape Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 [as "cf. crucigera"] Manthey 2008 Smith 1935 Stuart & Emmett 2006   Acanthosaura lepidogaster Brown Pricklenape Daltry & Dany 2000 David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Smith 1935   Acanthosaura nataliae Gia Lai Pricklenape Stuart, Rowley, Neang, Emmett & Som 2010 Genus Bronchocela (Long-tailed Agamas)   Bronchocela smaragdina Emerald Long-tailed Agama Daltry & Dany 2000 David & Ineich in Bourret 2009 Hallermann 2005 Manthey 2008 Mulcahy, Cota, Makchai & Stuart 2021 Nguyen, Ho & Nguyen 2009 Smith 1935 Stuart, Sok & Neang 2006 Genus Calotes (South Asian Forest Agamas)   Calotes bachae Eastern Blue Crested Agama Bezuijen, Vinn & Seng 2009 [as C. mystaceus] David & Ineich in Bourret 2009 [as C. mystaceus (part)] Neang, Grismer, Hun & Phan 2015 Nguyen, Ho & Nguyen 2009 [as C. mystaceus (part)] Smith 1935 [as C. mystaceus (part)] Stuart, Sok & Neang 2006 [as C. mystaceus]   Calotes emma Forest Crested Agama Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Nguyen, Ho & Nguyen 2009 Stuart & Emmett 2006 Stuart, Sok & Neang 2006   Calotes goetzi Cambodian Blue Crested Agama Daltry & Dany 2000 [as C. mystaceus] David & Ineich in Bourret 2009 [as C. mystaceus (part)] Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as C. mystaceus] Grismer, Neang, Chav & Grismer 2008 [as C. mystaceus] Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 [as C. mystaceus] Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 [as C. mystaceus] Nguyen, Ho & Nguyen 2009 [as C. mystaceus (part)] Smith 1935 [as C. mystaceus (part)] Stuart & Emmett 2006 [as C. mystaceus] Wagner, Ihlow, Hartmann, Flecks, Schmitz & Boehme 2021   Calotes irawadi Ayeyarwady Forest Agama Gowande, Pal, Jablonski, Masroor, Phansalkar, Dsouza, Jayarajan & Shanker 2021   Calotes versicolor South Indian Crested Agama Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Nguyen, Ho & Nguyen 2009 Stuart, Sok & Neang 2006 Genus Draco (Gliding Dragons)   Draco indochinensis Indochinese Gliding Dragon David & Ineich in Bourret 2009 Grismer, Neang, Chav & Grismer 2008 Musters 1983 Nguyen, Ho & Nguyen 2009 Stuart, Sok & Neang 2006 Wermuth 1967   Draco maculatus Black-spotted Gliding Dragon Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Manthey 2008 [as "cf. maculatus"] Musters 1983 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Nguyen, Ho & Nguyen 2009 Stuart & Emmett 2006   Draco taeniopterus Narrow-lined Gliding Dragon Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Manthey 2008 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Stuart & Emmett 2006 Genus Leiolepis (Butterfly Agamas)   Leiolepis belliana Common Butterfly Agama Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as "cf. belliana"] Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009   Leiolepis reevesii Eastern Butterfly Agama Cox, Dijk, Nabhitabhata & Thirakhupt 1998 Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Nguyen, Ho & Nguyen 2009   Leiolepis rubritaeniata Red-banded Butterfly Agama Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Hartmann, Sothanin, Handschuh & Boehme 2012 Genus Physignathus (Indochinese Water Dragon)   Physignathus cocincinus Indochinese Water Dragon Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Manthey 2010 Nguyen, Ho & Nguyen 2009 Stuart & Emmett 2006 Stuart, Sok & Neang 2006 Genus Pseudocalotes (Long-headed Agamas)   Pseudocalotes floweri Chanthaburi Long-headed Agama Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Neang, Chav & Grismer 2008 Hallermann & Boehme 2000 Manthey 2010 Smith 1935  Family Dibamidae (Blind Skinks) Genus Dibamus (Oriental Blind Skinks)   * Dibamus dalaiensis Phnom Dalai Blind Skink Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 [as D. sp.] Neang, Holden, Eastoe, Seng, Ith & Grismer 2011   Dibamus montanus Mountain Blind Skink Ineich 1999 [as "cf. montanus"]  Family Gekkonidae (Typical Geckos) Genus Cnemaspis (Asian Rock and Forest Geckos)   Cnemaspis chanthaburiensis Chanthaburi Rock Gecko David & Ineich in Bourret 2009 Grismer, Grismer & Chav 2010 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Grismer, Wood, Anuar, Riyanto, Ahmad, Muin, Sumontha, Grismer, Chan, Quah & Pauwels 2014 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Quah, Wood, Anuar, Muin & Grismer 2020   * Cnemaspis neangthyi Cambodian Rock Gecko Grismer, Grismer & Chav 2010 Grismer, Wood, Anuar, Riyanto, Ahmad, Muin, Sumontha, Grismer, Chan, Quah & Pauwels 2014 Quah, Wood, Anuar, Muin & Grismer 2020 Genus Cyrtodactylus (Australasian Bent-toed Geckos)   * Cyrtodactylus auralensis Phnom Aural Bent-toed Gecko Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer 2019   * Cyrtodactylus bokorensis Bokor Bent-toed Gecko Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer 2019   * Cyrtodactylus cardamomensis Cardamom Mountains Bent-toed Gecko Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer 2019   * Cyrtodactylus kohrongensis Koh Rong Bent-toed Gecko Grismer, Onn, Oaks, Neang, Sokun, Murdoch, Stuart & Grismer 2020   * Cyrtodactylus kulenensis Mount Kulen Bent-toed Gecko Grismer, Geissler, Neang, Hartmann, Wagner & Poyarkov 2021   * Cyrtodactylus laangensis Phnom Laang Bent-toed Gecko Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer 2019   * Cyrtodactylus phnomchiensis Phnom Chi Bent-toed Gecko Neang, Henson & Stuart 2020   Cyrtodactylus pseudoquadrivirgatus Thua Thien-Hue Bent-toed Gecko Stuart, Rowley, Neang, Emmett & Som 2010   * Cyrtodactylus thylacodactylus Phnom Dalai Bent-toed Gecko Murdoch, Grismer, Wood, Neang, Poyarkov, Ngo, Nazarov, Aowphol, Pauwels, Nguyen & Grismer 2019 Genus Dixonius (Southeast Asian Leaf-toed Geckos)   Dixonius siamensis Siamese Leaf-toed Gecko Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Nguyen, Ho & Nguyen 2009   Dixonius vietnamensis Vietnamese Leaf-toed Gecko David & Ineich in Bourret 2009 Nguyen, Ho & Nguyen 2009 Stuart, Sok & Neang 2006 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Neang, Chav & Grismer 2008 Nguyen, Ho & Nguyen 2009 Stuart, Sok & Neang 2006 Genus Gekko (Tokays)   Gekko gecko Tokay Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Nguyen, Ho & Nguyen 2009 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Stuart, Sok & Neang 2006   Gekko petricolus Ubon Sandstone Gecko David & Ineich in Bourret 2009 Harbig 2006 Neang, Grismer, Hun & Phan 2015 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011   Gekko tokehos Cambodian Gliding Gecko David & Ineich in Bourret 2009 [as Ptychozoon lionotum] Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 [as Ptychozoon lionotum] Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as Ptychozoon lionotum] Grismer, Neang, Chav & Grismer 2008 [as Ptychozoon lionotum] Grismer, Wood, Grismer, Quah, Neang, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza & Brown 2019 [as Ptychozoon tokehos] Herr & Lee 2016 [as Ptychozoon lionotum] Nguyen, Ho & Nguyen 2009 [as Ptychozoon lionotum] Stuart & Emmett 2006 [as Ptychozoon lionotum]   Gekko trinotaterra Three-banded Gliding Gecko Grismer, Wood, Grismer, Quah, Neang, Phimmachak, Sivongxay, Seateun, Stuart, Siler, Mulcahy, Anamza & Brown 2019 [as Ptychozoon trinotaterra] Hartmann, Betts, Greef & Ihlow 2014 [as Ptychozoon trinotaterra] Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013   Hemidactylus garnotii Fox Gecko Neang, Chhin, Kris & Hun 2011   Hemidactylus platyurus Flat-tailed Gecko Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Nguyen, Ho & Nguyen 2009 Stuart, Sok & Neang 2006   Hemidactylus vietnamensis Vietnamese House Gecko Nguyen, Ho & Nguyen 2009  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   Takydromus sexlineatus Six-lined Grass Lizard Bezuijen, Vinn & Seng 2009 Bischoff 1992 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Stuart & Emmett 2006 Stuart, Sok & Neang 2006  Family Scincidae (Skinks) Genus Dasia (Indo-Malayan Tree Skinks)   Dasia olivacea Olive Tree Skink David & Ineich in Bourret 2009 Greer 1970 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Stuart & Emmett 2006 Genus Eutropis (South Asian Sun Skinks)   Eutropis longicaudata Long-tailed Sun Skink Bezuijen, Vinn & Seng 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Neang, Chav & Grismer 2008   Eutropis macularia Speckled Forest Skink Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Stuart, Sok & Neang 2006   Eutropis multifasciata Many-lined Sun Skink Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 O'Shea, Kusuma & Kaiser 2018 Stuart, Sok & Neang 2006 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia microcerca Laotian Striped Skink Bezuijen, Vinn & Seng 2009 [as vittigera] Daltry & Dany 2000 [as vittigera] David & Ineich in Bourret 2009 [as vittigera] Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as vittigera] Grismer, Neang, Chav & Grismer 2008 [as vittigera] Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 [as vittigera] Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 [as vittigera] Mahony 2008 [as vittigera] Poyarkov, Geissler, Gorin, Dunayev, Hartmann & Suwannapoom 2019 Stuart & Emmett 2006 [as vittigera] Stuart, Sok & Neang 2006 [as vittigera]   Lipinia trivittata Vietnamese Three-striped Skink Poyarkov, Geissler, Gorin, Dunayev, Hartmann & Suwannapoom 2019   Lipinia vassilievi Kon Tum Striped Skink Poyarkov, Geissler, Gorin, Dunayev, Hartmann & Suwannapoom 2019 Genus Lygosoma (Southeast Asian Writhing Skinks)   Lygosoma isodactylum Even-toed Writhing Skink Cox, Dijk, Nabhitabhata & Thirakhupt 1998 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011   Lygosoma siamense Siamese Writhing Skink Daltry & Dany 2000 [as L. quadrupes] David & Ineich in Bourret 2009 [as L. quadrupes] Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011 [as L. quadrupes] Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as L. quadrupes] Grismer, Neang, Chav & Grismer 2008 [as L. quadrupes] Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 [as L. quadrupes] Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer 2018 Stuart, Sok & Neang 2006 [as L. quadrupes] Genus Plestiodon (Blue-tailed Skinks)   Plestiodon quadrilineatus Four-striped Long Skink Daltry & Dany 2000 Nguyen, Ho & Nguyen 2009 Genus Riopa (Asian Gracile Skinks)   Riopa angeli Dong Nai Gracile Skink Neang, Morawska & Nut 2015   Riopa haroldyoungi Banded Gracile Skink Geissler, Farkas, Samorn, Nanthawichianrit, Chea & Neang 2022   * Riopa veunsaiensis Veun Sai Gracile Skink Geissler, Hartmann & Neang 2012 Genus Scincella (South Asian and American Ground Skinks)   Scincella melanosticta Black-spotted Ground Skink Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Stuart & Emmett 2006   Scincella nigrofasciata Black-striped Ground Skink Neang, Chan & Poyarkov 2018   Scincella reevesii Speckled Ground Skink Daltry & Dany 2000 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010   Scincella rufocaudata Red-tailed Forest Skink David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Stuart & Emmett 2006 Stuart, Sok & Neang 2006   Scincella rupicola Rock-dwelling Ground Skink Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus annamiticus Annam Forest Skink David & Ineich in Bourret 2009 [as stellatus] Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as stellatus] Grismer, Neang, Chav & Grismer 2008 [as stellatus] Grismer, Wood, Quah, Anuar, Poyarkov, Neang, Orlov, Thammachoti & Hun 2019 Stuart & Emmett 2006 [as stellatus]   Sphenomorphus buenloicus Buon Luoi Forest Skink Neang & Poyarkov 2016   Sphenomorphus indicus Common Indian Forest Skink Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008   Sphenomorphus lineopunctulatus Southeast Asian Spot-lined Forest Skink Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Hartmann, Nguyen, Ohler, Chippich, Handschuh & Boehme 2010   Sphenomorphus maculatus Spotted Forest Skink Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Smith 1935 Stuart & Emmett 2006 Stuart, Sok & Neang 2006   * Sphenomorphus preylangensis Prey Lang Forest Skink Grismer, Wood, Quah, Anuar, Poyarkov, Neang, Orlov, Thammachoti & Hun 2019 Genus Subdoluseps (Asian Agile Skinks)   Subdoluseps bowringii Short-legged Agile Skink Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Geissler, Nguyen, Phung, Van Devender, Hartmann, Farkas, Ziegler & Boehme 2011 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Le, Nguyen, Phan, Rujirawan, Aowphol, Vo, Murphy & Nguyen 2021 Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer 2018 Stuart & Emmett 2006 Genus Tropidophorus (Stream Skinks)   Tropidophorus cocincinensis Indochinese Stream Skink Chuaynkern, Nabhitabhata, Inthara, Kamsook & Somsri 2005 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Hartmann, Chipich, Handschuh, Nguyen & Boehme 2009 Stuart, Rowley, Neang, Emmett & Som 2010   Tropidophorus microlepis Small-scaled Stream Skink Chuaynkern, Nabhitabhata, Inthara, Kamsook & Somsri 2005 David & Ineich in Bourret 2009 Stuart, Sok & Neang 2006  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus nebulosus Clouded Monitor Auffenberg 1994 [as V. bengalensis] Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 [as V. bengalensis] David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013   Varanus salvator Asian Water Monitor Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 David & Ineich in Bourret 2009 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Smith 1935

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Nguyen, Ho & Nguyen 2009 Saint Girons 1972 Wallach, Williams & Boundy 2014   Acrochordus javanicus Java File Snake Daltry & Dany 2000 McDiarmid, Campbell & Toure 1999 Nguyen, Ho & Nguyen 2009 Saint Girons 1972  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla fusca Dark Whip Snake David, Nadolski, Ganesh, Adhikari & Srikanthan 2022   Ahaetulla nasuta Long-nosed Green Whip Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Saint Girons 1972 Tran & Nguyen 2021 Wallach, Williams & Boundy 2014   Ahaetulla prasina Short-nosed Whip Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Saint Girons 1972 Stuart & Emmett 2006 Stuart, Sok & Neang 2006 Wallach, Williams & Boundy 2014 Genus Amphiesma (South Asian Keelbacks)   Amphiesma stolatum Buff-striped Keelback Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Saint Girons 1972 Stuart & Emmett 2006 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga cyanea Green Cat Snake Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Ganesh, Mallik, Achyuthan, Shanker & Vogel 2021 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Orlov & Ryabov 2002 Saint Girons 1972 Stuart & Emmett 2006 Stuart, Sok & Neang 2006   Boiga guangxiensis Guangxi Cat Snake Neang, Chan, Chhin, Samorn, Poyarkov, Stephens, Daltry & Stuart 2017   Boiga melanota Western Mangrove Cat Snake Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as B. dendrophila] Grismer, Neang, Chav & Grismer 2008 [as B. dendrophila] Nguyen, Ho & Nguyen 2009 Orlov & Ryabov 2002 Stuart & Emmett 2006 [as B. dendrophila]   Boiga multomaculata Marbled Cat Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Orlov & Ryabov 2002 Saint Girons 1972 Stuart & Emmett 2006   Boiga siamensis Grey Cat Snake Daltry & Dany 2000 Ganesh, Mallik, Achyuthan, Shanker & Vogel 2021 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Kroon 1973 [as B. ocellata] Nguyen, Ho & Nguyen 2009 Orlov & Ryabov 2002 Saint Girons 1972 [as cynodon] Smith 1943 [as cynodon] Stuart & Emmett 2006 Genus Calamaria (Oriental Reed Snakes)   Calamaria pavimentata Brown Reed Snake Daltry & Dany 2000 Inger & Marx 1965 Saint Girons 1972 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Mertens 1968 Saint Girons 1972 Stuart & Emmett 2006 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus flavolineatus Yellow-striped Rat Snake Grismer, Neang, Chav & Grismer 2008   Coelognathus radiatus Copper-headed Rat Snake Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Saint Girons 1972 Schulz 1996 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis ngansonensis Nganson Bronzeback Neang, Grismer, Hun & Phan 2015   Dendrelaphis pictus Southern Painted Bronzeback Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Saint Girons 1972 Vogel & Rooijen 2008 Vogel & Rooijen 2011   Dendrelaphis subocularis Burmese Mountain Bronzeback Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Saint Girons 1972 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops rubescens Keel-bellied Vine Snake Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Nguyen & Tran 2021 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe taeniura Stripe-tailed Rat Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Schulz 2010 Genus Fowlea (Fishing Keelbacks)   Fowlea flavipunctata Yellow-spotted Keelback Amarasinghe, Bandara, Weerakkody, Campbell, Marques, Danushka, Silva & Vogel 2022 Bezuijen, Vinn & Seng 2009 [as piscator] Brooks, Allison, Gill & Reynolds 2009 [as piscator] Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Saint Girons 1972 Vogel & David 2006 Vogel & David 2012 Genus Gongylosoma (Southeast Asian Ground Snakes)   Gongylosoma scriptum Indochinese Ground Snake Lalremsanga, Bal, Vogel & Biakzuala 2022 Neang, Grismer, Hun & Phan 2015 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Saint Girons 1972 Schulz 1996 Stuart & Emmett 2006 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius boulengeri White-browed Keelback David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 [as "cf. boulengeri"] David, Vogel & Rooijen 2013 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as "cf. khasiense"] Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Nguyen, Ho & Nguyen 2009 [as khasiense] Purkayastha & David 2019 Stuart & Emmett 2006 [as khasiense] Wallach, Williams & Boundy 2014   Hebius leucomystax White-lipped Keelback Stuart, Rowley, Neang, Emmett & Som 2010 Wallach, Williams & Boundy 2014 Genus Liopeltis (Ringneck Snakes)   Liopeltis stoliczkae Northeast Indian Ringneck Snake Stuart, Sok & Neang 2006 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 [as aulicus] Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Iskandar & Colijn 2001 Neang, Hartmann, Hun, Souter & Furey 2014 Nguyen, Ho & Nguyen 2009 O'Shea, Kusuma & Kaiser 2018 Saint Girons 1972   Lycodon cardamomensis Cardamom Mountains Wolf Snake Daltry & Wuster 2002 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Neang, Hartmann, Hun, Souter & Furey 2014 Vogel & David 2019   Lycodon davisonii Burmese Bridle Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Saint Girons 1972 Stuart & Emmett 2006   Lycodon laoensis Laotian Wolf Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Neang, Hartmann, Hun, Souter & Furey 2014 Nguyen, Ho & Nguyen 2009 Saint Girons 1972 Wang, Qi, Lyu, Zeng & Wang 2020   Lycodon septentrionalis Northern Big-toothed Snake Das & Vasudevan 2015 Neang, Hartmann, Hun, Souter & Furey 2014 Nguyen, Ho & Nguyen 2009 Stuart, Sok & Neang 2006   Lycodon subcinctus Malayan Banded Wolf Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Neang, Hartmann, Hun, Souter & Furey 2014 Nguyen, Ho & Nguyen 2009 Saint Girons 1972   * Lycodon zoosvictoriae Phnom Samkos Wolf Snake Neang, Hartmann, Hun, Souter & Furey 2014 Genus Oligodon (Kukri Snakes)   Oligodon annamensis Annam Kukri Snake Neang & Hun 2013 Nguyen, Tran, Nguyen, Neang, Yushchenko & Poyarkov 2020 Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021   Oligodon barroni Chonburi Kukri Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972   Oligodon cinereus Black-barred Kukri Snake Daltry & Dany 2000 David, Das & Vogel 2011 David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Saint Girons 1972 Smith 1943 Stuart, Sok & Neang 2006   Oligodon deuvei Orange-striped Kukri Snake David, Vogel & Rooijen 2008 Nguyen, Ho & Nguyen 2009   Oligodon eberhardti Tam Dao Kukri Snake Daltry & Dany 2000 Nguyen, Ho & Nguyen 2009   Oligodon fasciolatus Small-banded Kukri Snake Daltry & Dany 2000 David, Das & Vogel 2011 David, Vogel & Pauwels 2008 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Neang, Chav & Grismer 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972 [as smithi] Stuart & Emmett 2006   Oligodon inornatus Southeast Asian Plain Kukri Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Saint Girons 1972 Stuart & Emmett 2006   * Oligodon kampucheaensis Phnom Samkos Kukri Snake Neang, Grismer & Daltry 2012   Oligodon mouhoti Cambodian Kukri Snake David, Vogel & Rooijen 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972   Oligodon ocellatus Ocellated Kukri Snake Daltry & Dany 2000 David, Vogel & Pauwels 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972 Stuart, Sok & Neang 2006   Oligodon saintgironsi Dong Nai Blotched Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 David, Vogel & Pauwels 2008 Nguyen, Ho & Nguyen 2009   Oligodon taeniatus Striped Kukri Snake Daltry & Dany 2000 David, Vogel & Rooijen 2008 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Saint Girons 1972 Genus Oreocryptophis (Red Mountain Rat Snake)   Oreocryptophis porphyraceus Red Mountain Rat Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas carinata Malayan Keeled Rat Snake Nguyen, Ho & Nguyen 2009 Stuart & Emmett 2006 Stuart, Sok & Neang 2006   Ptyas korros Indochinese Rat Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Saint Girons 1972 Stuart, Sok & Neang 2006   Ptyas mucosa Common Oriental Rat Snake Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Saint Girons 1972 Sindaco, Venchi & Grieco 2013 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis chrysargos Speckled-bellied Keelback Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 Saint Girons 1972 Stuart, Sok & Neang 2006   Rhabdophis nigrocinctus Black-banded Keelback Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009 Stuart & Emmett 2006   Rhabdophis siamensis Siamese Red-necked Keelback Daltry & Dany 2000 [as subminiatus] David & Vogel 2021 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 [as subminiatus] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as subminiatus] Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as subminiatus] Grismer, Neang, Chav & Grismer 2008 [as subminiatus] Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 [as subminiatus] Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 [as subminiatus] Saint Girons 1972 [as subminiatus] Stuart, Sok & Neang 2006 [as subminiatus] Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis collaris Collared Black-headed Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009   Sibynophis triangularis Triangled Black-headed Snake Stuart, Sok & Neang 2006 Genus Xenochrophis (Oriental Painted Keelbacks)   Xenochrophis trianguligerus South Asian Triangle Keelback Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Nguyen, Ho & Nguyen 2009 Stuart & Emmett 2006  Family Cylindrophiidae (Asian Pipe Snakes) Genus Cylindrophis (Oriental Pipe Snakes)   Cylindrophis ruffus Red-tailed Pipe Snake Brooks, Allison, Gill & Reynolds 2009 Daltry & Dany 2000 McDiarmid, Campbell & Toure 1999 Nguyen, Ho & Nguyen 2009 Saint Girons 1972  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus candidus Malayan Krait Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972 Stuart, Sok & Neang 2006   Bungarus fasciatus Banded Krait Bezuijen, Vinn & Seng 2009 Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972 Stuart, Sok & Neang 2006 Genus Calliophis (South Asian Coral Snakes)   Calliophis maculiceps Small-spotted Coral Snake Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as Maticora maculiceps atrofrontalis] Nguyen, Ho & Nguyen 2009 Saint Girons 1972 Genus Naja (Typical Cobras)   Naja kaouthia Monocellate Cobra Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Saint Girons 1972 Wuster & Thorpe 1992 Wuster 1998   Naja siamensis Siamese Spitting Cobra Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Grismer, Neang, Chav & Grismer 2008 Nguyen, Ho & Nguyen 2009 Teynie & David 2007 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Saint Girons 1972 Genus Sinomicrurus (East Asian Coral Snakes)   Sinomicrurus macclellandii Southeast Asian Coral Snake Neang, Chan, Chhin, Samorn, Poyarkov, Stephens, Daltry & Stuart 2017 Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Aipysurus (Olive Sea Snakes)   Aipysurus eydouxii White-spotted Sea Snake Sanders, Rasmussen, Elmberg, Mumpuni, Guinea, Blias, Lee & Fry 2012 Genus Hydrophis (Typical Sea Snakes)   Hydrophis schistosus Beaked Sea Snake Saint Girons 1972 Smith 1943   Hydrophis torquatus Grey Sea Snake Ineich 1996 Nguyen, Ho & Nguyen 2009 Genus Kolpophis (Granular Sea Snake)   Kolpophis annandalei Granular Sea Snake Nguyen, Ho & Nguyen 2009 Taylor 1965 Genus Thalassophis (Scale-snouted Sea Snake)   Thalassophis anomalus Scale-snouted Sea Snake Smith 1943 Taylor 1965  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Nguyen, Ho & Nguyen 2009 [as C. rynchops] Saint Girons 1972 [as C. rynchops] Genus Enhydris (Asian Mud Snakes)   Enhydris enhydris Striped Mud Snake Bezuijen, Vinn & Seng 2009 Brooks, Allison, Gill & Reynolds 2009 Daltry & Dany 2000 Gyi 1970 Murphy & Voris 2014 Murphy 2007 Saint Girons 1972   Enhydris innominata Tay Ninh Mud Snake Daltry & Dany 2000 Murphy 2007 Nguyen, Ho & Nguyen 2009   * Enhydris longicauda Long-tailed Mud Snake Bezuijen, Vinn & Seng 2009 Brooks, Allison, Gill & Reynolds 2009 Daltry & Dany 2000 Gyi 1970 Murphy & Voris 2014 Murphy 2007 Saint Girons 1972   Enhydris subtaeniata Mekong Mud Snake Murphy & Voris 2014 Murphy 2007 Nguyen, Ho & Nguyen 2009 Saint Girons 1972 [as E. jagori] Voris, Murphy, Karns, Kremer & O'Connell 2012 Genus Erpeton (Tentacled Snake)   Erpeton tentaculatum Tentacled Snake Bezuijen, Vinn & Seng 2009 Brooks, Allison, Gill & Reynolds 2009 Daltry & Dany 2000 Gyi 1970 Murphy & Voris 2014 Murphy 2007 Saint Girons 1972 Genus Fordonia (White-bellied Mangrove Snake)   Fordonia leucobalia White-bellied Mangrove Snake Nguyen, Ho & Nguyen 2009 Genus Gerarda (Glossy Marsh Snake)   Gerarda prevostiana Glossy Marsh Snake Murphy & Voris 2014 Genus Homalopsis (Masked Water Snakes)   Homalopsis mereljcoxi Thai Masked Water Snake Murphy & Voris 2014 Murphy, Voris, Murthy, Traub & Cumberbatch 2012   Homalopsis nigroventralis Mekong Masked Water Snake Bezuijen, Vinn & Seng 2009 Murphy & Voris 2014 Murphy 2007 Murphy, Voris, Murthy, Traub & Cumberbatch 2012 Stuart, Sok & Neang 2006 Genus Hypsiscopus (Orange-bellied Mud Snake and Matanna Mud Snake)   Hypsiscopus murphyi Northern Orange-bellied Mud Snake Bernstein, Voris, Stuart, Phimmachak, Seateun, Sivongxay, Neang, Karns, Andrews, Osterhage, Phipps & Ruane 2022   Hypsiscopus plumbea Southern Orange-bellied Mud Snake Bernstein, Voris, Stuart, Phimmachak, Seateun, Sivongxay, Neang, Karns, Andrews, Osterhage, Phipps & Ruane 2022 Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Saint Girons 1972 Stuart, Sok & Neang 2006 Genus Subsessor (Southeast Asian Mud Snake)   Subsessor bocourti Southeast Asian Mud Snake Brooks, Allison, Gill & Reynolds 2009 Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Gyi 1970 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Murphy & Voris 2014 Murphy 2007 Saint Girons 1972 Stuart & Emmett 2006  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Saint Girons 1972 Stuart & Emmett 2006 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis indochinensis Indochinese Sand Snake Hartmann, Handschuh & Boehme 2011 Nguyen, Ho & Nguyen 2009  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas berdmorei Mon Slug Eater Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022   Pareas margaritophorus White-spotted Slug Eater Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Hauser 2017 Saint Girons 1972 Stuart & Emmett 2006 Stuart, Sok & Neang 2006 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 McDiarmid, Campbell & Toure 1999 Saint Girons 1972 Genus Python (Typical Pythons)   Python bivittatus Burmese Python Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Iskandar & Colijn 2001 Nguyen, Ho & Nguyen 2009 Saint Girons 1972  Family Typhlopidae (Typical Blind Snakes) Genus Argyrophis (Southeast Asian Blind Snakes)   Argyrophis diardii Greater Blind Snake Daltry & Dany 2000 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Nguyen, Ho & Nguyen 2009 Saint Girons 1972   Argyrophis muelleri Malayan Bicoloured Blind Snake Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Stuart & Emmett 2006   Argyrophis siamensis Siamese Blind Snake Daltry & Dany 2000 McDiarmid, Campbell & Toure 1999 Saint Girons 1972 Genus Ramphotyphlops (Western Pacific Blind Snakes)   Ramphotyphlops albiceps Southeast Asian White-headed Blind Snake Neang, Chan, Chhin, Samorn, Poyarkov, Stephens, Daltry & Stuart 2017 Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 McDiarmid, Campbell & Toure 1999 Saint Girons 1972 Wallach 2009 Wallach 2020  Family Viperidae (Vipers) Genus Calloselasma (Malayan Pitviper)   Calloselasma rhodostoma Malayan Pitviper Bezuijen, Vinn & Seng 2009 Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Gloyd & Conant 1990 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Saint Girons 1972 Stuart & Emmett 2006 Stuart, Sok & Neang 2006 Genus Daboia (Daboias)   Daboia siamensis Siamese Chain Viper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 [as russelii] Iskandar & Colijn 2001 Neang, Grismer, Hun & Phan 2015 Saint Girons 1972 Wuster, Otsuka, Malhotra & Thorpe 1992 Genus Ovophis (Asian Mountain Pitvipers)   Ovophis convictus Indo-Malayan Mountain Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Neang, Chhin, Kris & Hun 2011 Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus albolabris Southeast Asian White-lipped Tree Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Regenass & Kramer 1981 Saint Girons 1972 Stuart & Emmett 2006 Stuart, Sok & Neang 2006   Trimeresurus cardamomensis Cardamom Mountains Green Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Malhotra, Thorpe, Mrinalini & Stuart 2011   Trimeresurus macrops Big-eyed Green Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Malhotra, Thorpe, Mrinalini & Stuart 2011   Trimeresurus rubeus Ruby-eyed Green Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Malhotra, Thorpe, Mrinalini & Stuart 2011 Genus Viridovipera (Green Tree Pitvipers)   Viridovipera vogeli Khao Yai Green Tree Pitviper Cox, Hoover, Chanhome, Thirakhupt & Pongcharoen 2018 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as Trimeresurus s. stejnegeri] Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Neang, Grismer, Chan, Grismer, Wood & Youmans 2010 [as "cf. vogeli"] Nguyen, Ho & Nguyen 2009 Regenass & Kramer 1981 [as Trimeresurus s. stejnegeri] Stuart & Emmett 2006 Stuart, Sok & Neang 2006  Family Xenodermidae (Asian Litter Snakes) Genus Fimbrios (Southern Bearded Snakes)   Fimbrios klossi Lesser Bearded Snake Ziegler, David, Miralles, Doan & Nguyen 2008  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis unicolor Common Asian Sunbeam Snake Daltry & Dany 2000 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 [as concolor] Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 McDiarmid, Campbell & Toure 1999 Orlov 2000 Saint Girons 1972 Stuart & Emmett 2006 Stuart, Sok & Neang 2006

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Diamond, Blanco & Duncan 2012 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Diamond, Blanco & Duncan 2012 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Diamond, Blanco & Duncan 2012 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, An & Dijk 2002  Family Geoemydidae (Eastern Pond Turtles) Genus Batagur (River Terrapins)   Batagur affinis Southern River Terrapin Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as baska] Grismer, Neang, Chav & Grismer 2008 [as baska] Moll, Platt, Chan, Horne, Platt, Praschag, Chen & Dijk 2015 Platt, Stuart, Sovannara, Kheng, Kalyar & Kimchhay 2003 [as baska] Praschag, Holloway, Georges, Packert, Hundsdoerfer & Fritz 2009 Praschag, Sommer, McCarthy, Gemel & Fritz 2008 [as baska] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schoppe & Das 2011 Stuart, Dijk & Hendrie 2001   Cuora bourreti Central Indochinese Box Turtle Lehr, Fritz & Obst 1998 McCord & Joseph-Ouni 2002 Nguyen, Ho & Nguyen 2009   Cuora picturata Southern Indochinese Box Turtle Lehr, Fritz & Obst 1998 McCord & Joseph-Ouni 2002 Nguyen, Ho & Nguyen 2009 Genus Cyclemys (Asian Leaf Turtles)   Cyclemys atripons Southeastern Leaf Turtle Fritz & Havas 2006 Fritz, Guicking, Auer, Sommer, Wink & Hundsdoerfer 2008 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Iverson & McCord 1997 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Sok & Neang 2006   Cyclemys oldhamii Dark-throated Leaf Turtle Fritz, Guicking, Auer, Sommer, Wink & Hundsdoerfer 2008 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 [as "C. cf. oldhamii"] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Rowley, Neang, Emmett & Som 2010   Cyclemys pulchristriata Striped Leaf Turtle Fritz, Guicking, Auer, Sommer, Wink & Hundsdoerfer 2008 Nguyen, Nguyen & Nguyen 2018 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Heosemys (Forest Turtles)   Heosemys annandalii Yellow-headed Temple Turtle Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Fritz & Havas 2006 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001   Heosemys grandis Giant Asian Pond Turtle Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Fritz & Havas 2006 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001 Genus Malayemys (Southeast Asian Snail-eating Turtles)   Malayemys subtrijuga Mekong Snail-eating Turtle Bezuijen, Vinn & Seng 2009 Brophy 2005 Dawson, Ihlow & Platt 2020 Fritz & Havas 2006 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Notochelys (Malayan Flat-shelled Turtle)   Notochelys platynota Malayan Flat-shelled Turtle Daltry & Dany 2000 King & Burke 1989 Genus Siebenrockiella (Black Marsh Turtle and Philippine Pond Turtle)   Siebenrockiella crassicollis Black Marsh Turtle Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001  Family Platysternidae (Big-headed Turtle) Genus Platysternon (Big-headed Turtle)   Platysternon megacephalum Big-headed Turtle Daltry & Dany 2000 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Indotestudo (Asian Tortoises)   Indotestudo elongata Elongated Tortoise Bezuijen, Vinn & Seng 2009 Daltry & Dany 2000 Fritz & Havas 2006 Geissler, Hartmann, Ihlow, Neang, Seng, Wagner & Boehme 2019 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 Ihlow, Dawson, Hartmann & Som 2016 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001 Genus Manouria (Indochinese Tortoises)   Manouria impressa Impressed Tortoise Daltry & Dany 2000 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 King & Burke 1989 Lehr & Holloway 2000 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Trionychidae (Softshell Turtles) Genus Amyda (Southeast Asian Softshell Turtles)   Amyda ornata Black-rayed Softshell Turtle Auliya, Dijk, Moll & Meylan 2016 [as A. cartilaginea] Bezuijen, Vinn & Seng 2009 [as A. cartilaginea] Daltry & Dany 2000 [as A. cartilaginea] Fritz, Gemel, Kehlmaier, Vamberger & Praschag 2014 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 [as A. cartilaginea] Grismer, Neang, Chav & Grismer 2008 [as A. cartilaginea] Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008 [as A. cartilaginea] Hartmann, Ihlow, Edwards, Sovath, Handschuh & Boehme 2013 [as A. cartilaginea] King & Burke 1989 [as A. cartilaginea] Nguyen, Ho & Nguyen 2009 [as A. cartilaginea] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001 [as A. cartilaginea] Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Daltry & Dany 2000 Das 2008 Grismer, Chav, Neang, Wood, Grismer, Youmans, Ponce, Daltry & Kaiser 2007 Grismer, Neang, Chav & Grismer 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Stuart, Dijk & Hendrie 2001