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ę Rune Midtgaard


Taxonomy of the family Phrynosomatidae
Bibliography of the genus Urosaurus
Biodiversity of the family Phrynosomatidae








Genus
Urosaurus

North American Tree Lizards, Brush Lizards

Nordamerikanische Baumleguane

Nordamerikanske TrŠleguaner

1854 Urosaurus Hallowell (type species: Urosaurus graciosus Hallowell 1854)
1856 Phymatolepis Dumeril (type species: Phymatolepis bi-carinatus Dumeril 1856; syn. Smith & Taylor 1950)
Contents: 9 species, of which 5 (55.6%) are endemic.
Distribution: W. USA, Mexico.
Reported from: Mexico (Aguascalientes, Baja California Norte, Baja California Sur, Chiapas, Chihuahua, Coahuila, Colima [incl. Revillagigedo Islands (Clarion, Socorro)], Durango, Guerrero, Jalisco, Mexico, Michoacan, Morelos, Nayarit [incl. Islas Marias (Maria Madre, Maria Magdalena, San Juanito)], Nuevo Leon, Oaxaca, Puebla, Sinaloa, Sonora, Tamaulipas, Zacatecas), USA (Arizona, California, Colorado, Nevada, New Mexico, Texas, Utah, Wyoming).

Urosaurus auriculatus

Socorro Island Tree Lizard, Socorro Island Rock Lizard

Socorro-Baumleguan*

Socorro-trŠleguan*

1871 Uta auriculata Cope
Urosaurus auriculatus Mittleman 1942

Distribution: Mexico (Colima: Revillagigedo Islands: Socorro).


Urosaurus bicarinatus

Tropical Tree Lizard, Bicarinate Tree Lizard

Doppelgekielter Baumleguan*

Tok°let TrŠleguan*

1856 Phymatolepis bi-carinatus Dumeril
Uta bicarinata Cope 1864
Urosaurus bicarinatus Mittleman 1942
1921 Uta nelsoni Schmidt (Smith & Taylor 1950)
Uta bicarinata nelsoni Mittleman 1941
Urosaurus bicarinatus nelsoni Mittleman 1942
1921 Uta tuberculata Schmidt (Smith & Taylor 1950)
Uta bicarinata tuberculata Mittleman 1941
Urosaurus bicarinatus tuberculatus Mittleman 1942
1940 Uta anonymorpha Mittleman (Smith & Taylor 1950)
Uta bicarinatus anonymorpha Mittleman 1941
Urosaurus bicarinatus anonymorphus Mittleman 1942
1941 Uta unica Mittleman (Hardy & McDiarmid 1969, Oliver 1943, Tanner 1987, Wiens 1993)
Urosaurus unicus Mittleman 1942
1954 Urosaurus bicarinatus spinosus Bumsahem & Smith

Other common names:
anonymorphus: Mittleman's Tree Lizard
bicarinatus: Carinate Tree Lizard
nelsoni: Nelson's Tree Lizard
spinosus: Spiny Tree Lizard
tuberculatus: Tubercular Tree Lizard
Distribution: Mexico (Aguascalientes, Chiapas, Chihuahua, Colima, Durango, Guerrero, Jalisco, Mexico, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Sinaloa, Sonora, Zacatecas).


Urosaurus clarionensis

Clarion Island Tree Lizard, Clarion Island Rock Lizard

Clarion-Baumleguan*

Clarion-trŠleguan*

1890 Uta clarionensis Townsend
Urosaurus clarionensis Mittleman 1942

Distribution: Mexico (Colima: Revillagigedo Islands: Clarion).


Urosaurus gadovi

Jaliscan Tree Lizard*, (Gadow's Tree Lizard)

Jalisco-Baumleguan*

Jalisco-trŠleguan*

1921 Uta gadovi Schmidt
Urosaurus gadovi Mittleman 1942

Remarks: Duellman (1958c) considered the type locality (CofradÝa, Jalisco) to be an error based on confusion between two different villages of the same name, and he restricted it to CofradÝa in Michoacan.
Distribution: Mexico (Guerrero, Jalisco, Michoacan).


Urosaurus graciosus

Long-tailed Brush Lizard

Nordamerikanische Langschwanz-Baumleguan*

Nordamerikansk Langhalet TrŠleguan*

1854 Urosaurus graciosus Hallowell
Uta graciosa Baird 1859
Uta gratiosa Coues 1875 [invalid emendation]
Urosaurus ornatus graciosus Mittleman 1942
1955 Urosaurus graciosus shannoni Lowe

Other common names:
graciosus: Western Long-tailed Brush Lizard
shannoni: Arizona Long-tailed Brush Lizard
Remarks: Although included for Jalisco by Cruz-Sßenz, Guerrero, Lazcano & TÚlles-Lˇpez (2009), presence in the state is considered undocumented herein, following Cruz-Sßenz, Mu˝oz-Nolasco, Mata-Silva, Johnson, GarcÝa-Padilla & Wilson (2017), who did not list the species for Jalisco.
Distribution: Mexico (Baja California Norte, Sonora), USA (Arizona, California, Nevada, Utah).


Urosaurus lahtelai

Catavi˝a Brush Lizard, (Baja California Brush Lizard)

Catavi˝a-Baumleguan*

Catavi˝a-trŠleguan*

1977 Urosaurus lahtelai Rau & Loomis

Distribution: Mexico (Baja California Norte).


Urosaurus microscutatus

Small-scaled Tree Lizard

Kleinschuppiger Baumleguan*

FinskŠllet TrŠleguan*

1894 Uta microscutata Van Denburgh
Urosaurus microscutatus Mittleman 1942
1900 Uta parviscutata Cope [substitute name for Uta microscutata Van Denburgh 1894]

Remarks: Aquirre-L., Morafka & Murphy (1999) and Grismer (1999) placed microscutatus in the synonymy of nigricaudus, but Feldman, Flores-Villela & Papenfuss (2011) retained them as separate species, pending further studies.
Distribution: Mexico (Baja California Norte), USA (California).


Urosaurus nigricaudus

Baja California Brush Lizard, (Black-tailed Brush Lizard)

Niederkalifornischer Baumleguan*

Baja California-trŠleguan*

1864 Uta nigricauda Cope
Urosaurus nigricaudus Mittleman 1942

Remarks: Some authors regard microscutatus as a synonym of nigricaudus.
Distribution: Mexico (Baja California Norte, Baja California Sur), USA (California).


Urosaurus ornatus

Common Tree Lizard

Gew÷hnlicher Baumleguan*

Almindelig TrŠleguan*

1852 Uta ornata Baird & Girard
Urosaurus ornatus Mittleman 1942
1857 Uta ornata linearis Baird
Urosaurus ornatus linearis Mittleman 1942
1858 Uta schottii Baird (Smith & Taylor 1950)
Uta ornata schottii Mittleman 1941
Urosaurus ornatus schottii Mittleman 1942
1858 Uta symmetrica Baird (Schmidt 1953)
Uta ornata symmetrica Schmidt 1921
Urosaurus ornatus symmetricus Mittleman 1942
1883 Uta lateralis Boulenger (Lemos-Espinal & Smith 2007)
Uta ornata lateralis Van Denburgh 1922
Urosaurus ornatus lateralis Zweifel & Norris 1955
1884 Uta gularis Cragin (Smith & Taylor 1950)
1890 Uta levis Stejneger (Schmidt 1953)
Uta ornata levis Schmidt 1953
Urosaurus ornatus levis Hotton 1955
1921 Uta wrighti Schmidt (Schmidt 1953)
Uta ornata wrighti Schmidt 1953
Urosaurus ornatus wrighti Maslin 1947
1935 Uta caerulea Smith (Smith & Taylor 1950)
Urosaurus ornatus caeruleus Mittleman 1942
1940 Uta ornata schmidti Mittleman
Urosaurus ornatus schmidti Mittleman 1942
1941 Uta ornata chiricahuae Mittleman
Urosaurus ornatus chiricahuae Mittleman 1942

Other common names:
caeruleus: Blue Tree Lizard
lateralis: Mexican Tree Lizard
levis: Smooth Tree Lizard, Canyon Tree Lizard
linearis: Lined Tree Lizard
ornatus: Texas Tree Lizard, Eastern Tree Lizard
schmidti: Big Bend Tree Lizard
schottii: Schott's Tree Lizard
symmetricus: Colorado River Tree Lizard
wrighti: Northern Tree Lizard
Remarks: Reports of schotti from Chihuahua refer to lateralis (Lemos-Espinal & Smith 2007). Although included for Jalisco by Cruz-Sßenz, Guerrero, Lazcano & TÚlles-Lˇpez (2009), presence in the state is considered undocumented herein, following Cruz-Sßenz, Mu˝oz-Nolasco, Mata-Silva, Johnson, GarcÝa-Padilla & Wilson (2017), who did not list the species for Jalisco.
Distribution: Mexico (Baja California Norte, Chihuahua, Coahuila, Durango, Nayarit [incl. Islas Marias (Maria Madre, Maria Magdalena, San Juanito)], Nuevo Leon, Sinaloa, Sonora, Tamaulipas), USA (Arizona, California, Colorado, Nevada, New Mexico, Texas, Utah, Wyoming).