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Taxonomy of the genus Vipera Biodiversity of the family Viperidae

Bibliography of the genus
Vipera (Typical Vipers or Adders)

(Reptilia: Serpentes: Viperidae)

Note: In order to limit redundancy, relevant literature indexed in the related bibliographies in the left column may not have been included in this page. For a comprehensive search of literature, these bibliographies should therefore also be consulted.


Vipera in general

Angelici, F.M.; Luiselli, L. 1998. Ornithophagy in Italian snakes: a review. Bulletin de la Societe Zoologique de France Evolution et Zoologie 123(1): 15-22.

Bakiev, A.G.; Ratnikov, V.Y.; Zinenko, A.I. 2007. [About forming of vipers' fauna of the Volga river basin]. (In Russian). Izvestiya Samarskogo Nauchnogo Centra Rossiskaya Akademii Nauk 9(1)(19): 163-170.

Bakradze, M.A. 1975. [Distribution of Vipera laurenti in south Georgia]. Vestnik Gosudarstvennogo Muzeya Gruzii (A) 28: 411-412.

Basoglu, M. 1949. On some varieties of Vipera berus from the extreme N.E. Anatolia. Revue Faculte des Sciences Universite d'Istanbul 1948: 143-160.

Bea, A. 1986. A general review of the dorsal scales microornamentation in Vipera species (Reptilia: Viperidae). pp. 367-371. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European herpetological meeting (3rd ordinary general meeting of the Societas Europaea Herpetologica), Prague 1985. Charles University, Prague. 754 pp.

Bea, A. 1987. The microonamentation unities of the Oberhauchten of the dorsal scales in Vipera species (Reptilia: Viperidae). pp. 59-62. In: van Gelder, J.J., Strijbosch, H. & Bergers, P.J.M. (eds.). Proceedings of the 4th Ordinary General Meeting of the Societas Europaea Herpetologica. Societas Europaea Herpetologica, Faculty of Sciences, Nijmegen. 473 pp.

Bea, A.; Brana, F.; Baron, J.P.; Saint-Girons, H. 1992. Regimes et cycles alimentaires des viperes europeennes (Reptilia, Viperidae). Etude comparee. Annee Biologique 31(1): 25-44.

Bergman, J.; Norstrom, M. 1994. Some notes on the genus Vipera (Serpentes: Viperidae: Viperinae) in Anatolia, Turkey. Litteratura Serpentium (English Edition) 14(6): 166-175.

Boyer, J. 1945. Les vipères de France. Nature (Paris) 1941(3072): 257-260.

Brito, J.C.; Rebelo, A.; Crespo, E.G. 2001. Viper killings for superstitious reasons in Portugal. Boletin de la Asociacion Herpetologica Espanola 12(2): 101-104.

Bruno, S. 1985. Le vipere d'Italia e d'Europa. Edagricole, Bologna: V-Viii, 1-269.

Damme, R. van; Verheyen, R. 1987. Parameters in the thermal ecology of the common lizard, Lacerta vivipara. (Mistitled for: Thermal dependence of chemoreceptive predator-recognition inLacerta vivipara). pp. 421-424. In: van Gelder, J.J.; Strijbosch, H. & Bergers, P.J.M. (eds.). Proceedings of the 4th Ordinary General Meeting of the Societas Europaea Herpetologica. Societas Europaea Herpetologica, Faculty of Sciences, Nijmegen. 473 pp.

Denis, J. 1936. Observations sur l'altitude atteinte par les vipères dans les Pyrenées. Bulletin de la Société Zoologique de France 61: 404-405.

Duguy, R. 1972. Biologie des vipères dans les Pyrénées. Science et Nature (Paris) 1972(109): 9-16.

Duron, P.; Acolat, L. 1956. Le couleuvre verte et jaune (Zamenis viridiflavus viridiflavus Lacep.) peut contribuer a la destruction des vipères. Bull. Soc. Hist. nat. Doubs 58: 179-183.

Duron, P.; Acolat, L. 1957. La couleuvre verte et jaune (Zamenis viridiflavus (Lacep.) peut contribuer a la destruction des vipères. Ann. Sci. Univ. BesanYon (2) Zool. et Phys. 6: 179-183.

Ettling, J.A.; Aghasyan, A.L.; Aghasyan, L.A. 2015. The conservation of rare Armenian vipers Montivipera raddei and Pelias spp. pp. 81–88. In: Zoological Society of London (ed.). International Zoo Yearbook. Vol. 49. Reptile Conservation. Zoological Society of London, London. 442 pp.

Ferchaud, A.L.; Ursenbacher, S.; Cheylan, M.; Luiselli, L.M.; Jelic, D.; Halpern, B.; Major, A.; Kotenko, T.; Najme, K.; Behrooz, R.; Crnobrnja-Isailovic, J.; Tomovic, L.; Ghira, I.; Ioannidis, Y. 2012. Phylogeography of the Vipera ursinii complex (Viperidae): mitochondrial markers reveal an east-west disjunction in the Palaearctic region. Journal of Biogeography 39(10): 1836-1847.

Feuillee-Billot, A. 1929. Les ennemis naturels des vipères du Herisson a la couleuvre lisse. pp. 177-178. Nature, Paris.

Gasc, J.P. 1968. Morphologie des hemipenis chez Vipera ursinii ursinii (Bonaparte) et discussion biogeographique sur la repartition des espèces du genre Vipera en Europe occidentale. Bulletin du Muséum National d'Histoire Naturelle (Paris) 40: 95-101.

Grano, M.; Meier, G.; Cattaneo, C. 2017. Vipere Italiane – Gli ultimi studi sulla sistematica, l´ecologia e la storia naturale. Publisher unknown, Aicurzio. 197 pp.

Herrmann, H.W.; Joger, U.; Nilson, G. 1992. Molecular phylogeny and systematics of viperine snakes. 1. General phylogeny of European vipers (Vipera sensu stricto). pp. 219-224. In: Korsos, Z. & Kiss, I. (eds.): Proceedings of the 6th Ordinary General Meeting of the Societas Europaea Herpetologica, 19-23 August 1991, Budapest, Hungary. Hungarian Natural History Museum, Budapest. 531 pp.

Herrmann, H.W.; Joger, U.; Nilson, G.; Sibley, C.G. 1987. First steps towards a biochemically based reconstruction of the phylogeny of the genus Vipera. pp. 195-200. In: van Gelder, J.J., Strijbosch, H. & Bergers, P.J.M. (eds.). Proceedings of the 4th Ordinary General Meeting of the Societas Europaea Herpetologica. Societas Europaea Herpetologica, Faculty of Sciences, Nijmegen. 473 pp.

Istituto Erpetologico Italiano di Verona 1974. Vipère: un pericola crescente. Natura Alp. 1974(3): 164-170.

Jirousek, V. 1983. Systematic survey of adders of genus Vipera. Fauna Bohemiae Septentrionalis 8: 71-94.

Joger, U.; Herrmann, H.W.; Nilson, G. 1992. Molecular phylogeny and systematics of viperine snakes. 2. A revision of the Vipera ursinii complex. pp. 239-244. In: Korsos, Z. & Kiss, I. (eds.): Proceedings of the 6th Ordinary General Meeting of the Societas Europaea Herpetologica, 19-23 August 1991, Budapest, Hungary. Hungarian Natural History Museum, Budapest. 531 pp.

Joger, U.; Lenk, P.; Baran, I.; Böhme, W.; Ziegler, T.; Heidrich, P.; Wink, M. 1997. The phylogenetic position of Vipera barani and of V. nikolskii within the Vipera berus complex. pp. 185-194. In: Böhme, W.; Bischoff, W. & Ziegler, T. (eds.). Herpetologia Bonnensis. Proceedings of the 8th Ordinary General Meeting of the Societas Europaea Herpetologica, 23-27 August 1995, Bonn, Germany. Societas Europaea Herpetologica, Bonn. 414 pp.

Kaletzkaya, M.L. 1956. [Contribution to the biology of viper]. (In Russian). Priroda (Moscow) 5: 101-102.

Kalyabina, S.A.; Schweiger, S.; Joeger, U.; Mayer, W.; Orlov, N.L.; Wink, M. 2004. Phylogeny and systematics of adders (Vipera berus complex). Mertensiella 15: 7-15.

Keller-Zschokke, J.V. 1928. Von den Giftschlangen in Westlichen Schweizer Jura. A. Die Kreuzotter in den Kantonen Neuenburg und Waadt, pp. 83-87. B. Die Vipèrerie in Baulmes, Kanton Waadt. Mitt. Naturf. Ges. Solothurn 8: 88-93.

Kemier, L. 1945. Notes au sujet de l'hieroglyphe et des vipères dans l'Egypte ancienne. L. Keimer. Etud D'Egyptol. Le Caire Fasc. 7: 1-52.

Knoefel, P.K. 1991. Abati's work on the amazing nature of the viper and its miraculous powers. Toxicon 29(3): 301-309.

Leenders, T. 1994. The changing face of the genus Vipera. Litteratura Serpentium (English Edition) 14(5): 145-152.

Massia, G. 1928. Note sur l'habitat des vipères. Bull. Soc. linn. Lyon 7(5): 39.

Mizsei, E.; Jablonski, D.; Roussos, S.A.; Dimaki, M.; Ioannidis, Y.; Nilson, G.; Nagy, Z.T. 2017. Nuclear markers support the mitochondrial phylogeny of Vipera ursinii-renardi complex (Squamata: Viperidae) and species status for the Greek meadow viper. Zootaxa 4227(1): 75-88.

Naulleau, G. 1982. Action de la temperature sur la digestion chez les viperes espagnoles du genre Vipera. Publicaciones del Centro Pirenaico de Biologia Experimental 13: 89-94.

Naulleau, G. 1983. Action de la temperature sur la digestion chez cinq especes de viperes europeennes du genre Vipera. Bulletin de la Societe Zoologique de France 108(1): 47-58.

Nikolsky, A.M. 1910. De nova g. Viperae specie e Caucaso. Mitt. Kaukas. Mus. (Tiflis) 51: 81-84.

Nilson, G. 1998. Venomous snakes in west Asia: applicability of current species concepts. Russian Journal of Herpetology 5(2): 92-96.

Nilson, G.; Tuniyev, B.; Andren, C.; Orlov, N.L.; Joger, U.; Herrmann, H.W. 1999. Taxonomic position of the Vipera xanthina complex. Kaupia Darmstädter Beiträge Zur Naturgeschichte 8: 99-102.

Obst, F.J. 1983. Zur Kenntnis der Schlangengattung Vipera (Reptilia, Serpentes, Viperidae). Zoologische Abhandlungen (Dresden) 38(13) [1982]: 229-235.

Orth, K. 2002. Einige Vipern aus Europa im Terrarium. Reptilia (D) 7(33): 26-30.

Orth, K. 2002. European vipers in the terrarium. Reptilia (GB) 22: 25-29.

Phisalix, M. 1928. Capture des vipères. Bull. Soc. Hist. nat. du Doubs Besancon 1927(36): 43-45.

Phisalix, M. 1937. Les vipères au jardin du Roy et a l'academie des sciences. Bulletin de la Société Zoologique de France 62: 5-21.

Phisalix, M. 1969. La livree des vipères de France. Bulletin du Muséum National d'Histoire Naturelle (Paris) (Ser. 2) 40: 661-676.

Police, G. 1909. Caso di melanismo in una Vipera nel mezzogiorno d'Italia. Napoli Annuario Museo Zool. N.S. 3(6): 1-3.

Police, G. 1911. Due casi di morsicatura di Vipera. Napoli Boll. Soc. nat. 24 [1910]: 5-8.

Pozzi, A. 1985. Ma non sono tutte vipere. Natura Oggi 3(8): 124-135.

Redi, F. 1988. Francesco Redi on vipers. E.J. Brill, Leiden, New York etc.: I-Xviii, 1-86.

Reuss, T. 1927. Beschreibung neuer Vipern aus Jugoslawien. Zoologischer Anzeiger 71: 215-223.

Saint Girons, H. 1981. Le cycle des mues chez les viperes europeenes. Bulletin de la Societe Zoologique de France 105(4) [1980]: 551-559.

Saint Girons, H. 1982. Cycle annuel et survie de quelques viperes d'Europe. Influence des temperatures exceptionellement elevees de l'annee 1976. Vie et Milieu 31(1) [1981]: 59-64.

Saint Girons, H.; Duguy, R.; Detrait, J. 1984. Les viperes du sud du Massif Central: morphologie externe et venins. Bulletin de la Societe d'Histoire Naturelle de Toulouse 119 [1983]: 81-86.

Saint Girons, H.; Naulleau, G. 1981. Poids des nouveau-nes et strategies reproductrices des viperes europeennes. Terre et la Vie 35(4): 597-616.

Saint-Girons, H. 1975. Observations preliminaires sur la thermoregulation des vipères d'Europe. Vie et Milieu (C) 25(1): 137-168.

Saint-Girons, H. 1976. Les differentes types de cycles sexuels des males chez les vipères europeennes. Comptes Rendus Hebdomadaires des Seances Academie des Sciences (Paris) 282(10): 1017-1019.

Saint-Girons, H. 1978. Morphologie externe comparee et systematique des vipères d'Europe (Reptilia, Viperidae). Revue Suisse de Zoologie 85(3): 565-595.

Saint-Girons, H. 1978. Thermoregulation comparee des vipères d'Europe. Étude biotelemetrique. La Terre et la Vie (Paris) 32(3): 417-440.

Saint-Girons, H. 1979. Les cycles alimentaires des vipères europeennes dans des conditions semi-naturelles. Annales de Biologie Animale Biochimie Biophysique 19(1a): 125-134.

Schmidt, D. 1989. The snakes of Transcaucasia. Part 3. Eirenis, Telescopus and Vipera. Snake Keeper 3(1): 13-18.

Shine, R.; Madsen, T. 1994. Sexual dichromatism in snakes of the genus Vipera: a review and a new evolutionary hypothesis. Journal of Herpetology 28(1): 114-117.

Shiryaev, K.A. 2005. New data on reproductive biology of Caucasian species of the genus Vipera. Russian Journal of Herpetology 12: 213-215. (Supplement).

Starkov, V.G.; Osipov, A.V.; Utkin, Y.N. 2007. Toxicity of venoms from vipers of Pelias group to crickets Gryllus assimilis and its relation to snake entomophagy. Toxicon 49(7): 995-1001.

Stejneger, L.H. 1936. Types of the amphibian and reptilian genera proposed by Laurenti in 1768. Copeia 1936(3): 133-141.

Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Vanni, S.; Lanza, B. 1982. Note di erpetologia Italiana: Salamandra, Triturus, Rana, Phyllodactylus, Podarcis, Coronella, Vipera. Natura (Milano) 73(1-2): 3-22.

Zinenko, O.; Stümpel, N.; Mazanaeva, L.; Bakiev, A.; Shiryaev, K.; Pavlov, A.; Kotenko, T.; Kukushkin, O.; Chikin, Y.; Duisebayeva, T.; Nilson, G.; Orlov, N.L.; Sako, T.; Ananjeva, N.B.; Murphy, R.W.; Joger, U. 2015. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi). Molecular Phylogenetics and Evolution 84: 85-100.

Vipera altaica

Tuniyev, B.; Nilson, G.; Andrén, C. 2010. A new species of viper (Reptilia, Viperidae) from the Altay and Saur Mountains, Kazakhstan. Russian Journal of Herpetology 17(2): 110-120.

Zinenko, O.; Stümpel, N.; Mazanaeva, L.; Bakiev, A.; Shiryaev, K.; Pavlov, A.; Kotenko, T.; Kukushkin, O.; Chikin, Y.; Duisebayeva, T.; Nilson, G.; Orlov, N.L.; Sako, T.; Ananjeva, N.B.; Murphy, R.W.; Joger, U. 2015. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi). Molecular Phylogenetics and Evolution 84: 85-100.

Vipera ammodytes

Afsar, M.; Cicek, K.; Dincaslan, Y.E.; Ayaz, D.; Tok, C.V. 2013. New record localities of five snake species in Turkey. Herpetozoa 25(3-4): 179-183.

Ajtic, R.D. 2008. [Nose-horned viper (Vipera ammodytes) conservation problems in Serbia]. (In Serbian). Zashtita Prirode 60(1-2): 319-326.

Akkaya, A.; Ugurtas, I.H. 2012. Rediscovery of Vipera ammodytes (Linnaeus, 1758) at Uludag-Bursa, Turkey after 62 years. Herpetozoa 24(3-4): 181-185.

Arikan, H.; Göcmen, B.; Mermer, A.; Bahar, H. 2005. An electrophoretic comparison of the venoms of a colubrid and various viperid snakes from Turkey and Cyprus, with some taxonomic and phylogenetic implications. Zootaxa 1038: 1-10.

Arikan, H.; Kumlutas, Y.; Türkozan, O.; Baran, I. 2003. Electrophoretic patterns of some viper venoms from Turkey. Turkish Journal of Zoology 27(3): 239-242.

Baran, I. 1978. Some rare species of snakes from Turkey. Annalen des Naturhistorischen Museums in Wien 81 [1977]: 261-265.

Beek, H. van de 1991. Vipera ammodytes ammodytes - sandviper. Litteratura Serpentium (English Edition) 11(6): 144.

Beshkov, V.A. 1977. Über die Biologie und Ökologie der Schlangen im Maleshevskigebirge (Südwest-Bulgarien). 3. Nahrung und Fortpflanzung der Sandotter Vipera ammodytes meridionalis Boulenger). Ekologiya (Sofia) 3: 3-12.

Beshkov, V.A.; Nankinov, D.N. 1979. Materialen über die Ornithophagia der Schlangen im Gebirge 'Maleschevska' (Südwestbulgarien). Ekologiya (Sofia) 5: 55-63.

Biella, H.J. 1983. Die Sandotter Vipera ammodytes. Neue Brehm-Bücherei 558: 1-84.

Biella, H.J. 1983. Intermediäre und abnorme Sandvipern aus der Herpetologischen Sammlung des Staatlichen Museums für Tierkunde Dresden (Reptilia: Serpentes, Viperidae). Zoologische Abhandlungen (Dresden) 38(14) [1982]: 237-245.

Biella, H.J. 1984. Zum Erbgang qualitativer Merkmale bei der Kreuzung von Vipera ammodytes ammodytes mit Vipera ammodytes montandoni (Reptilia, Serpentes, Viperidae). Zoologische Abhandlungen (Dresden) 39(7) [1983]: 99-105.

Biella, H.J. 1995. Die Sandotter. (Reprint). VerlagsKG Wolf (formerly Westarp Wissenschaften), Magdeburg. 108 pp. (Die Neue Brehm-Bücherei 558).

Biella, H.J.; Blattler, E. 1989. Problematik der unterartlichen Eingliederung nord-und südwestlicher Vipera ammodytes - Populationen, dargestellt am Biespiel der südslowenischen Sandviper (Reptilia, Serpentes: Viperidae). Zoologische Abhandlungen (Dresden) 45(2): 11-22.

Biella, H.J.; Blattler, E. 1992. Zur geographischen Variation eidonomischer Merkmale bei der Sandviper (Vipera ammodytes). Mitteilungen aus dem Zoologischen Museum in Berlin 68(2): 249-258.

Bogdanov, O.P. 1971. New data on the reproduction of horned viper, Vipera ammodytes transcaucasiana Boulenger. Soobshcheniya Akademii Nauk Gruzinskoi SSR 64(3): 711-712.

Bogoescu, C. 1933. Vipera ammodytes in Jud. Bacau. Bull. Soc. Nat. Romania Bucuresti 3: 7-8.

Bolkay, S.J. 1923. On a case of cannibalism among Vipera ammodytes L. Glasn. Hrv. Pris. Drust. Zagreb 35: 16.

Boulenger, G.A. 1903. On the geographical variations of the sand-viper, Vipera ammodytes. Proceedings of the Zoological Society of London 1: 185-186.

Boulenger, G.A. 1904. On the sandviper of Roumania (Vipera ammodytes, var. montandoni). Annals and Magazine of Natural History 14: 134-135.

Boulenger, G.A. 1913. On the geographical races of Vipera ammodytes. Annals and Magazine of Natural History (London) 11: 283-287.

Bozhansky, A.T.; Kudryavtsev, S.V. 1986. [The sand viper Vipera ammodytes]. (In Russian). Priroda (Moscow) 1986(2): 46-47.

Bozhansky, A.T.; Kudryavtsev, S.V. 1986. Ecological observations of the rare vipers of the Caucasus. pp. 495-498. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European herpetological meeting (3rd ordinary general meeting of the Societas Europaea Herpetologica), Prague 1985. Charles University, Prague. 754 pp.

Bringsoe, H. 1985. Remarks on: 'Some amphibians and reptiles of Corfu with a special note on the occurrence of the nose-horned viper (Vipera ammodytes)' (Stafford P. The Herptile, 9(3) Sept 1984 pp. 105-108). Herptile 10(1): 23-24.

Bruno, S. 1967. Sulla Vipera ammodytes (Linnaeus 1758) in Italia. Memorie del Museo Civico di Storia Naturale di Verona 15: 289-336.

Buchholz, K.F.; Schultze-Westrum, T. 1964. Zur Kenntnis der Schlangenfauna der nördlichen Sporaden. Zoologischer Anzeiger 173: 127-136.

Calinescu, R.I. 1926. Vipera ammodytes Cuv. in Romania. Trav. Inst. Geograph. Univ. Cluj. Roumanie 2 [1924-1925]: 261-290.

Carruccio, A. 1909. Esemplari di Vipera ammodytes del Montenegro donati al Museo zoologico di Roma da S. M. la Regina Elena. Roma Boll. Soc. zool. ital. 10: 278-286.

Cattaneo, A. 1997. L'erpetofauna dell'isola greca di Skiathos (Sporadi settentrionali). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 136(2): 145-156.

Cattaneo, A. 1999. Gli anfibi e i rettili delle isole greche di Skyros, Skopelos e Alonissos (Sporadi settentrionali). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 139(2): 127-149.

Cattaneo, A. 2010. Note eco-morfologiche su alcune species ofidiche egee, con particolare riferimento alle popolazioni delle cicladi centro-orientali (Reptilia). Naturalista Siciliano 34(3-4): 319-350.

Christov, K.; Beshkov, V.A. 1999. On the subspecies morphological characteristics of the sandvipers (Vipera ammodytes) from different locations in Bulgaria. Acta Zoologica Bulgarica 51(2-3): 61-67.

Christov, K.; Matev, I.; Iliev, Y. 1997. Major characteristics of three Vipera ammodytes subspecies from laboratory conditions. Folia Zoologica 46(3): 286-288.

Clark, R.J. 1967. Centipede in stomach of young Vipera ammodytes meridionalis. Copeia 1967: 224.

Clark, R.J. 1968. Aussergewöhnliche Farbung und Zeichnung bei Vipera ammodytes mendionalis von griechischen Inseln. Salamandra 4: 69-72.

Crnobrnja-Isailovic, J.M. 2002. Notes of diurnal activity in Vipera ammodytes of the central Balkans. Biota (Race) 3(1-2): 9-16.

Crnobrnja-Isailovic, J.M.; Ajtic, R.D.; Tomovic, L.M. 2007. Activity patterns of the sand viper (Vipera ammodytes) from the central Balkans. Amphibia-Reptilia 28(4): 582-589.

Cubric, T.; Crnobrnja-Isailovic, J. 2017. Malformation of eye pupil in nose-horned viper (Vipera ammodytes) from a wild population. Herpetology Notes 10: 229-231.

Cundall, D. 2002. Envenomation strategies, head form and feeding ecology in vipers. pp. 149-161. In: Schuett, G.W.; Höggren, M.; Douglas, M.E.; Greene, H.W. (eds.). Biology of the vipers. Eagle Mountain Publishing, Eagle Mountain. 580 pp.

Debogovic, Z.; Heisinger Puric, A. 1981. Seasonal variations of the toxicity of Vipera ammodytes venom from the vicinity of Slunj over a five-year period. Veterinarski Arhiv 51 (Suppl.): 34-36.

Dobrilla, R.; Dolce, S. 1996. Studio morfologico delle popolazioni di Vipera ammodytes (L., 1758) di Friuli, Venezia-Giulia, Istria e Dalmazia. Atti del Museo Civico di Storia Naturale di Trieste 47: 285-300.

Eiselt, J.; Baran, I. 1970. Ergebnisse zoologischer Sammelreisen in der Türkei: Viperidae. Annalen des Naturhistorischen Museums in Wien 74: 357-369.

Esterbauer, H. 1987. Pflege und Nachzucht der westlichen Sand- oder Hornotter Vipera ammodytes (Linnaeus, 1758). Aquaria (St. Gallen) 34(9): 143-147.

Faoro, G. 1986. Bemerkungen zur Verbastardierung von Vipera ammodytes ammodytes mit Vipera aspis atra. Herpetofauna (Weinstadt) 8(43): 6-7.

Fuhn, I.E. 1986. [Melanism and head abnormalities in a specimen of Vipera ammodytes ammodytes (L., 1758) (Viperidae, Serpentes, Reptilia) in the surroundings of Resita]. (In Romanian). Studii Si Cercetari de Biologie Seria Biologie Animala 38(1): 7-10.

Georgescu, B.; Salajan, G.H.; Georgescu, D.; Mierlita, D.; Neagoe, E. 1997. Preliminary studies on the lethal action of the venom from Vipera ammodytes depending on the age. Buletinul Universitatii de Stiinte Agricole si Medicina Veterinara Cluj Napoca Seria Zootehnie si Biotehnologii si Medicina Veterinara 51(1): 15-18.

Georgescu, B.; Salajan, G.H.; Iozon, D.; Mierlita, D.; Odagiu, A. 2000. Cercetari privind evolutia consumului voluntar de hrana pe durata unui sezon activ la specia Vipera ammodytes in conimtiile alimentarii cu soareci albi de laborator. [Studies on voluntary food ingestion of white laboratory mice in Vipera ammodytes snakes during an active season]. (In Romanian). pp. 247-251. In: Man, C. (ed.). Simpozion National "Realizari si Perspective in Zootehnie si Biotehnologii": lucrari stiintifice: Vol. XXVI (1) Cluj-Napoca, 17-18 noiembrie 2000. [National symposium "Achievements and Expectations in Animal Husbandry and Biotechnologies": scientific debates: Vol.XXVI (1), Cluj-Napoca, 17-18 November 2000]. Editura AcademicPres, Cluj-Napoca. 259 pp.

Georgescu, B.; Salajan, G.H.; Iozon, D.; Mierlita, D.; Odagiu, A. 2003. Studies on the length and quality of digestion of the species Vipera ammodytes. Buletinul Universitatii de Stiinte Agricole si Medicina Veterinara Cluj Napoca Seria Zootehnie si Biotehnologii 59: 240-243.

Georgescu, B.; Salajan, G.H.; Mierlita, D.; Iozon, D.; Odagiu, A. 1999. Studies concerning the variations in the digestibility of food in the ammodytes ammodytes viper, in considering the used diet resource (mice, chicken, cattle meat). Buletinul Universitatii de Stiinte Agricole si Medicina Veterinara Cluj Napoca Seria Zootehnie si Biotehnologii si Medicina Veterinara 53: 37-40.

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Vipera anatolica

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Vipera aspis

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Bonnet, X.; Naulleau, G.; Lourdais, O. 2002. Benefits of complementary techniques: using capture-recapture and physiological approaches to understand costs of reproduction in the aspic viper (Vipera aspis). pp. 483-495. In: Schuett, G.W.; Höggren, M.; Douglas, M.E.; Greene, H.W. (eds.). Biology of the vipers. Eagle Mountain Publishing, Eagle Mountain. 580 pp.

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Caputo, V.; Guarino, F.M. 1992. L'erpetofauna del Cilento. Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano 132(22): 273-292.

Carlino, P.; Pauwels, T.C.P.; Pauwels, O.S.G. 2017. Natural history notes: Vipera aspis hugyi (South-Italian Asp Viper). Diet. Herpetological Review 48(1): 221.

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Castanet, J.; Naulleau, G. 1985. La squelettochronologie chez les reptiles. 2. Resultats experimentaux sur la signification des marques de croissance squelettiques chez les serpents. Remarques sur la croissance et la longevite de la vipere aspic. Annales des Sciences Naturelles Zoologie et Biologie Animale 7(1): 41-62.

Castella, B.; Golay, J.; Monney, J.C.; Golay, P.; Mebert, K.; Dubey, S. 2013. Melanism, body condition and elevational distribution in the asp viper. Journal of Zoology (London) 290(4): 273-280.

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Vipera barani

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Vipera berus

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Bednarz, Z. 2002. Swiat gadow w zyciu Lasowiakow - jak wykorzystac w ochronie przyrody strach przed zmijami. [The world of reptiles in the life of Lasowiaks: how to make the most of the fear of vipers for nature conservation]. (In Polish, English summary). Przeglad Przyrodniczy 13(4): 153-157.

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Vipera darevskii

Agasyan, L.A.; Agasyan, A.L. 2008. New data on the distribution and conservation of Darevsky’s Viper, (Vipera darevskii Vedmederja, Orlov et Tuniyev, 1986). pp. 7-10. In: Ananjeva, N.B.; Danilov, I.G.; Dunayev, E.A.; Ischenko, V.G.; Lada, G.A.; Litvinchuk, S.N.; Orlova, V.F.; Smirina, E.M.; Tuniyev, B.S. & Khalikov, R.G. (eds.). The problems of herpetology. Proceedings of the 3rd meeting of the Nikolsky Herpetological Society, 9-13 October 2006, Putschino. Russian Academy of Sciences, St. Petersburg. 467 pp.

Avci, A.; Ilgaz, C.; Baskaya, S.; Baran, I.; Kumlutas, Y. 2010. Contribution to the distribution and morphology of Pelias darevskii (Vedmederja, Orlov et Tuniyev 1986) (Reptilia: Squamata: Viperidae) in northeastern Anatolia. Russian Journal of Herpetology 17(1): 1-7.

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Orlov, N.L.; Tuniyev, B.S. 1986. The recent areas, their possible genesis and the phylogeny of three viper species of Eurosiberian group of the Vipera kaznakowi complex in the Caucasus. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 157: 107-135.

Orlov, N.L.; Tuniyev, B.S. 1990. Three species in the Vipera kaznakowi complex (Eurosiberian group) in the Caucasus: their present distribution, possible genesis, and phylogeny. Asiatic Herpetological Research 3: 1-36.

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Tuniyev, B.S; Avci, A.; Ilgaz, C.; Olgun, K.; Petrova, T.V.; Bodrov, S.Y.; Geniez, P.; Teynié, A. 2018. On taxonomic status of Shield-head Vipers from Turkish Lesser Caucasus and East Anatolia. Proceedings of the Zoological Institute RAS 322(1): 3–44.

Tuniyev, S.B.; Iremashvili, G.; Heras, B. de las; Tuniyev, B. 2014. About type locality and finds of Darevsky’s Viper [Pelias darevskii (Vedmederja, Orlov et Tuniyev, 1986), Reptilia: Viperinae] in Georgia. Russian Journal of Herpetology 21(4): 281-290.

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Vipera dinniki

Orlov, N.L.; Tuniyev, B.S. 1986. The recent areas, their possible genesis and the phylogeny of three viper species of Eurosiberian group of the Vipera kaznakowi complex in the Caucasus. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 157: 107-135.

Orlov, N.L.; Tuniyev, B.S. 1990. Three species in the Vipera kaznakowi complex (Eurosiberian group) in the Caucasus: their present distribution, possible genesis, and phylogeny. Asiatic Herpetological Research 3: 1-36.

Orlov, N.L.; Tuniyev, B.S. 2005. Vipera dinniki - Dinniks Kaukasusotter. pp. 311-323. In: Joger, U. & Stümpel, N. (eds.). Handbuch der Reptilien und Amphibien Europas. Schlangen III. Viperidae. AULA-Verlag, Wiebelsheim. 420 pp.

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Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Tuniyev, B.S.; Volcik, S. 1995. On placing and thermobiology of polymorphic population of Vipera dinniki (Nikolsky, 1913) (Serpentes, Viperidae) at uppermost Msymta-river valley. pp. 244-249. In: Llorente, G.A.; Montori, A.; Santos, X. & Carretero, M.A. (eds.). Scientia Herpetologica: papers submitted from the 7th Ordinary General Meeting of Societas Europaea Herpetologica, Barcelona, September 15-19, 1993. Asociacion Herpetologica Espanola, Barcelona. 385 pp.

Tzarewsky, S.T.; Tzarewsky, S.F. 1916. Formes nouvelles du genre Vipera, trouvees dans l'Empire Russe. Ann. Mus. Zool. Petrograd 21: 34-39.

Vedmerja, V.J.; Orlov, N.L.; Tuniyev, B.S. 1986. On taxonomy of three viper species of the Vipera kaznakowi complex. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 157: 55-61.

Vipera ebneri

Dely, O.G.; Stohl, G. 1989. Ergänzende Angaben zur Diagnose der Unterart Vipera ursinii ebneri Knoepffler et Sochurek, 1955 (Serpentes, Viperidae). Vertebrata Hungarica 23: 21-27.

Hutwalker, A. 2017. Herpetologische Eindrücke einer Wanderreise im zentralen Elbursgebirge, Nordiran. Sauria (Berlin) 39(3): 25-30.

Nilson, G.; Andrén, C. 2001. The meadow and steppe vipers of Europe and Asia: the Vipera (Acridophaga) ursinii complex. Acta Zoologica Academiae Scientiarum Hungaricae 47(2-3): 87-267.

Rajabizadeh, M.; Nilson, G.; Kami, H.G.; Naderi, A.R. 2011. Distribution of the subgenus Acridophaga Reuss, 1927 (Serpentes: Viperidae) in Iran. Iranian Journal of Animal Biosystematics 7(1): 83-87.

Zinenko, O.; Stümpel, N.; Mazanaeva, L.; Bakiev, A.; Shiryaev, K.; Pavlov, A.; Kotenko, T.; Kukushkin, O.; Chikin, Y.; Duisebayeva, T.; Nilson, G.; Orlov, N.L.; Sako, T.; Ananjeva, N.B.; Murphy, R.W.; Joger, U. 2015. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi). Molecular Phylogenetics and Evolution 84: 85-100.

Vipera eriwanensis

Arikan, H.; Göcmen, B.; Mermer, A.; Bahar, H. 2005. An electrophoretic comparison of the venoms of a colubrid and various viperid snakes from Turkey and Cyprus, with some taxonomic and phylogenetic implications. Zootaxa 1038: 1-10.

Baran, I.; Tok, C.V.; Olgun, K.; Iret, F.; Avci, A. 2005. On viperid (Serpentes: Sauria) specimens collected from Northeastern Anatolia. Turkish Journal of Zoology 29(3): 225-228.

Gvozdik, V.; Jandzik, D.; Cordos, B.; Rehak, I.; Kotlik, P. 2012. A mitochondrial DNA phylogeny of the endangered vipers of the Vipera ursinii complex. Molecular Phylogenetics and Evolution 62(3): 1019-1024.

Krecsák, L. 2007. Re-description of the holotype of Vipera eriwanensis (Reuss, 1933) (Serpentes: Viperidae). Revue Suisse de Zoologie 114(3): 655-662.

Kukushkin, O.V.; Iskenderov, T.; Axmedov, S.; Bunyatova, S.N.; Zinenko, O.I. 2012. Additions to the distribution of Vipera eriwanensis (Serpentes: Viperidae) in Transcaucasia, with comments on the identity of vipers in northeastern Azerbaijan. Herpetology Notes 5: 423-427.

Kutrup, B.; Bülbül, U.; Yilmaz, N. 2005. On the distribution and morphology of the steppe viper, Vipera eriwanensis (Reuss, 1933), from Gavur Mountain (Gümüshane). Turkish Journal of Zoology 29: 321-325.

Rajabizadeh, M.; Nilson, G.; Kami, H.G.; Naderi, A.R. 2011. Distribution of the subgenus Acridophaga Reuss, 1927 (Serpentes: Viperidae) in Iran. Iranian Journal of Animal Biosystematics 7(1): 83-87.

Reuss, F.A.T. 1935. Fortsetzung und Schluss der Originalberichte. Nach. Bl. Aq. Terr. Berlin 1933: 372-373.

Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Zinenko, O.; Stümpel, N.; Mazanaeva, L.; Bakiev, A.; Shiryaev, K.; Pavlov, A.; Kotenko, T.; Kukushkin, O.; Chikin, Y.; Duisebayeva, T.; Nilson, G.; Orlov, N.L.; Sako, T.; Ananjeva, N.B.; Murphy, R.W.; Joger, U. 2015. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi). Molecular Phylogenetics and Evolution 84: 85-100.

Vipera graeca

Korsos, Z.; Barina, Z.; Pifko, D. 2008. First record of Vipera ursinii graeca in Albania (Reptilia: Serpentes, Viperidae). Acta Herpetologica 3(2): 167-173.

Mizsei, E.; Balint, U. 2012. Novel defensive behaviours of both sexes of Vipera ursinii graeca (Serpentes: Viperidae). Herpetology Notes 5: 481-483.

Mizsei, E.; Jablonski, D.; Roussos, S.A.; Dimaki, M.; Ioannidis, Y.; Nilson, G.; Nagy, Z.T. 2017. Nuclear markers support the mitochondrial phylogeny of Vipera ursinii-renardi complex (Squamata: Viperidae) and species status for the Greek meadow viper. Zootaxa 4227(1): 75-88.

Mizsei, E.; Üveges, B.; Vági, B.; Szabolcs, M.; Lengyel, S.; Pfliegler, W.P.; Nagy, Z.T.; Tóth, J.P. 2016. Species distribution modelling leads to the discovery of new populations of one of the least known European snakes, Vipera ursinii graeca, in Albania. Amphibia-Reptilia 37(1): 55-68.

Nilson, G.; Andrén, C. 1988. A new subspecies of the subalpine meadow viper, Vipera ursinii (Bonaparte) (Reptilia, Viperidae), from Greece. Zoologica Scripta 17(3): 311-314.

Zinenko, O.; Stümpel, N.; Mazanaeva, L.; Bakiev, A.; Shiryaev, K.; Pavlov, A.; Kotenko, T.; Kukushkin, O.; Chikin, Y.; Duisebayeva, T.; Nilson, G.; Orlov, N.L.; Sako, T.; Ananjeva, N.B.; Murphy, R.W.; Joger, U. 2015. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi). Molecular Phylogenetics and Evolution 84: 85-100.

Vipera kaznakovi

Afsar, M.; Afsar, B. 2009. A new locality for Vipera (Pelias) kaznakovi Nikolsky, 1909 (Reptilia, Viperidae) in the North-Eastern Anatolia. Russian Journal of Herpetology 16(2): 155-158.

Arikan, H.; Göcmen, B.; Mermer, A.; Bahar, H. 2005. An electrophoretic comparison of the venoms of a colubrid and various viperid snakes from Turkey and Cyprus, with some taxonomic and phylogenetic implications. Zootaxa 1038: 1-10.

Arikan, H.; Kumlutas, Y.; Türkozan, O.; Baran, I. 2003. Electrophoretic patterns of some viper venoms from Turkey. Turkish Journal of Zoology 27(3): 239-242.

Baran, I.; Tok, C.V.; Olgun, K.; Iret, F.; Avci, A. 2005. On viperid (Serpentes: Sauria) specimens collected from Northeastern Anatolia. Turkish Journal of Zoology 29(3): 225-228.

Bozhansky, A.T. 1984. [The viper Vipera kaznakowi]. (In Russian). Priroda (Moscow) 1984(6): 53-55.

Bozhansky, A.T.; Kudryavtsev, S.V. 1986. An experience of Caucasian viper breeding in captivity. Vestnik Zoologii 1986(3): 78-81.

Bozhansky, A.T.; Kudryavtsev, S.V. 1986. Ecological observations of the rare vipers of the Caucasus. pp. 495-498. In: Rocek, Z. (ed.). Studies in herpetology. Proceedings of the European herpetological meeting (3rd ordinary general meeting of the Societas Europaea Herpetologica), Prague 1985. Charles University, Prague. 754 pp.

Chiszar, D.; Brown, J.; Chiszar, A.; Smith, H.M. 1999. Strike-induced chemosensory searching in the Caucasus viper (Vipera kaznakovi) and the ocellate mountain viper (Vipera wagneri). Bulletin of the Maryland Herpetological Society 35(3): 85-91.

Dostal, I.; Duska, S.; Koropecky, I. 1979. [The snake Vipera kaznakowi]. (In Czech). Ziva 27(3): 106-107.

Göcmen, B.; Cicek, K.; Igci, N.; Akman, B. 2013. Data on the reproduction of a Caucasian Viper, Vipera kaznakovi Nikolsky, 1909 (Serpentes: Viperidae) from Hopa (northeastern Anatolia, Turkey). Ecologia Balkanica 5(2): 99-101.

Gül, S.; Kumlutas, Y.; Ilgaz, C. 2016. Predicted distribution patterns of Pelias kaznakovi (Nikolsky, 1909) in the Caucasus hotspot with a new locality record from Turkey. Russian Journal of Herpetology 23(3): 224-230.

Kramer, S. 1961. Variation, Sexualdimorphismus, Wachstum und Taxionomie von Vipera ursinii (Bonaparte, 1835) und Vipera kaznakovi Nikolskij, 1909. Revue Suisse de Zoologie 68: 627-725.

Kretz, J. 1971. Über Vipera kaznakovi Nikolskij 1909 aus Nordostanatolien (Reptilia, Viperidae). Jahrbuch Naturh. Mus. Bern 4 [1969-1971]: 125-134.

Mamet, S.; Kudryavtsev, S.V. 1997. Caucasian viper: husbandry and reproduction in captivity. Russian Journal of Herpetology 4(2): 198-199.

Norström, M. 1988. The Caucasian viper in Turkey. Fauna Och Flora (Stockholm) 83(1): 26-33.

Orlov, N.L.; Tuniyev, B.S. 1986. The recent areas, their possible genesis and the phylogeny of three viper species of Eurosiberian group of the Vipera kaznakowi complex in the Caucasus. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 157: 107-135.

Orlov, N.L.; Tuniyev, B.S. 1990. Three species in the Vipera kaznakowi complex (Eurosiberian group) in the Caucasus: their present distribution, possible genesis, and phylogeny. Asiatic Herpetological Research 3: 1-36.

Orlov, N.L.; Tuniyev, B.S. 2005. Vipera kaznakovi - Gemeine Kaukasusotter. pp. 325-341. In: Joger, U. & Stümpel, N. (eds.). Handbuch der Reptilien und Amphibien Europas. Schlangen III. Viperidae. AULA-Verlag, Wiebelsheim. 420 pp.

Stumpel, A.; Podloucky, R.; Corbett, K.; Andren, C.; Bea, A.; Nilson, G.; Oliveira, M.E. 1992. Threatened reptiles in Europe requiring special conservation measures. pp. 25-34. In: Korsos, Z. & Kiss, I. (eds.). Proceedings of the 6th Ordinary General Meeting of the Societas Europaea Herpetologica, 19-23 August 1991, Budapest, Hungary. Hungarian Natural History Museum, Budapest. 531 pp.

Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Vedmederya, V.I. 1984. [Distribution, variation and ecological features of Vipera kaznakowi]. pp. 8-9. (In Russian). In: Ishchenko, V.G. (ed.). [The species and its productivity within its distribution range. Part 5. Questions of herpetology. Material from the 4th All-Union Symposium 3-7 April 1984]. Urals Scientific Centre, Academy of Sciences USSR, Sverdlovsk. 54 pp.

Vedmerja, V.J.; Orlov, N.L.; Tuniyev, B.S. 1986. On taxonomy of three viper species of the Vipera kaznakowi complex. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR 157: 55-61.

Zinyakova, M.P.; Trofimov, A.G. 1977. [On the biology of reproduction in the Caucasian viper Vipera kasnakovi Nik.]. pp. 4-5. In: Darevsky, I.S. [Fourth All-Union Herpetological Conference. Questions in Herpetology]. Akademiya Nauk SSSR, Zoologicheskij Institut. Izdatel'stvo 'Nauka', Leningrad. 243 pp. (In Russian).

Vipera latastei

Alonso-Zarazaga, M.A. 2013. Nomenclatural notes. Graellsia 69(1): 129-131.

Amat, F. 2004. Analysis of the morphological variation of Vipera latastei: Taxonomic and biogeographic implications. Russian Journal of Herpetology 11(3): 198-202.

Bernis, F. 1969. La culebra de las Islas Columbretes: Vipera latastei. Boletin R. Soc. Esp. Hist. nat. (Sec. Biol.) 66: 115-133.

Brito, J.C. 2001. A record of melanism in Vipera latasti. Herpetological Bulletin 76: 28-29.

Brito, J.C. 2003. Seasonal and daily activity patterns of Vipera latastei in northern Portugal. Amphibia-Reptilia 24(4): 497-508.

Brito, J.C. 2003. Seasonal variation in movements, home range, and habitat use by male Vipera latastei in northern Portugal. Journal of Herpetology 37(1): 155-160.

Brito, J.C. 2004. Feeding ecology of Vipera latastei in northern Portugal: Ontogenetic shifts, prey size and seasonal variations. Herpetological Journal 14(1): 13-19.

Brito, J.C.; Alvares, F.J. 2004. Patterns of road mortality in Vipera latastei and V. seoanei from northern Portugal. Amphibia-Reptilia 25(4): 459-465.

Brito, J.C.; Crespo, E.G. 2002. Distributional analysis of two vipers (Vipera latastei and V. seoanei) in a potential area of sympatry in the northwestern Iberian Peninsula. pp. 129-138. In: Schuett, G.W.; Höggren, M.; Douglas, M.E.; Greene, H.W. (eds.). Biology of the vipers. Eagle Mountain Publishing, Eagle Mountain. 580 pp.

Brito, J.C.; Fahd, S.; Martínez-Freiria, F.; Tarroso, P.; Larbes, S.; Pleguezuelos, J.M.; Santos, X. 2011. Climate change and peripheral populations: predictions for a relict Mediterranean viper. Acta Herpetologica 6(1): 105-118.

Brito, J.C.; Rebelo, R. 2003. Differential growth and mortality affect sexual size dimorphism in Vipera latastei. Copeia 2003(4): 865-871.

Brito, J.C.; Santos, X.; Pleguezuelos, J.M.; Fahd, S.; Llorente, G.A.; Parellada, X. 2006. Morphological variability of the Lataste's viper (Vipera latastei) and the Atlas dwarf viper (Vipera monticola): patterns of biogeographical distribution and taxonomy. Amphibia-Reptilia 27(2): 219-240.

Busack, S.D.; Salvador, A. 1984. Nuevos datos sobre la distribucion de cuatro epsecies de reptiles en la provincia de Cadiz. Doñana Acta Vertebrata 11(2): 322-325.

Damme, R. van; Castilla, A.M. 1996. Chemosensory predator recognition in the lizard Podarcis hispanica: effects of predation pressure relaxation. Journal of Chemical Ecology 22(1): 13-22.

Fahd, S.; Barata, M.; Benitez, M.; Brito, J.C.; Caro, J.; Carvalho, Silvia; Chirosa, M.; Feriche, M.; Herrera, T.; Marquez-Ferrando, R.; Nesbitt, D.; Pleguezuelos, J.M.; Reques, R.; Paz, R. 2007. Presencia de la vibora hocicuda Vipera latastei en el Atlas Medio (Marruecos) y otras citas herpetologicas para la region. Boletin de la Asociacion Herpetologica Espanola 18: 26-34.

Fahd, S.; Benitez, M.; Brito, J.C.; Caro, J.; Chirosa, M.; Feriche, M.; Fernandez-Cardenete, J.R.; Martinez-Freiria, F.; Marquez-Ferrando, R.; Nesbitt, D.; Pleguezuelos, J.M.; Reques, R.; Rodriguez, M.P.; Santos, X.; Sicilia, M. 2006. Distribucion de Vipera latasti en el Rif y otras citas herpetologicas para el norte de Marruecos. Boletin de la Asociacion Herpetologica Espanola 16(1-2): 19-25.

Ferreira, D.; Mateus, C.; Santos, X. 2016. Responses of reptiles to fire in transition zones are mediated by bioregion affinity of species. Biodiversity and Conservation 25(8): 1543-1557.

Freitas, I.; Fahd, S.; Velo-Antón, G.; Martínez-Freiría, F. 2018. Chasing the phantom: biogeography and conservation of Vipera latastei-monticola in the Maghreb (North Africa). Amphibia-Reptilia 39(2): 145–161.

Gragera Ramos, S.; Fernandez Cardenete, J.R. 2013. Un caso de comportamiento arborícola en Vipera latastei. Boletin de la Asociacion Herpetologica Espanola 24(1): 45-47.

Hopkins, P.W. 1974. Sobre la herpetofauna de la Sierra de Estrella (Portugal) con especial referencia a Coronella austriaca austriaca y Vipera latasti. Doñana Acta Vertebrata 1(1): 11-17.

Lopez-Jurado, L.F.; Ruiz Caballero, M. 1981. Predacion de Vipera latastei sobre Mustela nivalis. Donana Acta Vertebrata 8: 298-299.

Loureiro, A. 2007. Cabecas de vibora - Köpfe von Vipera latastei als Handelsgut in Portugal. Zeitschrift für Feldherpetologie 14(1): 77-86.

Malkmus, R. 1997. Slangen van Portugal (10): de Wipneusadder (Vipera latasti). Lacerta 56(2) [1997-1998]: 44-48.

Martinez, F.; Brito, J.C.; Lizana Avia, M. 2006. Intermediate forms and syntopy among vipers (Vipera aspis and V. latastei) in northern Iberian Peninsula. Herpetological Bulletin 97: 14-18.

Martinez-Freiria, F.; Brito, J.C.; Lizana-Avia, M. 2006. Ophiophagy and cannibalism in Vipera latastei Bosca, 1878 (Reptilia, Viperidae). Herpetological Bulletin 96: 26-28.

Martínez-Freiria, F.; Lizana, M.; Amaral, J.P. do; Brito, J.C. 2010. Spatial and temporal segregation allows coexistence in a hybrid zone among two Mediterranean vipers (Vipera aspis and V. latastei). Amphibia-Reptilia 31(2): 195-212.

Martínez-Freiria, F.; Pardavila, X.; Lamosa, A. 2012. [A new case of melanism in Vipera latastei]. (In Spanish). Boletin de la Asociacion Herpetologica Espanola 23(1): 51-54.

Martínez-Freiría, F.; Santos, X. 2015. Assessing the heritability of dorsal pattern shape in Vipera latastei. Amphibia-Reptilia 36(3): 313-317.

Martinez-Freiria, F.; Santos, X.; Pleguezuelos, J.M.; Lizana, M. 2009. Geographical patterns of morphological variation and environmental correlates in contact zones: a multi-scale approach using two Mediterranean vipers (Serpentes). Journal of Zoological Systematics and Evolutionary Research 47(4): 357-367.

Martinez-Freiria, F.; Sillero, N.; Lizana, M.; Brito, J.C. 2008. GIS-based niche models identify environmental correlates sustaining a contact zone between three species of European vipers. Diversity and Distributions 14(3): 452-461.

Mediani, M.; Amezian, M.; Tattou, M.I.; Benhoussa, A.; Rguibi Idrissi, H.; El Agbani, M.A.; Qninba, A. 2009. Nouvelles citations de deux especes reliques palearctiques, Emys orbicularis Linnaeus, 1758 et Vipera latastei Bosca, 1878 dans la Peninsule Tingitane (Rif occidental, Maroc). Bulletin de l'Institut Scientifique Section Sciences de la Vie (Rabat) 31(2): 99-102.

Niskanen, M.; Mappes, J. 2005. Significance of the dorsal zigzag pattern of Vipera latastei gaditana against avian predators. Journal of Animal Ecology 74(6): 1091-1101.

Parellada, X. 1995. About the apparent inexistence of a spring mating in the Catalan population of Vipera latasti (Reptilia: Viperidae), and note about the reproductive success. pp. 250-254. In: Llorente, G.A.; Montori, A.; Santos, X. & Carretero, M.A. (eds.). Scientia Herpetologica: papers submitted from the 7th Ordinary General Meeting of Societas Europaea Herpetologica, Barcelona, September 15-19, 1993. Asociacion Herpetologica Espanola, Barcelona. 385 pp.

Parellada, X. 1995. Status of Vipera aspis and Vipera latasti (Viperidae, Reptilia) in Catalonia (NE Spain). pp. 328-334. In: Llorente, G.A.; Montori, A.; Santos, X. & Carretero, M.A. (eds.). Scientia Herpetologica: papers submitted from the 7th Ordinary General Meeting of Societas Europaea Herpetologica, Barcelona, September 15-19, 1993. Asociacion Herpetologica Espanola, Barcelona. 385 pp.

Parellada, X.; Santos, X. 2002. Caudal luring in free-ranging adult Vipera latasti. Amphibia-Reptilia 23(3): 343-347.

Pires Ceríaco, L.M. 2011. Geographic distribution: Vipera latastei (Lataste's Viper). Herpetological Review 42(4): 575.

Pleguezuelos, J.M.; Santos, X.; Brito, J.C.; Parellada, X.; Llorente, G.A.; Fahd, S. 2007. Reproductive ecology of Vipera latastei, in the Iberian Peninsula: Implications for the conservation of a Mediterranean viper. Zoology (Jena) 110(1): 9-19.

Real, R.; Luz Márquez, A.; Olivero, J.; Estrada, A. 2010. Species distribution models in climate change scenarios are still not useful for informing policy planning: an uncertainty assessment using fuzzy logic. Ecography 33(2): 304-314.

Saint Girons, H. 1989. Vipera latastei. pp. 173. In: Castanet, J. & Guyetant, R. Atlas de repartition des amphibiens et reptiles de France. Societe Herpetologique de France, Paris. 191 pp.

Saint-Girons, H. 1954. Le cycle d'activite et ses facteurs chez Vipera latastei Bosca. Vie et Milieu 5: 513-528.

Saint-Girons, H. 1977. Systematique de Vipera latastei latastei Bosca, 1878 et description de Vipera latastei gaditana. subsp. n. (Reptilia, Viperidae). Revue Suisse de Zoologie 84(3): 599-607.

Salvador, A.; Busack, S.D.; McDiarmid, R.; Ineich, I.; Brito, J.C. 2014. Case 3629. Vipera latastei Bosca, 1878 (Reptilia, Serpentes, Viperidae): request for conservation of the original spelling. Bulletin of Zoological Nomenclature 71(1): 22-25.

Santos, X. 2011. Amenaces i causes de la desaparicio de l'escurco iberica la peninsula Iberica. L'Atzavara 20: 77-82.

Santos, X.; Brito, J.C.; Pleguezuelos, J.M.; Llorente, G.A. 2007. Comparing Filippi and Luiselli's (2000) method with a cartographic approach to assess the conservation status of secretive species: the case of the Iberian snake-fauna. Amphibia-Reptilia 28(1): 17-23.

Santos, X.; Brito, J.C.; Sillero, N.; Pleguezuelos, J.M.; Llorente, G.A.; Fahd, S.; Parellada, X. 2006. Inferring habitat-suitability areas with ecological modelling techniques and GIS: a contribution to assess the conservation status of Vipera latastei. Biological Conservation 130(3): 416-425.

Santos, X.; Llorente, G.A.; Pleguezuelos, J.M.; Brito, J.C.; Fahd, S.; Parellada, X. 2007. Variation in the diet of the Lataste's viper Vipera latastei in the Iberian Peninsula: seasonal, sexual and size-related effects. Animal Biology (Leiden) 57(1): 49-61.

Santos, X.; Pleguezuelos, J.M.; Brito, J.C.; Fahd, S.; Llorente, G.A.; Parellada, X. 2004. La vibora hocicuda: una especie desconocida y amenazada de la fauna mediterranea. Quercus 216: 32-39.

Santos, X.; Pleguezuelos, J.M.; Brito, J.C.; Llorente, G.A.; Parellada, X.; Fahd, S. 2008. Prey availability drives geographic dietary differences of a Mediterranean predator, the Lataste's viper (Vipera latastei). Herpetological Journal 18(1): 16-22.

Santos, X.; Vidal-García, M.; Brito, J.C.; Soumia, F.; Llorente, G.A.; Martínez-Freiria, F.; Parellada, X.; Pleguezuelos, J.M.; Sillero, N. 2014. Phylogeographic and environmental correlates support the cryptic function of the zigzag pattern in a European viper. Evolutionary Ecology 28(4): 611-626.

Sanz, T. 2008. Vibora hocicuda, Vipera latastei Bosca, 1878: cita en el sector nororiental de la provincia de Leon. Boletin de la Asociacion Herpetologica Espanola 19: 62-63.

Schwarzer, U. 1999. Zur Verbreitung der Stülpnasenotter, Vipera latastei gaditana (Saint-Girons, 1977) in Südwest-Portugal (Squamata: Serpentes: Viperidae). Herpetozoa 11(3-4): 181-184.

Soler Massana, J.; Amill Franch, I.; Martinez-Silvestre, A.; Barrull Ventura, J.; Mate Alonso, I. 2006. Nuevos datos de distribucion para 9 especies de reptiles en la Comarca del Priorat (sudoeste de Cataluna). Boletin de la Asociacion Herpetologica Espanola 17(2): 66-73.

Stümpel, N.; Brito, J.C.; Saint-Girons, H. 2005. Vipera latastei - Stülpnasenotter. pp. 187-212. In: Joger, U. & Stümpel, N. (eds.). Handbuch der Reptilien und Amphibien Europas. Schlangen III. Viperidae. AULA-Verlag, Wiebelsheim. 420 pp.

Tarroso, P.; Pereira, R.J.; Martínez-Freiria, F.; Godinho, R.; Brito, J.C. 2014. Hybridization at an ecotone: ecological and genetic barriers between three Iberian vipers. Molecular Ecology 23(5): 1108-1123.

Trutnau, L. 1994. Zur Lebensweise, Pflege und Nachzucht der Stülpnasenotter Vipera latasti Bosca, 1878. Herpetofauna (Weinstadt) 16(91): 19-24.

Valkonen, J.K.; Nokelainen, O.; Mappes, J. 2011. Antipredatory function of head shape for vipers and their mimics. PLoS ONE 6(7): e22272, 1-4.

Velo-Anton, G.; Godinho, R.; Harris, D.J.; Santos, X.; Martínez-Freiria, F.; Fahd, S.; Larbes, S.; Pleguezuelos, J.M.; Brito, J.C. 2012. Deep evolutionary lineages in a western Mediterranean snake (Vipera latastei/monticola group) and high genetic structuring in southern Iberian populations. Molecular Phylogenetics and Evolution 65(3): 965-973.

Zaldivar, C.; Gosa, A.; Bergerandi, A. 1992. Primera cita de Vipera latastei Bosca, 1878 en La Rioja. Zubia 10: 190-192.

Vipera lotievi

Kidov, A.A.; Pikhov, S.G.; Tishina, G.V. 2009. [On biology of Lotiev's viper Vipera lotievi Nilson et al., 1995 (Reptilia, Serpentes, Viperidae) in the northern Ossetia]. (In Russian). Nauchnye Issledovaniya v Zoologicheskikh Parkakh 25: 158-160.

Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Zinenko, O.; Stümpel, N.; Mazanaeva, L.; Bakiev, A.; Shiryaev, K.; Pavlov, A.; Kotenko, T.; Kukushkin, O.; Chikin, Y.; Duisebayeva, T.; Nilson, G.; Orlov, N.L.; Sako, T.; Ananjeva, N.B.; Murphy, R.W.; Joger, U. 2015. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi). Molecular Phylogenetics and Evolution 84: 85-100.

Vipera magnifica

Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Tuniyev, B.S.; Ostrovskikh, S.V. 2001. Two new species of vipers of "kaznakovi" complex (Ophidia, Viperinae) from the western Caucasus. Russian Journal of Herpetology 8(2): 117-126.

Vipera monticola

Beerli, P.; Billing, H.; Schätti, B. 1986. Taxonomischer Status von Vipera latasti monticola Saint Girons, 1953 (Serpentes, Viperidae). Salamandra 22(2-3): 101-104.

Brito, J.C.; Santos, X.; Pleguezuelos, J.M.; Fahd, S.; Llorente, G.A.; Parellada, X. 2006. Morphological variability of the Lataste's viper (Vipera latastei) and the Atlas dwarf viper (Vipera monticola): patterns of biogeographical distribution and taxonomy. Amphibia-Reptilia 27(2): 219-240.

Freitas, I.; Fahd, S.; Velo-Antón, G.; Martínez-Freiría, F. 2018. Chasing the phantom: biogeography and conservation of Vipera latastei-monticola in the Maghreb (North Africa). Amphibia-Reptilia 39(2): 145–161.

Fritz, P.; Scheler, B. 2017. Das Porträt: Vipera monticola Saint Girons, 1953. Sauria (Berlin) 39(1): 1-2.

Herrmann, H.W. 1993. Bemerkungen zum Beutespektrum von Vipera monticola. Salamandra 29(1): 90-91.

Pillet, J.M. 1994. Nouvelles donnees sur la repartition et l'ecologie de la vipere naine du Haut Atlas Vipera monticola Saint Girons, 1954 (Reptilia, Viperidae). Revue Suisse de Zoologie 101(3): 645-653.

Saint-Girons, H. 1953. Une vipère naine: Vipera latastei montana. Bulletin de la Société Zoologique de France 78: 24-28.

Saint-Girons, H. 1954. Note rectificative au sujet de Vipera latastei montana, n. subsp. Bulletin de la Société Zoologique de France 78: 475.

Saint-Girons, H. 1973. Nouvelles donnees sur la vipère naine du Haut Atlas, Vipera latastei monticola. Bulletin Soc. Sci. nat. phys. Maroc 53(1-2): 111-118.

Velo-Anton, G.; Godinho, R.; Harris, D.J.; Santos, X.; Martínez-Freiria, F.; Fahd, S.; Larbes, S.; Pleguezuelos, J.M.; Brito, J.C. 2012. Deep evolutionary lineages in a western Mediterranean snake (Vipera latastei/monticola group) and high genetic structuring in southern Iberian populations. Molecular Phylogenetics and Evolution 65(3): 965-973.

Westerström, A. 2010. Early record of the elusive atlas dwarf viper Vipera monticola Saint Girons, 1953, in the Moroccan High Atlas. Herpetozoa 23(1-2): 103-104.

Vipera nikolskii

Bakiev, A.G.; Böhme, W.; Joger, U. 2005. Vipera nikolskii - Waldsteppenotter. pp. 293-309. In: Joger, U. & Stümpel, N. (eds.). Handbuch der Reptilien und Amphibien Europas. Schlangen III. Viperidae. AULA-Verlag, Wiebelsheim. 420 pp.

Bondarenko, Z.S.; Zinenko, O.I. 2016. Individual growth rates of Nikolsky's Viper, Vipera berus nikolskii (Squamata, Viperidae). Vestnik Zoologii 50(1): 65-70.

Efimov, R.V.; Zavialov, E.V.; Tabachishin, V.G.; Pomazenko, O.A. 2009. [Molecular-genetic analysis of rare and protected species of Volga vipers]. (In Russian). Izvestiya Samarskogo Nauchnogo Centra Rossiskaya Akademii Nauk 11(1) (27): 429-432.

Efimov, R.V.; Zavyalov, E.V.; Velikov, V.A.; Tabachishin, V.G. 2008. Genetic divergence of Vipera berus and Vipera nikolskii (Reptilia: Viperidae, Vipera) populations in lower Volga and adjacent territories assessed according to the sequences of cytochrome oxidase III and 12S ribosome RNA genes. Russian Journal of Genetics 44(2): 240-243.

Gumprecht, A. 1994. Zur Kenntnis der Russischen Waldsteppenotter Vipera nikolskii Vedmederja, Grubant & Rudajeva, 1986. Sauria (Berlin) 16(4): 27-33.

Jandzik, D. 2007. Husbandry and captive reproduction in Vipera nikolskii (Viperidae). Herpetological Review 38(2): 171-172.

Joger, U.; Lenk, P.; Baran, I.; Böhme, W.; Ziegler, T.; Heidrich, P.; Wink, M. 1997. The phylogenetic position of Vipera barani and of V. nikolskii within the Vipera berus complex. pp. 185-194. In: Böhme, W.; Bischoff, W. & Ziegler, T. (eds.). Herpetologia Bonnensis. Proceedings of the 8th Ordinary General Meeting of the Societas Europaea Herpetologica, 23-27 August 1995, Bonn, Germany. Societas Europaea Herpetologica, Bonn. 414 pp.

Loparev, S.A.; Sytnik, A.I. 2008. The comparative ecology of coexistent populations of Vipera berus and 'Vipera nikolskii' in the forest steppe on Dnieper’s right riverbank of Ukraine. pp. 258-261. In: Ananjeva, N.B.; Danilov, I.G.; Dunayev, E.A.; Ischenko, V.G.; Lada, G.A.; Litvinchuk, S.N.; Orlova, V.F.; Smirina, E.M.; Tuniyev, B.S. & Khalikov, R.G. (eds.). The problems of herpetology. Proceedings of the 3rd meeting of the Nikolsky Herpetological Society, 9-13 October 2006, Putschino. Russian Academy of Sciences, St. Petersburg. 467 pp.

Mizsei, E.; Jablonski, D.; Roussos, S.A.; Dimaki, M.; Ioannidis, Y.; Nilson, G.; Nagy, Z.T. 2017. Nuclear markers support the mitochondrial phylogeny of Vipera ursinii-renardi complex (Squamata: Viperidae) and species status for the Greek meadow viper. Zootaxa 4227(1): 75-88.

Oleynik, Y.V.; Nekrasova, O.D. 2012. [New data on the distribution of Vipera nikolskii (Reptilia, Viperidae) in Kyiv Region]. (In Russian). Vestnik Zoologii 46(3): 258.

Ramazanova, A.S.; Starkov, V.G.; Osipov, A.V.; Ziganshin, R.H.; Filkin, S.Y.; Tsetlin, V.I.; Utkin, Y.N. 2009. Cysteine-rich venom proteins from the snakes of Viperinae subfamily - Molecular cloning and phylogenetic relationship. Toxicon 53(1): 162-168.

Sokolov, A.S. 2005. On the taxonomic status of the common adder of the partially wooded steppe of the Oka-Don Plain. Russian Journal of Herpetology 12: 96-99. (Supplement).

Sos, T.; Hegyeli, Z. 2011. Morphotype of Vipera berus nikolskii Vedmederja, Grubant & Rudaeva, 1986, present in the Buzau Subcarpathians, Romania. Herpetozoa 24(1-2): 91-95.

Tabachishin, V.G.; Tabachishina, I.E.; Zavialov, E.V. 2003. Modern habitats and some ecological aspects of the Forest-steppe Viper in the north of the Lower Volga region. Russian Journal of Ecology 2003(1): 82-86.

Tabachishin, V.G.; Zavialov, E.V. 2003. Forest-steppe Viper distribution in the south of the Podolsk height. Russian Journal of Ecology 2003(2): 202-203.

Tabatschischin, W.G.; Sawjalow, E.W. 2004. Zur Präzisierung der südlichen Grenze des Verbreitungsareals der Waldsteppenotter (Vipera nikolskii) im Europäischen Tteil Russlands. Mauritiana 19(1): 83-85.

Tabatschischina, I.E.; Tabatschischin, W.G.; Sawjalow, E.W. 2002. Wachstumsdynamik bei Vipera nikolskii im Gebiet Saratow. Mauritiana 18(2): 203-206.

Vedmederya, V.I.; Grubant, V.N. 1986. [On the question of the name of the black viper in the forest steppes of the European part of the USSR]. (In Russian). Vestnik Khar'kovskogo Universiteta 288: 83-85.

Velikov, V.A.; Yefimov, R.V.; Zavialov, E.V.; Kuznetsov, P.E.; Tabachisin, V.G.; Shlyakhtin, G.V.; Kaybeleva, E.I. 2006. [Genetic divergence of some Vipera species (Reptilia: Viperidae, Vipera) from gene sequention of NADN-dehydrogenase and 12S ribosomal RNA]. (In Russian, English summary). Sovremennaya Gerpetologiya 5-6: 41-49.

Yefimov, R.V.; Zavialov, E.V.; Tabachishin, V.G. 2008. [Aspects of ecological segregation and technology of specific identification of viper snakes (Reptilia: Viperidae, Vipera) in Volga region by genotyping]. (In Russian). Povolzhskii Ekologicheskii Zhurnal 2: 147-153.

Zavialov, E.V.; Kaybeleva, E.I.; Tabachishin, V.G. 2006. [A comparative karyological characteristics of forest-steppe viper (Vipera (Pelias) nikolskii) from the small river flood-lands of the Volga and Don basins]. (In Russian, English summary). Sovremennaya Gerpetologiya 5-6: 100-103.

Zavialov, E.V.; Yefimov, R.V.; Tabachishin, V.G.; Pomazenko, O.A. 2009. [Genetic characteristics of Vipera renardi (Reptilia: Viperidae) populations in the lower-Volga region and adjacent territories from sequenation of mitochondrial genes of cytochrome b and c, NADN-dehydrogenase and 12s ribosomic RNA. (In Russian). Sovremennaya Gerpetologiya 9(1-2): 12-17.

Zinenko, O.I. 2003. [First generation hybrids between the Nikolsky's adder, Vipera nikolskii, and the common adder, Vipera berus (Reptilia, Serpentes, Viperidae)]. (In Russian, English summary). Vestnik Zoologii 37(1): 101-104, 107.

Zinenko, O.I. 2004. New data about hybridization between Vipera nikolskii Vedmederya, Grubant & Rudaeva, 1986 and Vipera berus berus (Linnaeus, 1758) and their contact zones in Ukraine. Mertensiella 15: 17-28.

Zinenko, O.I. 2006. Habitats of Vipera berus nikolskii in Ukraine. pp. 205-209. In: Vences, M.; Köhler, J.; Ziegler, T. & Böhme, W. (eds.). Herpetologia Bonnensis II. Proceedings of the 13th Congress of the Societas Europaea Herpetologica, 27 September - 2 October 2005, Bonn, Germany. Societas Europaea Herpetologica, Bonn. 262 pp.

Zinenko, O.I.; Turcanu, V.; Strugariu, A. 2010. Distribution and morphological variation of Vipera berus nikolskii Vedmederja, Grubant et Rudaeva, 1986 in western Ukraine, the Republic of Moldova and Romania. Amphibia-Reptilia 31(1): 51-67.

Vipera olguni

Tuniyev, S.B.; Avci, A.; Tuniyev, B.S.; Agasian, A.L.; Agasian, L.A. 2012. Description of a new species of Shield-head Vipers - Pelias olguni sp. nov. from basin of upper flow of the Kura River in Turkey. Russian Journal of Herpetology 19(4): 314-332.

Vipera orlovi

Ostrovskikh, S.V. 2008. Colour morph of Orlov’s Viper Vipera (Pelias) orlovi (Ophidia, Viperinae). pp. 337-340. In: Ananjeva, N.B.; Danilov, I.G.; Dunayev, E.A.; Ischenko, V.G.; Lada, G.A.; Litvinchuk, S.N.; Orlova, V.F.; Smirina, E.M.; Tuniyev, B.S. & Khalikov, R.G. (eds.). The problems of herpetology. Proceedings of the 3rd meeting of the Nikolsky Herpetological Society, 9-13 October 2006, Putschino. Russian Academy of Sciences, St. Petersburg. 467 pp.

Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Tuniyev, B.S.; Ostrovskikh, S.V. 2001. Two new species of vipers of "kaznakovi" complex (Ophidia, Viperinae) from the western Caucasus. Russian Journal of Herpetology 8(2): 117-126.

Vipera pontica

Baran, I.; Tosunoglu, M.; Kaya, U.; Kumlutas, Y. 1997. Camlihemsin (Rize) civarinin herpetofaunasi hakkinda. [On the herpetofauna of the vicinity of Camlihemsin.] (In Turkish, English summary). Turkish Journal of Zoology 21(4): 409-416.

Billing, H.; Nilson, G.; Sattler, U. 1990. Vipera pontica sp.n., a new viper species in the kaznakovi group (Reptilia, Viperidae) from northeastern Turkey and adjacent Transcaucasia. Zoologica Scripta 19(2): 227-231.

Kutrup, B. 2003. The identification of new specimens of Vipera from Trabzon, Turkey with affinities to Vipera barani and V. pontica. Herpetological Review 34(1): 28-31.

Vipera renardi

Aliev, T.R.; Ganiev, F.R. 1985. [Distribution and ecological-morphological features of Renard's viper - Vipera ursini Bonaparte, 1835 in Azerbaidjan]. (In Russian). Izvestiya Akademii Nauk Azerbaidzhanskoi SSR Seriya Biologicheskikh Nauk 1985(1): 44-50.

Bakiev, A.G. 2012. [Viper Vipera renardi (Christoph, 1861): about the species first description and its author - Hugo Christoph]. (In Russian). Izvestiya Samarskogo Nauchnogo Centra Rossiskaya Akademii Nauk 14(5): 155-158.

Bakiev, A.G.; Garanin, V.I.; Litvinov, N.A.; Pavlov, A.V.; Ratnikov, V.Y. 2004. [Vipera renardi (Christoph, 1861)]. pp. 61–91. (In Russian). In: Garanin, V.I.; Pavlov, A.V. & Bakiev, A.G. [Snakes of Volga-Kama Region]. Samara Sci. Center, Samara.

Bakiev, A.G.; Litvinov, N.A.; Shurshina, I.V. 2010. [On diet of Renard's viper Vipera renardi (Christoph, 1861) in the Volga river basin]. (In Russian). Sovremennaya Gerpetologiya 10(1-2): 54-56.

Basoglu, M. 1947. On some varieties of Vipera berus from the extreme north-eastern Anatolia. Revue Faculte des Sciences Universite d'Istanbul 12B(3): 182-190.

Boulenger, G.A. 1894. On Vipera renardi, Christoph. Proceedings of the Zoological Society of London 1894: 757-760.

Efimov, R.V.; Zavyalov, E.V.; Velikov, V.A.; Tabachishin, V.G. 2008. Genetic divergence of Vipera berus and Vipera nikolskii (Reptilia: Viperidae, Vipera) populations in lower Volga and adjacent territories assessed according to the sequences of cytochrome oxidase III and 12S ribosome RNA genes. Russian Journal of Genetics 44(2): 240-243.

Fomina, M.I. 1965. On the feeding habits of the steppe viper in spring and summer periods. Zoologicheskii Zhurnal 44: 1100-1103.

Fomina, M.I. 1968. Habitats and resting places of the Renard's viper (Vipera ursini) in Chu-Ili. Interiver Region. Zool. Zh. 47: 782-785.

Gvozdik, V.; Jandzik, D.; Cordos, B.; Rehak, I.; Kotlik, P. 2012. A mitochondrial DNA phylogeny of the endangered vipers of the Vipera ursinii complex. Molecular Phylogenetics and Evolution 62(3): 1019-1024.

Joger, U.; Dely, O.G. 2005. Vipera renardi - Steppenotter. pp. 343-354. In: Joger, U. & Stümpel, N. (eds.). Handbuch der Reptilien und Amphibien Europas. Schlangen III. Viperidae. AULA-Verlag, Wiebelsheim. 420 pp.

Kidov, A.A. 2009. [Renard's viper Vipera (Pelias) renardi Christoph, 1861 (Reptilia, Serpentes: Viperidae) from northwestern foothills of the Caucasus]. (In Russian). Nauchnye Issledovaniya v Zoologicheskikh Parkakh 25: 161-165.

Kotenko, T.I. 1981. [On Vipera ursinii renardi in the south of Ukraine]. pp. 73. (In Russian). In: Darevsky, I.S., Ananjeva, N.B., Barkagan, Z.S., Borkin, L.Y., Sokolova, T.M. & Shcherbak, N.N. (eds.). The problems of herpetology: abstracts. Nauka, Leningrad. 163 pp.

Kotenko, T.I. 2002. [New finds of the steppe viper (Vipera renardi) in the Sivash area of the Crimea]. (In Russian). Vestnik Zoologii 36(2): 100.

Kotenko, T.I. 2007. [On distribution of the steppe viper, Vipera renardi (Reptilia, Viperidae), in the western part of the lowland Crimea]. (In Russian). Vestnik Zoologii 41(5): 422.

Krasavtzev, B.A. 1943. Zur Biologie der Steppenotter im Nord-kaukasus. Zoologischer Anzeiger 142: 141-144.

Krecsák, L.; Toth, T. 2005. Biting among male steppe vipers Vipera renardi (Christoph, 1861). Herpetozoa 18(3-4): 188-189.

Kukushkin, O.V. 2009. [Vipera renardi puzanovi ssp. nov. (Reptilia: Serpentes: Viperidae) as a new subspecies of steppe viper from mountain Crimea]. (In Russian). Sovremennaya Gerpetologiya 9(1-2): 18-40.

Kukushkin, O.V.; Zinenko, O.I. 2006. Morphological peculiarities and their possible bearing on the taxonomic status of the Crimean montane populations of the steppe viper, Vipera renardi (Christoph, 1861). pp. 61-66. In: Vences, M.; Köhler, J.; Ziegler, T. & Böhme, W. (eds.). Herpetologia Bonnensis II. Proceedings of the 13th Congress of the Societas Europaea Herpetologica, 27 September - 2 October 2005, Bonn, Germany. Societas Europaea Herpetologica, Bonn. 262 pp.

Laister, A. 1908. [Zur Fauna des Gouv. Erivan. Über das Vorkommen von Vipera renardi Christoph. in Transkaukasien]. (In Russian). Erivan, Pam. Knizka Erivansk. gub. 1908: 223-226.

Mazuch, T. 2004. Nekolik poznamek ke komplexu zmiji stepnich 'Vipera (Acridophaga) ursinii' s durazem na biologii zmije stepni Vipera renardi renardi Christoph, 1861 v chovu a prirode (2). [Some notes on the complex of steppe snakes 'Vipera (Acridophaga) ursinii' with emphasis on the biology of Renard's Viper Vipera renardi renardi Christoph, 1861 in captivity and in the wild (2).]. (In Czech). Akvarium Terarium 47(4): 63-67.

Mizsei, E.; Jablonski, D.; Roussos, S.A.; Dimaki, M.; Ioannidis, Y.; Nilson, G.; Nagy, Z.T. 2017. Nuclear markers support the mitochondrial phylogeny of Vipera ursinii-renardi complex (Squamata: Viperidae) and species status for the Greek meadow viper. Zootaxa 4227(1): 75-88.

Nilson, G.; Andrén, C. 2001. The meadow and steppe vipers of Europe and Asia: the Vipera (Acridophaga) ursinii complex. Acta Zoologica Academiae Scientiarum Hungaricae 47(2-3): 87-267.

Nilson, G.; Andren, C.; Joger, U. 1993. Re-evaluation of the taxonomic status of the Moldavian steppe viper based on immunological investigations, with a discussion of the hypothesis of secondary intergradation between Vipera ursinii rakosiensis and Vipera (ursinii) renardi. Amphibia-Reptilia 14: 45-57.

Ostrovskikh, S.V. 1997. Different forms of melanism and its development with age in the populations of steppe viper Vipera renardi (Christoph, 1861). Russian Journal of Herpetology 4(2): 186-191.

Ostrovskikh, S.V. 2006. [Morphological variability of eastern meadow viper - Vipera (Pelias) renardi in the North-Western Caucasus]. (In Russian, English summary). Sovremennaya Gerpetologiya 5-6: 61-70.

Pavlov, A. 2005. Steppe viper (Vipera renardi) in the northern point of its area. Russian Journal of Herpetology 12: 87-89. (Supplement).

Pavlov, A.V.; Zinenko, O.I.; Joger, U.; Stuempel, N.; Petrova, I.V.; Malenyov, A.L.; Zaitseva, O.V.; Shurshina, I.V.; Bakiev, A.G. 2011. [Natural hybridization of the eastern steppe viper Vipera renardi and the common adder V. berus]. (In Russian). Izvestiya Samarskogo Nauchnogo Centra Rossiskaya Akademii Nauk (Suppl. 43) 13(5): 172-178.

Puzachenko, A.Y.; Baklushinskaya, I.Y.; Lyapunova, E.A.; Vlasov, A.A. 1997. Karyotype of Vipera ursini (Reptilia, Viperidae) from Streletskaya steppe. (In Russian, English summary). Vestnik Zoologii 31(3): 81-82.

Tabachishina, I.E.; Tabachishin, V.G.; Shlyakhtin, G.V. 2007. [On revision of the northern boundary of the Vipera renardi habitat in the Volga region]. (In Russian, English summary). Povolzhskii Ekologicheskii Zhurnal 3: 271-277.

Toktosunov, A.T.; Mazik, E.Y.; Kassymalieva, E.B.; Imanalieva, N.D. 1992. Specific characters of karyotypes of some snake species (Colubridae, Viperidae, Crotalidae) from the northern Tien-Shan. Zoologicheskii Zhurnal 71(5): 118-127.

Tsaruk, O.I. 1981. [New data on the distribution of some Amphibia and reptile species in southeastern Kazakhstan]. pp. 143-144. In: Darevsky, I.S.; Ananjeva, N.B.; Barkagan, Z.S.; Borkin, L.Y.; Sokolova, T.M. & Szczerbak, N.N. (eds.). The problems of herpetology: abstracts. Nauka, Leningrad. 163 pp. (abstract, in Russian).

Tseitgamel', Y.S. 1976. [Renard's viper Vipera renardi]. (In Russian). Zashchita Rastenii (Moscow) 1976(6): 61.

Tuniyev, B.S. 2016. Rare species of Shield-head Vipers in the Caucasus. Nature Conservation Research 1(3): 11–25.

Ujvari, B.; Madsen, T.; Kotenko, T.I.; Olsson, M.; Shine, R.; Wittzell, H. 2002. Low genetic diversity threatens imminent extinction for the Hungarian meadow viper (Vipera ursinii rakosiensis). Biological Conservation 105(1): 127-130.

Velikov, V.A.; Yefimov, R.V.; Zavialov, E.V.; Kuznetsov, P.E.; Tabachisin, V.G.; Shlyakhtin, G.V.; Kaybeleva, E.I. 2006. [Genetic divergence of some Vipera species (Reptilia: Viperidae, Vipera) from gene sequention of NADN-dehydrogenase and 12S ribosomal RNA]. (In Russian, English summary). Sovremennaya Gerpetologiya 5-6: 41-49.

Wettstein, O. von 1953. Vipera renardi aus Persien. Zoologischer Anzeiger 150: 266-268.

Zavialov, E.V.; Yefimov, R.V.; Tabachishin, V.G.; Pomazenko, O.A. 2009. [Genetic characteristics of Vipera renardi (Reptilia: Viperidae) populations in the lower-Volga region and adjacent territories from sequenation of mitochondrial genes of cytochrome b and c, NADN-dehydrogenase and 12s ribosomic RNA. (In Russian). Sovremennaya Gerpetologiya 9(1-2): 12-17.

Zinenko, A.I.; Bakiev, A.G. 2007. [About change of the northern boundary of distribution of the steppe viper, Vipera renardi (Reptilia, Viperidae) in European Russia]. (In Russian). Vestnik Zoologii 41(5): 478.

Zinenko, O.; Stümpel, N.; Mazanaeva, L.; Bakiev, A.; Shiryaev, K.; Pavlov, A.; Kotenko, T.; Kukushkin, O.; Chikin, Y.; Duisebayeva, T.; Nilson, G.; Orlov, N.L.; Sako, T.; Ananjeva, N.B.; Murphy, R.W.; Joger, U. 2015. Mitochondrial phylogeny shows multiple independent ecological transitions and northern dispersion despite of Pleistocene glaciations in meadow and steppe vipers (Vipera ursinii and Vipera renardi). Molecular Phylogenetics and Evolution 84: 85-100.

Vipera sakoi

Tuniyev, B.S; Avci, A.; Ilgaz, C.; Olgun, K.; Petrova, T.V.; Bodrov, S.Y.; Geniez, P.; Teynié, A. 2018. On taxonomic status of Shield-head Vipers from Turkish Lesser Caucasus and East Anatolia. Proceedings of the Zoological Institute RAS 322(1): 3–44.

Vipera seoanei

Acuna, A.; Martínez-Freiría, F.; Velo-Antón, G. 2013. Ingesta de tres especies de herpetos (Bufo calamita, Anguis fragilis, Vipera seoanei) por un ejemplar de águila culebrera (Circaetus gallicus) abatido en un parque eólico. Boletin de la Asociacion Herpetologica Espanola 24(1): 59-60.

Bea, A. 1978. Contribution to the systematics of Vipera seoanei Lataste, 1879 Reptilia, Viperidae. 1. Ultrastructure of the scales cuticle. Butlletí de la Institució Catalana d'Història Natural (Sec. Zool.) 42: 107-118.

Bea, A.; Bas, S.; Brana, F.; Saint Girons, H. 1984. Morphologie comparee et repartition de Vipera seoanei Lataste, 1879, en Espagne. Amphibia-Reptilia 5(3-4): 395-410.

Bedriaga, J. 1895. Notice sur le "Peliade Portugais". Ann. Sci. nat. Porto 2: 114-116.

Billing, H. 1983. Polymorphismus bei Vipera berus seoanei. Herpetofauna (Weinstadt) 5(24): 31-33.

Boudarel, P.; Haffner, P.; Hippolyte, J.; Leconte, M. 1994. A propos de la presence de la vipere de seoane (Vipera seoanei Lataste, 1879) (Ophidia, Viperidae) en Haute Vallee des Aldudes (Pays-Basque, Pyrenees-Atlantiques) et de sa limite orientale en Pyrenees. Bulletin de la Societe Herpetologique de France 65-66 [1993]: 15-21.

Brana, F.; Bas, S. 1983. Vipera seoanei cantabrica ssp. n. Munibe 35(1-2): 87-88.

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Vipera walser

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